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eee upubsetan tirade tha iinet 


as gna 90 Oee 
peer nines OT une 
est proses enet 
aabegrarsee eter ee ts: 


deg peier ern enae anit ene es SALTS TS De Tien 
Mitesh i leneenaatatereestyry siete te mont ieee pcabeasdeeedeeobene rereenes® 
Rp iecarette : os ‘s ter ope eebeasereeetetsae 
pa arberet > cb pe tet tt ean err Dyk ianesd eee od A re ETT TE hae 
spaa vat enb ay tae reat snaraunt ciemeemenennarest oot rowss oes 

uy poeeneperse see ety panne Lai aeeratesegrpoettedt s bber¢: pe reee pine hr 
‘ ce Slerpsmyeeates sStacducorbranersenstntene negra fal ranibey anes heen ari ore ensaeueerees hi 

hd too paar hints bahar ot sole Presb Poe sae ae et 

sdeecha preset i ait ogy ee rhe peeerinns cra dt oi raparsecd sts! 
o> paeebeentye reel Meets ee ee hh Oot eet oh fee cnediee nists 
pared habbo ea ivrs err peesoyeeniy oct ppepaons mt raven st vo 8 eae eieteenesereniaee rt 
7 penvens pyenen snghe beeetee whe eae Met Pest yi aehonterpbyey ere 442 denne 
iavaresordedotooyesnsdebehts mice rarueabeprarpessrenrgrrronett ett ty ererey 
” bisaeseeeeneeenernenrrneT ry. > a Peering a babar eerntr iON eT oi ed spel abet ots 
ppmpad og gpneneenepiteddet 1 TS Givers Pra oh wibeenrly al Sc omet ot a ae cages bel et 
pent parbrbeevtuoat orenboeeted ete ey ej ebeteneneh si) 
oho, he te rare nea epaseeer yt 
abe ener eres? 
Bisbee epee ret twenties: 


ere vegas) a me 1 
ote ptetede Lippe yb ret tSehorcisgceescrmectenetst of ae et eye tw Meh ered te bye sated 
tad abebae be aarees hme th dedoreeanreyeratge! ve reaeee eet pabsbot ngieee st Oye epieiaetebel teh pbs dip enpeey teem Dee Eth ates 
rin bs boys ot ate Ribtintuneeeettereipeeriten pesstaes ta we nese ere ey wabebreed cha¥ bre vyTer ee ened eral possuensett 
ot “ty a re panos otipener entre RE ataneperepeetyes Taser ee et bor weetet er Teles yeatoeet ery boternesee tye leben ead 
4 peers paneyetentiess + reader ayreneeeeve sane se prentrppe ete eé ne Tf es 232 gh eee e ett eet ep voobet oh See eee eieiapeneretete’ wh 8 9 
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L6r othe hot Lexeed Meares oar att te yprheeier toes Hehe isi aie 
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oboe s' Ary ead sepenenyee Sanpeomnn hyp y 
itera pape re mpatepeeey te Seren) rrp te be 
are ert a ameernererenyr ht rs iol ibe 
pene arche tet ohbe 


dente andes oor: 
rdeyreeted 
sprhritaty +i Oro 


M4 
Pilea ey nian r 
ephbeeseeser sana ht erer wasksaee 


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ped a ues 


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x ! } abe cnet et Mae e" oo 
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po Mee 9 pee to ah S “- ; sad te ctesteptant rs oe 
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| : Huei wh ont me eh re pavegetats 
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oussabpalited rhb > preeanprastiagseee) 0°: bee beet abpaee shes 
gievantennet cits earartane maaan oe a aeteathoee a et pe 
rie bar aeeerepavege sores fer oeegter eri 4 
gorereennces pratiose oe 


press 
sae eeeeprrnasante tery bor sheneduln pat ¢ ee anaes! 
US aecnaree paveperentytye yee. piehiseeahi sot at hh ete raareabebede rats 
behed seve 4 ae eye Pobre teres vente Teaavestetdoeastprpe neh (+ 

,! apiditnisben erie ret tree ania aiiverrebadebet 


ea vib enor oF yeh 
sie tH -wer el 
poised ged fyevererere ta! 
1 OTE eae ier preeeretedl 
Seeeeeebed 


ah 
meeeneye 
Teeeketon eee 
morass riatat te Sanrpasberee ect Teanaate rat aee 
Pitas yore te jeyreeyet researc epeuaean wansrenreanieetstgt cht. 
oatiegteo re aerit a otad en beet Seathey sy ened eer pisharot peut ara ots 
priser haseeenesy 2) « on 1 ee tplaumy serreareaeientttats ie Te alppeseecerree aeieeret 09 10 ee est pees 
phaehy tenes ey vate ; phan sessed . pcselaatisiepsente v4 peep anateet eh SIT DAA martserigeeare ey court trent 
ween a perensareenesy Rar pubsbesrbeseh ye Cotes She Mee Teh Tels dates is 
? myth! spemetedet i phere Peper oy Paris deena oer 11 1E Te ee ibebrneoet dehy blgas eset rears eth ty 
vend 4 pes Ory rs hes’ Ti rueptnenre ern have) te qertige the tthe 
pris vi seiepersetnped etarsbresdet Oot je re t Ott ee igeasererbabeereel penstrosl syriverestetats 
Ee eae es ee jebetes sirieiet-tahe) Srateget pipe yesnerinye 
part ipleeiy wien y 
Tenn aahaaena| > 


rp raadet egal benwpertrar ey. 
Uppate bsg stencenneeett 
sargibesal senaentse® 
agatha 4 eroneee) 


pidabepeese ery? 
bot 


bods posane Aenrwnetaty rAbet 
pelelessisbepet efeeatatirenss #0 208) Tpabed gerert sarees 
" pone peanrwap sie Taseeaepeereyeireeren Py 
. Paberererireehenpene rrapenin eh ETT SS eeT 


nants heoeitetertettalst erect ek 
ran eshe Lavehetetls peered erves ener? eRe me hee 
ee H eectiahspestateeeny fepeneen rer ppraaet ses) 7) 
cheat shone Cttgtginemassichects tia eteeater et 
doves Me haheaed pate sete edte Fat Fy. + vere 
me bat ri stateh 0 nse abeovben ttt 


ap eaneeny 
eqegeert 


oe 
aeeey 


"ts 
ag aged aoe! 


amet iatean ac gepmeettat se rosy) 
dee vege EEE 


ra jaagaerraeieneen ale 
farsa reeset bettraneedy seerarieaetet oat 
preven es emanate eit 
reriveasciopetmearseceseatie Pameapertter tins anauasreeet 
reparierisensnetecanereetseeenen terest Srremeeteet tarcremers pies auace ge oot) 1089) 
craic terms oer 


« 
4 : + ne 
uy ss aba yey temetetotesypersy at 
or ppvnarenest 2 eealara maerrent Pee eee 
1 ote pheaveched 
aetna! 


serdehe-ehabrnctsameenete 
Se pcrrereeerern 
epemeaegen® 
ye ay etstalasacinebeovenninereniterente 
pent ciyn asthe sty a aegat id rbepshabetepat’ Parsee Aetiyons 
9 ja . + 
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goto be og rpg oer erethtataechasmati ta ara siyevesOeveen reoteheasaresdne¥4) saver itndeeretty 
yeeewn ep! ST aah oaae ey om 
ao renipermerett? Mseererese tM ohapeeyer en iti nye resid erase 088 + 0 Ae ee ee ice depemnivereed acai as eke iohehors eet 
‘ ve panteasopberinne SPUN i A924 90 ob Loehesepeaerienrveespewretyt “ Slanasaeterpeee rer etetene ner tecrepceene rescbtye 2 
"i sao aee read neta oH Leb ‘Tbvbb eed scare ese ett sr atitpenneevounseseetotterttrcese tot Te eects alaLasl 
Lehshvet4 pabtpery OF ee ee ee a ohehe) acetet bi veoh + 
ess oy pabiasti8 ipetven ve.berere aH siecorardegeeetvtron ba yecpenenat ve 
aoe rbarayenery tty Saye ett 
nti, ny There Lac mse higae aay’ Ree  hidachebabesenedetee' 
ores ver rhs aged yeeey Lepeveesen ees 
setstectteryay Athy babvintot ve sropegabrncbeidntet eet ETT hog spliarane ieee eet 
beresaeey Deaeeo@be4 peaegapaaenabeprtecs torah tt! epeseen avers 
oss wie repianmiby pyesaues pect’ 
rite athcnelcheneoemeeneee 


guaee: pores eoetoy 
spate eapeted 
pera 


este me 
sr ed oe EB OD OP aeearata ne 
M ee fs ot 
“is tart tnes: aoe 
daiae oe) iss 
re 


2s 


oni 


ote hash teed Men ePer eT Fy 
eaieaatennatintiennist 


bepsargeter 
ranger ey eee 


ans rt 
oabia re anemes sel ee' 
ae 


peice 


ge 
sb ebawrd 
an sunbed eobeeet 


sprpneby sr 
aooae enietontde 


” 
nSrdetpehegeuprert ve 
eh lee eenea: bey 
ded qolpededeter 
pevsere ferro 


pans petit 
Pe Ee hahel 
aera neton bh oh 


” 
pupepnenlyys Pat! 9 re 
Ee has 


dap nprmeeers 
ged veaeepeee! 

54 babe 

+4 

prpotenaee? #9 
ae 


ve ere ra peg 
che pe penenrereer 
eat 


Hane 

444 ine dtaegadin Oro ede s 

Se tarboredae tt pens pes bal 
Se ee saearttt samo snyeert 
% 


re 

eras eed) ee 
retails ieeeyei®? Vaud-oe 
Wintojepeangueenyeye se ¢ 


read red aen dah 


pat tripege poner 4 
proegnsegenedeee! FP wp abegateengentposeaad 70% 
lee argent 4 oravevarset ee iwe od segs! arerene ety at 
7 4 ope joadn tear cee 
ole pid eee 
banana he 


tsatentaen oT 
Hy 
pepaaureh eersebeete lt ay) ‘ 
hua aagep qenem onder eertts aprei@ringed 
dogonenesoooen ented? 2h ede 
cpebedenageereste ate 
opee yathened re 
Dati ie 
npr pi pat 7 
‘ MaseePr erry phoitapanetene beer tet es rinprencane seat ea ‘ 
oye) eereve.ty ae be ree? 
st + Me aedigenepeiesnagenee® eet “  Iebeegaya beeen babeeid sorts rei sisal Sabet vibe sheet Shermans pesosesi sen stciet sh esPese eect 
pgsergeday serene’ sebeayave nearer yee sbrehy sesat eae eiecerdrsterteattans! apt nesey ted we St aceenurssesere? 
rnianla taste Seaeeratent renters aatortatienaeenaeeet Pibeiattiah cto ei 
end Dh etee ° ie tte pe pdteae! rep 4 1 peeesae (ieee) mest 
hi pense tt anh en o tS 
erry pr yee nba eipe§ ; PPR red entrees pbs 
wiigeths Marsejeerartedie sabcouirensngesten TORII rec opget Sa phgranee ee 
ite wate ry ethos spre ere ‘ Te eee cheeepat onan eaeha peceest 
peegene ated 
bepeend 


oes tte y+ +? ras 4 
swe bedbegegersd 
sin 4 “ ‘ orese ; " pene r 
jevere isiaarente 
writ ott Sstarichsareet pagegeortpeine® 
serene 


Pry 
aeepenet ie 
saan ener re 
# Ie te 
[ha eeeaerey 
Poedeisee m oaecbentee hae ney iF meaner iste paeae ene) oege! ees 
pad ebcayptingarien vetoes! CA Pe i ‘Ni e0 484 
eer essary ees Heraehegens drat reat mean + hee atseas haneenée ie “ny tore hatatsty ‘Y ‘ 
sTursedtegsameneyhyiet ty any iy os yejeqe ints ancien mera sea eee 
aa Lett shud aniie rene Catia Pra ea Seaktnerettci ae te he het 
ee aa lacoeastsistett its Pte BH teh ieee he biti quai ieee’ ay 
H viene erred. 
MY bettas rhviaa sr se ite a) ieseteherecoreabetersy ett. ; 
Witetetecoese Uhcbeitestete protease eth 
ant epeqedrnserplaes 
+ Cie abe rs viene pepeangens ree pos 
oe ere meaeedl op onahane ga? yes eee" 
seernetewierrs Perceatunteaate a betttngetawnanne fs paornne ree eestor 
caapmieeprs agente oF oe oooh ig alter meveyetsy + es areniees Be we 
‘ Lasisetanbes I ba honors pepageqercnneleyes yenenegey 
sop pj svar evelenet 
+ 
if hae + Senete uae pares 
- tee 
Iereysbeoedea oer sterghtcet eugene 


pete tedesne ers “er 


gepepete Pele 
Hester 


epee 
pepannn ether eee Ceeni treaty Mitt 
ae obs berdrabeenpnesveuedtt dete: Henrive ere aeons # jee 8 
sap enaniatt ateatan eee 
tet niet yen hed she genetsa " Chvhepennaneey 4 2 
seieenttied oetete 


ay atetiee wy bg oye 
shasta sbudh iveerentegnyre sest® 


prereie te arts 
poe cia gngene tt eee! ie ae 
pene tb mestnerieetty Pee rarersersiss bebe rise peeteded 
oe preplelyerd bs pas hene caepae petted opera eter ates ce ntey =a ni av emnen toned ty 
+4 
+ 
~ La bt oe spobanpyt veyt 


grawins oy 9 
oad oes 


Seat 
Dade dete | bbe ee 


parerer re 
- 

Sueescs peaveni graven yeti ave ie 
ohare Eby et Poteet ere ee - 


m oponey ofp nee sr Pet 
Se TL manatee’ Saori hareet 


Troawneatl fonds 

ptt 

ecient: 
va 


whey 
pera erty yet] 
wot 


ne 
saqebered) teat 
reprereet 
bel rhe) apes 
* 


erpennerat 
eee bd 


pt eee aehanenenel 
phen bicnte rye ree 
sepbieveses (oe? 


daray rtp oibote feat 
tgivetat 

wT band 

can peecennerer 


fe rk bed bee pos 
papteginagar! reve 


jae 5 
Ae serpoeedr holed HY ae th ogee 4 of panned en eee 
ere ae deg Sartor wpegrornngstsrcest eseree lier bie 
nd ek od coy ORONO wat aeee) pyrosuapes tied 
ty ea rtent eh bee pete vty se een dog apes or 
ay eres 
event tataestataes 
oo senere ee? 
neree 
papeqvi ene te ee is 
Agence teen 


a bbinerrert 


* 
av ey ashe 
lgvuagee ni raved obret sere 
maierent at ai te daar 
we be Meee parernevnsanerer hansery tenet yee t) meeres sere et 
reese! ) MT Tacha weeeend pisan ret satabyages 
42 4s deep pee eeree 
soae Tach. haaeaneaneee 
eeerrnresenr®” 


pene tr eee 


habe epee es dyer three 
Pen aegh pet da eees 
pe vee geyeyapee hte 


Poe a saben vy ot 
aneeraeyon pend Vie 


el navel 
pede etiee 
Taney rere penvee 


eprierey 


4 apeeteened 6 
Ned 


pica 

, prewar ese p air OP peed 

pearance ath 
arson spent Pre pep ah genertey OD ee ied 

peso 0884 rade th Gent ve" bed -eooem 


eae? borer at anes 4 
er o 
ware t rel riead waimuey oer? 
ictmeareter ry he rade sand gape oneard peste Nitrates 
pT boa anaed payers Salary 
eae pe aiotegsbeecwrersegcounns ei ent Hert titbnotts Sfititectercb tet? viet: base seorsh od shat one coagaenrwengcques te eeeety 
ea ee ea cane MaiLbee pert ty pee yaya poverd 024 0202 {Te beleneneeent eters tes yorerens 
nevisu riven PEPE caeafigiy sees penbeacterreanreiy et yr eerie Veet eh TT Sec pan-easysoeacrasne ties 
werner’ arantnenreeatisatae setae 1s reap tenth eereteeeaey etauattegtcniee gira tint neanee erprea a ancegt eeterhny 
overt ooetee ” or yee we. 
Seedee ne wens ee “ bepsentot wt asst pas one Sites abscesien satnuernveet yt 
> aad pacman he te pyeyt] aegeus eee Leases sapreeen eee at pra rrens ttt peer Lr, shah Lheabe eee eerrree 
Vnssenreeniy 1 eeonvanenrt! hel sa. pedewry loved Seeatecireuaneneer est ep gatguns fs trysEe gw rieseteogenr et snet 
a bot Pobeb pan esey pawererst porrereeere Sth proaneanyt sanay yg be ney 
Senter earvnsgit ent saree tet ap Lp Penbeye by Sepa 4 ee rne ee 
meee i peener peek PPP d hte? bine de? 
abet Ada wrtemee- dye ph 


Shep t neue rye joie vet 
‘ qeetcaes } prbenrnniener S20. e2 bade e Tj epeneiet=! 
n wen iat jeeepourbonenng relieve Swit tarenewtrt ttre 
annie? yrange finsarethtetiee copes 


Ara 
ere emrenane and 
tng 
het 
pose ery tacos veer eerie 
ase 198 ei wEarenes: stchertoricasesen cea ptrel eat 
rere Je setnget 
in ~ rath Latates tagueern yeverry rte} eapeprier yr ren yi. boeenant rg hon erenip eropre neg ’t' Them ere ead errant 
Nps bos Peet obi de treet beter ey rte pulp bonbened eeea'h ye) sraeyer ope et peace nacre’ a babe tere canbyh 
hod ave brad ne awe 4 aeener r sa otoh Shave Techehieseery erttt da ght et breepessene rere berets 
pans gtapr meryeet ee tas rete ad edrd ed sont Pa eee Pee 1H) are drvapeoty pope perarent er Sah pepetersereny a paprewerecerereeyriry 1. 
rr cehtarrarauced peceennete etter oettomtarty 4 pingpecscaserta cn prent poseiesies pers LT eee ra cataepes eet 
gel edrberrp eben’ eigeredee wee verge bidet ey eh etet: 4 oH of a paren werneasntin Seliry 
Tpestbeded ye ard teat sehen a rade bra nme peer eee Terie Tarapanseansncpeeh cen pre siren ene yit a eer se 
pod ep meyeben ee sued Myatt chaps caanecttregepeeti tt tee Seba be bebe +404 Hepeeen re Joe mb bettie te pnsaben tei bepeaeeeenenenantiner lees ryt tt 
eererersi Se INTL baabryeneteed fr eet de pep ae epee rt ree ar rr Hebe py nes bette ry 4 nsede Aeaeberaante ert ene yrrere pene: fret’ & 
eee obpeabe v4 ted eet’ pepe aastenneti Teh leerrn hat ebabner tet ee neens te Pabbbegegnte ie rere ater 
eer ae prenheenyey Aap eatieescedpar eed sel ath hognees geatiene Pt pn ipe stein 
{Teer eau ener more ier tee os Salona aoveeareetebree sett 
L piel Aoamenanent eet bpent yt Peebe see riat 
Perret ataberrelerte st irerie rede pRiraria inert ibenveepessore sieht en 
‘LL « "ie Lae _ + oeaet te reebeneyerrp may 
o vhereeet S88) rhe) aseesgeysiree veaiere| 


oer rps | 
ae ae 


Seat eee 


wees. 


as 
ou 


4 


Sea 


RECORDS 


of the 


INDIAN MUSEUM 


(A JOURNAL OF INDIAN ZOOLOGY ) 
Vol. IV, 1910—1912. 


EDITED BY 


THE SUPERINTENDENT 
OF THE 


INDIAN MUSEUM. 


Calcutta : 
PUBLISHED BY ORDER OF THE TRUSTEES OF THE INDIAN MUSEUM, 
BAPTIST MISSION PRESS. 


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= 


CONTENTS. 


Ay 


ee 


Vv 
No. I, FEBRUARY, I9QI0. 


Second Report on the collection of Culicidae in the Indian 
Museum, with descriptions of new genera and species 


Nos. II—III, Marcu, Ig10. 


The Indian species of Papataci Fly (Phlebotomus) 
Taxonomic values in Culicidae 


No: LV; JULY, IQrOo: 


Revision of the Oriental Bloodsucking Muscidae (Stomoxtnae, 
Philaematomyia, Aust., and Pristirhynchomyta, gen. nov.) 


No. V, NOVEMBER, IQIO. 


A new arrangement of the Indian Anophelinae 


No. VI, MARCH, IQII. 


A revision of the species of Tabanus from the Oriental Region 
including notes on species from surrounding countries 


No: VIE JUNE; Torr: 
New Oriental Nemocera aA on is ie 
Miscellanea (pp. 317—320) :— 


Synonymy in Corethrinae 
Further notes on Indian Pilenotare: 


Nos. VIII—IX, DECEMBER, IQII. 


A revision of the Oriental species of the genera of the Family 
Tabanidae other than Tabanus ae at 56 


Contributions to the Fauna of Yunnan— 
Part VII.—Tabanidae 


No. X, MARCH, 1912. 


Annotated Catalogue of Oriental Culicidae—Supplement 


Page 


i 


95 


PEE 


321 


401 


493 


esc 
CAG 
atl : 
y/ ak ; a 
r/R) \o 
dS WONG 
eb tae me 4\6 
Tp Sher ee epe } 
Me a RE 
A a orl lis 
y Rone /Ke 
\. i: e Ne ao < ae 
SL See 
ic as : tI a A: 
Py, 
£7ert® 
& 
= 
se 
a 
ne mf i 
‘ 
4 


es 


- 


Uf cle “hae ioe hs 


+: 


a 7 ~~ 
> + 


atte 
Hina 9 


= re 1 On ae ia Ve ie ee 


' . ; . 
* 
- 
ga. a & 
we ” 
= Feo 
in 
a3 


j,, Piel eee [aiid in ‘sel 


no 
= 
: ~s 
; 
Ras 
ome ys 


ue ortere ae ce 


“Enel | ik ‘fey ae, z 


ihe ‘eran? yas Nae ey 
ae aa ca nef! 


Bart OL ake 


ABs I fi cee fad iets way ate 


ul 


<e 


tones 


oh es 


, ean ieeac? pPeaeants 
st Mat BS vis te ain ‘Pee FOP Ra 2) 


au welt af Poor, Jeane Hit Wie ; 
aetna SHE SERS 
Basti ae < ith iad 2s wih oF 

eh Paar ba ; sails: 


AE Fie : 


EIS) Ob} PE ATES: 


——>-— 


Plates I—III (Culicidae) 

Plates IV—VI (Phlebotomi) 
Plates VII—VIII (Muscidae) 
Plates IX—XII (Anophelinae) .. 
Plates XIII—XIV (Tabanidae) 
Plates XV—XVIII (Tabanidae) 


Follow page 
34 
52 
o4 
110 
258 


402 


Nt ie 
cee 
aA ah 
{ 


ESE OF AULHORS: 


Annandale, N., D.Sc. 


Brunetti, EF. .. 


James, Major S. P., M.D. 


Ricardo, Gertrude 


Theobald, F. V., M.A. 


The Indian species of Papataci Fly (P/le- 
botomus), p. 35-—Further notes on 
Indian Phlebotomi, p. 319. 


Taxonomic values in Culicidae, p. 53.— 
Revision of the Oriental Bloodsucking 
Muscidae, p. 59.—New Oriental Nemo- 
cera, p. 259.—Synonymy in Core- 
thrinae, p. 317.—Annotated Catalogue 
of Oriental Culicidae, p. 403. 


A new arrangement of the Indian Ano- 
phelinae, p. 95. 


A revision of the species of Tabanus 
from the Oriental Region, including 
notes on species from surrounding 
countries, p. 111 —A revision of the 
Oriental species of the genera of the 
Family Tabanidae other than 7abanus, 
p. 321.—Contribution to the Fauna of 
Yunnan, Pt. VII, Tabanidae, p. 4or. 

Second Report on the collection of Culici- 
dae in the Indian Museum, with descrip- 
tions of new genera and species, p. I. 


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INDEX. 


——>-— 


[N.B.—An asterisk (*) preceding a line denotes a new variety or subspecies; a 
_ dagger (}) indicates a new species; and a double-dagger ({), a new genus; 


synonyms are priuted in Italics. } 


A 
Page 
Acalleomyia 3 487, 490, 491 
obscura ee AG 
**obscurus’’ 49! 
Acanthocera Ae Sees OE 
Aedeoniyia Be 487, 488 
catasticta ss 488 
squamipenna 7 405 
‘*“squammipenna’’ 26, 488 
‘“ squamipenuis ”’ 488 
Aedeomyinae 56, 413, 450, 
453, 486, 487 
Aedinae 407, 409, 450, 
485, 486, 487, 
504 
Aedes 54, 439, 487, 488, 480, 491 
butleri aa .. 489 | 
malaya 490 
nigrescens 490 
Aedimyia Fc e- 440) 
Aioretomyia 487, 489 
aedes 490 
ostentatio 490 
perdita 490 
singu'aris 490 
taeniata 490 
varietas an teh 
Aldvichia 95, 100, 106, 410, 433 
ervor 5S 100, 106, 407 
Aldrichine}la 410, 433 
error 433 
Amauromyia 450 
Ancylorhynchus 410 
Anisocheleomyia = 493 
albitarsis 494 
alboannulata 493 
Aukylorhynchus e A0 
Anopheles 54, 95—97, 106, 411, 412, | 
; 415 —417, 435, 517 
aconita.. ao bis 
aitkenii 96, 106, 415 
arabiensis 434 
barianensis 415 
bifurcatus 97 
cohaesa 415 
costalis 2 AZI 
culiciformis 434 
deceptor 434 
dthali 415, 419 
elegans 417 


Page 
Anopheles formosaensis 415, 416 
formosus .. 416 


96, 106, 416, 417 


gigas 

immaculatus 96, 106, 416 
indiensts si A2S 
jesoensis .. azo 
lindesayi 96, 106, 416, 417 


* var. maculata 1, 106, 416 


maculipennis 97, 407 
nigerrimus v5 425 
perplexeus a 403 
pictus 419, 434 
plumiger | 14.26 
pseudopictus 434 
punctulata se 452 
simlensis .. 416, 417 
subpictus . 434 
tesellaium 432 
treacherii 417 
vincenti 434 
wellcomei 417, 434 
Anophelina 410, 419, 433 


Anophelinae 55, 56, 95—109, 405—407, 
409, 412—415, 440 


Anthomyidae 56, 59 
Anthrax ss : 57 
tApistomyia trilineata 315 
Armiger a 441 
Armigeres AE 441 
Panalectoros 442 
Arribalzagia 415, 431 
Asilus 57 
Asiphonatae 55 
Asiphoniata ar re 55 
Atylotus II2)5 133) 135045, 
153, 159, 173, 184, 

ICO, 194, 195, 197, 

200, 209, 21T 212, 

217, 229, 220, 236, 

237, 244, 246—248 

251 

andamantcus 218 
bituberculatus 134 

(?) contcus 153, 154 
ditaeniatus 248 
flaviventris 214 

fulvus 133, 248 

fuscipes neyrade 
achrymans 156, 157 
latistriatus. . ao Pade 


433 


il 


Page 

Calvertina 410, 428 
lineata enn 4 333 
Calvertius a 410, 433 
Carolia a 410 
Carollia oP a5 | chit) 
Carrollia - 12, 14, 410, 454 
Cecidomyia she 57 
Cellia 54,95, 96, 99, 
100, IOI 102, 

105, 106, 107, 

428, 431 

flava syed PASI 
kochi 55 ZIT 
pulcherrima 3, 99, 102, 106, 

407 , 432 

punctulata eA G2 
Chaetomyia 452, 453 
Chagasia 2a 415, 433 
Chaoborus 317, 318, 504, 505 
antisepticus a0. BY 

asiatica 504, 505 

cornfordi .. SSO 
crystallina BZ ibs. 
manilensis £05 
plumicornis 317), 318 
punctipennis oe gy 
Chironomidae : 57 
Chironomus 57 
Christya 415 
{Christophersia 102, 103, 105, 


106, 107, 407, 

428, 431 
100, 102, 103, 
106, 407, 431 


halli 


Page 
Atylotus macer ae 150, 152 
microloma .. 152 
nigromaculatus snl S 4. 
phyrrhoceras 238, 239 
rufidens 244, 245 
Tusticus 248, 252, 253 
B 
Banksiella ie AIX AG, 
luteolateralis B47 
Bdellolarynx ae 61, 62 | 
sanguinolentus 62 
Bellardia 112, 149, 202 
SinztCUs 167, 170 
Bibio 273—28c 
tabdominalis 272270 
yapproximatus 274, 277 
bicolor 273 
clavipes 280 
}tdefectus 274, 280 
fdiscalis 273 27.3) 
jfuscitibia 273;.279 
thortulanoides 27:3 0274 | 
hortulanus ae M275 
johannis 273, 278 
laniger 3a) ze) 
marci ae 274—278 
obscuripennis 273—276, 278 | 
};proximus 274, 275 
trufifemur 27/3 3279 
vatripes 280 
venosus 277, 278 
Bibionidae 269 
Binotia 492 
Bironeila gracilis ae AS3 
Blanchardiomyia 440, 441 
apicalis 441 
aureolineata 441 | 
fusca 44r | 
joloensis 441 | 
jugraensis 441 
obturbans 441, 442 
panalectros 441, 442 
{Blepharocera indica 316 
Blepharoceridae 315 
{Bolbodeomyia a 31, 492 
ycomplex 31, 492 
Bolbodimyia ; 492 
Bombylidae 433 
Bombylius 57 
Brevipalpi ae 55 
Brevirhynchus ; 8, 442, 444 
fannulipalpis 6, 442 
fapicalis 7> 443 
magnus 6. 443 | 
Brunettia 289, 290, 291, 310, 311 | 
superstes 290, 310, 311 
travancorica 310—312 | 
C 
410, 433 
Calvertia 410 | 
lineata 


Chrysoconopas aurites sien A'S2 
Chrysoconops ake 479, 480 
aurites > W480 
brevicellulus 481 

conopas 481 

“*conopas’’ 481 

ochraceus 481 

pygmaeus 25, 481 

Chrysops 57, 365, 374s 377, 378, 401 
albicinctus .. eo, 

alter 374, 376, 387 
bifasciata 374, 377 378, 

380, 382 

cincta 374, 376, 393 
clavicrus 374, 393, 394 
costatus 22893378 
tdesignata 374, 376, 383, 401 

dispar 374 375; 378, 

380, 381, 382, 

386, 3900, 396 

fasciata 375, 376, 390, 

391, 393, 394 

fixissima 375, 376, 392, 393 


flaviventris 375, 3773 390 
flavocincta .. 375, 3775 395 
impar 375, 380, 382 
indiana 375, 376, 390 
avanensis 375, 378, 379 
ligata 375, 380, 381 
lunata : Sor. BS} 
manilensis .. 375» 377 
mlokosiewiczi 375, 378, 379, 

4OI 


Page 
Chrysops parallelus .. sO 
pellucidus .. 375, 376, 377, 
385, 386, 388 
tufitarsis 375, 376, 384, 394 
semicircula 375, 380, 381 
signifer 375, 376, 393, 394 
sinensis 375, 370, 388 
stimulans 375, 370, 389 
Siviata 375, 378, 379, 401 
terminalis 375, 380, 381 
translucens Bris oe 385, 
; 388, 396 
unizonata O75. ie 393 
punctifera .. 390 
flavipes 397 
Perea. 397 
Chrysozona 321 
Coelodiazesis barberi_ =. 405 
Colonemyia se 488, 498, 499 
caeruleocephala a 49D 
hybrida 498 
mendacis 498 
Conopomyia 453 
aurea 453 
hybrida . 453 
metallica 454 
Conops 63 
Corethra . 54, 56, E hive 318, 405, 
504, 505 
cinctipes 317, 318 
culiciformis 317, 318 
effoetus Ehily75 Shite: 
fusca ESL, 
pallida 317, 318 
plumicornts 8178, Sr 
punctipennis.. ees 
velutina 317, 318 
Corethrella brakeleyi Se Silks 
Corethridae 56, 413 
Corethrinae 55, 56, 317, 318, 409, 
411, 504 | 
Corizoneura 364, 365, 307 | 
longirostris 365, 367 
*subsp. varipes 367, 369 | 
rufa se S158, 
taprobanes 307 
Crematogaster difformis 504 
Culex I, 19, 31, 39, 40, 41, 43, 
54, 56, 57, 95, 410, 439, 
440, 445, 446, 459, 460, 

‘ 465, 497, 471, 477: 489 
albolineatus. 496 
albopictus 9 
albopleura 466 | 
albus 466 
ais)... 466 
angulatus NK 16/1406) | 
annulatus ai 412, 459 | 
annuliferus 466, 471 
annulioris ee 400 
annulus 466 | 
anxtfer 468 | 
arabiensis 4607 
argentinotus 497 
auratus 467 
aureopunctis 517 
aureostriatus 467, 471 


Page 

Culex biro 407 
caecus 467 
cantans 467 
christophersi 467 
concolor 20, 411, 468 
fatigans 5» 17, 406, 411, 442, 
468, 469, 475, 476, 483 

var. trilineatus 469 
flavifrons 4607 
foochowensis 469 
fragilis 469 
fuscanus Ae w= 469 
fuscocephalus .. 17, 469 
gelidus : 19 402 
graminis ss oc AZO 
enophodes 470 
halifaxii 470 
hirsuteron ve wo 
impellens 170574015 470 
imprimens 3. 47 
infula t..1 470 
japonicus : 471, 474 
var, ceylonica ES AE 
ylongifurcatas 19, 471, 474 
longipalpis A. 
longipes 471, 472 
loricatus ier AGT 
ludlowi 406, 471 
luteola So) 2G fp 
luteolateralis are, ATT 
macr pus 471, 472 
maculatus ; Be wiley, 
mediclineatus .. dg) Aye 
microannulatus 411, 472 
mMimeticus ‘ .. 9, 472 
minimus 472 
minor : 473 
multimaculosus 473 
nigricephalus 473 
occidentalis 407 
pallidostriatus .. 473 
pallidothorax ee 
;parascelos 18, 473 
perplexus = 5) eAt78 
tpettigrewii T5474 
pipiens 457, 469, 487 
pseudolongifurcatus 471, 474 
pseudostenoetrus an CATA 
pulchriventer 474 
pullus =» 474 
pungens 468, 469 
quasipipiens Me ATA. 
quasiunivittatus 474 
reesli 474 
rizali 474 
volanat 472 
sepositus meg GAs 
sericeus 411, 475 
sitiens pa PAs 
stenoetrus 475 
taytayensis ea 75 
tigripes 19, 411, 475 
tipuliformis sre Wn 
trilineatus 476 
trimaculatus 476 
tritaeniorhynchus 476 
uncus : 476 


Page 
Culex uniformis 476 
univittatus = 2470 
vagans 14, 476 
ventralis 442 
viridis 476 
viridiventer 476 
vishuui en 470 
Culicada 54, 461 
eruthrosops .. ser 402 
minuta ae O2 
nipponii 21, 462 
suknaensis 21, 462 
Culicales 409 439 
Culicidae cc 33, 40, 53—58, 


403—406, 408—414, 435, 504 


Culicimorphae 55 
Culicina eer EO 
Culicinae yy SO 405, 407, 409, 413, 
439, 440, 450, 485, 486, 
cee 504 
Culicini ac ake 54 
Culiciomyia ac 456, 474 
alis i 457 
annulata .. 457 
annuloabdominalis 457 
ceylonica.. 457 
inornata . 457 
minutissima 457 
nigerrima 457 
pulla 457 
Culiseta .. 54 
Cyathomyia 517 
jenseni .. 517 
Cyclolepidopteron grabhamii 415 
Cyrtoneura stabulans 67 
D 
Dactylomyia ceylonica 517 
Daniela 458 
Danielsia albotaeniata 458 
Dasymyia fusca 453 
Dasyneura 57 
Deinoceratinae 413 
Deinocerites ae oe) A S7 
Dendromyia 54, 5OI, 502, 503 
achaetae .. a5 O83 
aureochaeta 503 
communis 503 
scintillans 503 
Dendromyinae 410 
Desvoidea ae ot 6 
fapicalis .. sie 5 
obturbans.. a 4 
panalectros 5, 468 | 
“* Desvoidia ’’ ye Ct 
Desvoidya 440, 441, 453 
obturbans ; oo Chiat 
Diachlorus 321, 361, 303; 364 
flavipennis 364 
Diatomineura 364, 365 
Dichelacera re Or 
Dilophus graciosus - 280 
Dinoceratinae s« 409 
Diplonema 38> 41, 03L0 
Diptera 40, 42, 53, 58 


Ditylomyta 


379, 371 | 


iv 


Page 
Dixa a 54, 575 259, 264—269 
tbifasciata 265, 269 
fbistriata 265, 268, 269 
maculata e200 
fmaculipennis 265, 266 
fmoutana 264, 265 
tochrilineata 265, 267 
Dixidae 264 
Drymeia .. 56 
E 
Eatonisca sks AG 38 
Ectenopsis 373 
Ekrinomyia aureostriata 440 
Epialur.i Ss 409 
Erephopsis Be 365 
‘* Etorilepidomyia ’’.. 440 
Etiorleptiomyia ne 484 
completiva 484 
luzonensis -- 484 
‘« Etorlepitiomyia *’ 440, 453, 484 
Eucorethra underwoodi Jas ES 
Euculicidae 55 
Exoprosopa 57 
F 
Feltidia 5 2) 8 4OE 
Ficalbia .. ae 487, 494 
longirostris Te 404 
minima - 494 
simplex : 494 
tenax =o, SiL7, 
Filaria bancrofti ag 2G 
Finlaya 485, 517 
aranetana Soy 243C 
poicilia - 486 
Finlayia 440 
G 
Gastroxides 365, 370, 372 
ater 370, 371 
ornatus Bo sf! 
Gettonomyta 459 
Goeldia te A SOW 
Grabhamia 54; 461, 489 
ambigua Se) 46 
jamaciensis 461 
ochracea .. 461 
pulcripalpis 14 
spenceri 461 
sollicitans .. 461 
taeniarostris se 401 
Grahamia 27, 489 
H 
Haemagoginae Be CE 
Haemagogus 12, 488 
Haematobia 61, 62, 63, 66, 
82, 83 
extgua ac 86 
fevox 50 So- <5: 
geniculata bo (8 


Page | 


Haematobia irritans .. ein)’. 8 
jrufipes .. 64, 65 
sanguisugens 64, 65 

serrata GIONS 

stimulans 59, 63, 64, 65 

tibialis em 63 
Haematobosca 59, 60, 61, 62 
atripalpis Wea O2 
Haematopota 32I—360I1, 375, 401 
fannandalei 321, 335, 337, 

oe 338 

astatica 321, 340, 341 

yassamensis 321, 343, 345 

atomaria 321, 331, 332, 

; 333 

;bilineata 321, 331, 350 

borneana 321, 329 

brevis 322, 349 

cana . 322, 330 

cilipes . 322, 339, 360 


cingalensis 322, 349, 350, 
351 


cingulata 322, 326, 327 
cordigera 322, 351, 357 
decova .. 337 
}dissimilis 322, 331 
dorsalis Dow) able y/ 
yfasciata 322, 358 
fuscifrons 322, 357 
yimmaculata 322, 359 
yinconspicua 322, 358, 
359. 360 
indiana -+ 328 
irrorata 322, 352 
javana - 321, 322, 334, 
340, 341, 344. 
349 
lata 322, 336, 338, 
339, 344, 345. 
361, 401 
+latifascia 322, 356, 358 
limbata 322, 325, 326 
lunata . 380, 381 
lunulata 322, 354, 355, 
356 
+marginata 322, 347 
nigra 322, 340, 341 
pachycera 322, 336, 337 
pallens -» 348 
pluvialis 323, 340, 341 
punctifera 322, 327 
pungens 352, 353 
roralis 322, 323, 330 
rubida 322, 327, 336, 
Pighss 337, 338, 339 
sinensis 322, 345 
singularis 322, 327. 337, 
338, 339 
tessellata 322, 348, 350 
unizonata 322, 332 
tvalidicornis 322, 333, 335, 
337, 338 
_ Vittata.. aisle G7 
Harpagomyia 4445 504 
: splendens +» 504 
Heliconia brasiliensis 


404,442, 449 


‘* Hebotomus ’’ minutus 320 


Page 
Heinzmannia * 502, 503 
Heptaphlebomyinae .. 54, 409 
Hodgesia .. 30 487, 500 
malayi iSO 
quasisanguinae 501 
sanguinea 501 
Howardia ns i .423 
Howardina 54, 423, 485, 490 
chrysolineata 450 
greeni 450 
himalay.ana 456 
‘* Hulecoeteomyia’’ .. 455 
Hulecoetomyia S 455 
fluviatilis 455 
jugraensis 455 
pseudotaeniata 455 
trilineata I4, 455 
K 
Kerteszia mcgregori .. 431 
Kingia annandalei 443 
i 
Lamellibranchiata 55 
Laphria 57 
Laverania aoe 
Leicesteria ¥ 453, 454, 485 
anuulitarsis Som 454 
apicalis 454 
cingulata.. 454 
‘* dolicocephala "’ 455 
dolichocephala 455 
longipalpis ee 455 
Leicesteriomyia 452, 453 
flava .. Ee a3 
Lemna arrhiza 404 
minor st .. ~404 
Lepidotomyia 458, 459, 486, 487 
alboscutellata Be. Ke 
magna ., 458, 459 
taeniata he 450 
Leptosomatomyia lateralis . 488 
Leucomyia 411, 462, 463, 478, 480 
gelida ; sr 20 
var, cuneata : 20 
var. sinensis 20, 463 
sinensis ma 20 
Limatinae 407, 410 
Limatus .. yh cee 
Limnobiinae 35 
Longipalpi 55 
‘* Lophocelomyia *’ 427 
Lophoceratomyia -. 463 
}bicornuta 25, 463 
brevipalpis 25, 463 
eminentia sh 403 
fraudatrix 25, 26, 463 
mamunilifer 463 
minor 463 
niger 463 
rubithoracis 463 
sylvestris 465 
taeniata 465 
uniformis 465 
variata 465 


Page 
Lophomyia asiatica 427 
Lophoscelomyia : 424, 427 | 
asiatica 427 
Tudlowia. . 493 
chamberlaini. 493 | 
minima : 28493 
Lyperosia 60, 61, 66, 82, 83 
exigua 6o, 83, 86, 89 
;flavohirta 83, 89 
irritans 59, 83, 87, 89 
minuta 60, 83, 84, 88 
sevvata os 83 
titillans 83 
M 
Malaya .. re 501, 503 
genurostris 503 
Mansonia.. 440, 481, 483 
annulata aA S2 
annulifera Pease 
annulipes ae 411, 482, 483 
anopheloides .. 412 
arabica 482 
chrysogona 482 
nevo Be acl? 
seguini : 482, 483 
septempunctata Bee ksi} 
uniformis 22, 406, 482, 483 
Mansonioides 440, 482, 483 
annulifera 22, 483 
septemguttata 483 | 
Megalorhini Aes 409 
Megalorhinina 410 
Megarhinae 407, 409 
Megarhininae 413, 434 
Megarhinus Die ee 435 54375430 | 
ambotnensis 435 
ferox 438 
gilesit a eA O 
tmMisevicors 438, 439 
lewaldti 435 | 
minimus 435 
splendens 435 
subultfer 430 | 
Melanoconion 54, 484, 480, 490 
juxtapallidiceps 484 | 
ornatus 484. 
pallidiceps 484 
uniformis 484 
Mesomyia 373 
Metanototricha 409 
Metantotrichina ; 410 
Micrococcus melitensis Te eASO | 
Mimomyia 453, 487, 488, 492, 493 | 
chamberlaint 492, 493 
minuta 30, 492 
Mochlonyx 317, 318, 405 
effoetus joe» UY) 
velutina .. SS HRY/ 
Musca ‘ P 59, 91 
corvina ate aie 82 
domestica ee, 90 
pungens : 68 
Muscidae 56,5 59—93 
Mucidus 439, 440 
laniger 440 


vi 


| Page 
| Mucidus mucidus -- 440 
scatophagoides 4, 440 

| Muscinae as 56, 83, 493 
Mycetophila ; 57 
Mycteromyia : 365, 373 
nigrifacies Sm S15) 

_ Myzomyia 54,95, 97, 98, Ior, 
103—107, 415—418, 

421, 422, 517 

albirostris 418, 421 

annularis .. 418 

aurirostris 517 

azriki 418 
barbirostris 406 


christophersi 2, 98, 418, 420 
culicifacies 2), 190,975 98, 
106, 404, 407, 
418, 420, 422 


var. punjabensis 98, 106 
deceptor 418, 434 
dthali ea AIO) 
elegans 106, 109, 417, 419 
fuliginosus sal AOS 
funesta 400, 419, 424 

var. umbrosa 419 

var. sub-umbrosa.. 419 
indefinita .. 406, 419 
jehafti 419 
kumasit oe AO 
leptomeres 96, 97, 98, 106, 

419, 434 

leucophyrus 417, 420 
listoni 96, 97, 98, 106, 420 
ludlowii 406, 419, 420 
pseudobarbirostris 4.06 
punctulata 420, 421, 432 
rossii a2 OO a4 OO. 
_ 419, 421, 433 

var. indefinita 419 
tessellata .. 422 
thorntoni .. Bon LI 
turkhudi .. 96—g99, 106, 

; : 404, 422, 424 
sinensis See Cig 
vanus 400 

Myzorhynuchus 95, 99, 100, 
102, 103, 105, 

106, 107, 109, 

415, 424 

albotaeniatus 424 
alboannulatus 424 
barbirostris 3, 99, 102, 
103, 106, 407, 

424—427 

indiensis 427 
minutus 425 
nigerrimus 99, 102, 103, 
106, 425 

peditaeniatus 425 
philippinensts 426 
pictus. . 427 
pseudobarbirostris 426 
pseudopictus 426 
separatus en 420 
| sinensis 4, 99, 102, 103, 
106, 411, 424, 


425, 420, 427 


vil 


Page 


427 


Myzorhynchus umbrosus : 
102, 426, 427 


vanus 
N 
Neocellia 95, 96, 100, 
IOI, 105, 106, 
107 428, 432 
dudgeoni 100, 102, 106, 432 
indica TOO; 101, 102, 
106, 407, 432 
intermedia 100, 102, 106, 432 
stephenst 102, 106, 430 
willmori 102, 106, 430 
Neomacleaya indica .. 458 
var, simplex 458 
Neomyzomyia : 109, 418 | 
elegans ICQ, 417, 419, 
423 
leucophyrus 417, 420 
** Neonyssorhynchus ’’ SOx 
Nemocera ve ae 38 
Nemopalpus st ne Ke: 
{Neostethopheles 97, 98, 103, 
104, 106, 407, 
415, 422 
altkent 98, 106, 407, 
422 
culiciformis 98, 106, 
422, 423 
immaculatus 98, 106 
tNeotobanus ss 321, 363 
yceylonicus 5) SOK) 
Nepenthes alata 501 | 
rafflesiana 438 
{Nyssomyzomyia IOI, 104, 106, 
108, 418, 420 
ludlowt AQT eA22 
punctulata 421, 422, 
432 
vosstt 106, 407, 421, 
422 | 
Nyssorhynchus 95, 96, 99, 100, 
IOI, 104, 106, 
107, 109, 415, 
423, 428, 431 
fuliginosus 3, 99, 100, 
106, 428 
var. adiei 106 
var. nagpori 106 
var, pallida 428 
indiensis 428, 429 
jamesii 3, 99, 100, 106, 
429 
karwari 99, 100, 101, 
100, 429 
leucophyrus 417 
maculatus 3, 99, 100, 
106, 407, 429 
maculipalpis 99, 100, 
IOI, 106, 
Rees, 429 
var. indiensis 106 
niivipes 429, 430 
philippinensis 426, 430 
pseudowillmori 430 
punctulata 


432 | 


Page 
Nyssorhynchus stephensi 3, 99, 106, 430 
theobaldi 99, 100, 106, 


: 430 
willmori 3, 99, 106, 430 


O 


Oculeomyia fulleri .. ae ASS 
sarawakii 485 
‘* Oculiomyia ’’ 5 485 
Odontomyia oe Nace SSIS, 
Oestridae ae 8 59 
O’ Reillia ar Sah ASA. 
Orthopodomyia 440, 485, 480 
albipes 30, 486 
}maculata 29, 486 
maculipes 486 
nigritarsis 486 

P 
Pangonia 57+ 395, 366, 367, 373 
amboinensis 305, 306, 368 
conica Pat 373 
longivostris .. 365 
rufa : 367 
Pangoninae 364, 370, 374 
Panoplites Se Pe 
{Parabrunettia 289, 290, 291 
310, 311 
falbohumeralis 311, 312 
argenteopunctata 311, 
312 
artisquamis 311, 312 
fflavicollis 311, 312, 314 
flongichaeta 311, 312, 
314 
fo-notata 311, 312. 313 
squamipennis 311 
Pardomyia aurantia. . 460 
quadripunctis 469 
Parhaematopota 231750365 
cognata 337 
{Patagiamyia 97, 98, 104, 106, 107, 
109, 407, 415, 
416, 417 
gigas 98, 106, 407 
lindesayi 99, 106 
var. maculata 106 
Pecomyia 459, 460 
caeca af 459 
maculata a 400 
Pelorempis americana 318 
Penthetria 270 
melanaspis . te e205 
Pericoma 38, 39, 41, 289, 290, 291, 
30I—310 
annandalei 301, 304 
appendiculata 289, 302, 304 
bella : 289, 302, 304, 
A 230537507 
gilvipes 303, 306, 307, 308 
yimpunctata 303, 309 
lacteitarsis 303, 306, 307, 308 
margininotata 289, 290, 297, 


302, 304—308, 311 


vill 


j 


_ © Pseudomyzomyia ”’ 


Page 
Pericoma j;metatarsalis 302, 305 
jmixta 303, 306, 307 
+proxima .. 303, 307, 308 
spinicornis 289, 290, 302, 
ae 304 
+squaminervis 301, 303 
yunicolor .. 303, 309 
Phagomyia gubernatoris 456 
Philaematomyia insignis 90, 93 
Philodendromyia barkeri 503 
Phlebotomi 20 319 
Phlebotomiella As 37 
Phlebotomus 35—52, 289, 319, 504 
angustipennis 38, 51, 52 
argentipes 37, 38, 4244, 
46, 47, 49—51, 319, 320 
var. margin- 

atus 319 
Tbabu 37, 44, 49—52, 
319, 320 
*var. niger 320 
cruciatus see ey. 

duboscqui 37, 38 | 
+himalayensis 37, 44, 50, 
51, 319 
fmajor 37, 44, 46, 47, 51, 
319, 320 
*var. grisea 320 
;malabaricus 37, 44, 48 
50, 289 
marginatus oie) SHO) 
mascittii 37, 38 
minutus. 37 


papatasi 355 375 38, aa: Si, 
52, 289, 319, 320 


perturbans 35, 38, 44, 
47—49, 289 
tipuliformis =e 37, 
vexator .. oe 37 
Phlebotominae 38, 39, 289 
Philaematomyia 59, 89, gI 
Phoniomyia 501, 502 
bimaculipes - EES OZ 
caeruleocephala ven OZ 
indica 502 
‘* Phonomyia ”’ 502 | 
Plecia 270—272 | 
yatra 271, 272 | 
fulvicollis 27 O27. 
tindica 270, 271 
melanaspis 269, 271 | 
fobscura 271 e272 
subvarians 271 
tergorata 270, 271 
thoracica 271 
$Pleciomyia 269 
melanaspis 270, 271 
Polylepidomyia argenteiventris .. 502 
Popea RI 410, 485 
lutea 485, 517 
Poppea 410, 485 
Potisa x2 Sa SSH 
{Pristirhynchomyia . 59, 91 
tlineata QI, 92 
Promachus . 57 
t{Pseudocarrollia 12, 454 


ylophoventralis 1 3, 454 


Page 

Pseudograbhamia 460, 4890 
maculata 14, 460 
tPseudograhamia 26, 460, 489 
faureoventer 27, 460, 

489 

_ Pseudohowardina chrysoscuta 456 


95, 100, IOI. 
106, 421, 422 


rossi 95, 100, 

IOI, 106 

| Pseudoskusea Ae aD 488 
mediolineata 451 

multiplex 450 

nigrotarsis 450 
Pseudostegomyia : Ags 
Pseudotheobaldia niveitaeniata . aS 
Pseudouranotaenia parangensis.. 494 
ieee -. 494 

Psorophora ; So. zelc{o) 
Psorophorinae 413 
Psychoda 38, 30, 41, 289—301, 
310—315 

albonigra 289, 292, 294, 295 
albonotata 289, 294, 298 
yalbopicta 293, 296 
yapicalis 291, 294, 301 
argenteopunctata 200573KE 
artisquamis .. 310, 311 
bengalensis 230 200.8202. 

295, 303, 308, 312, 

314 

ydecora 294, 299 
;distans es 293, 296 
distincta 289, 290, 294, 208, 

iii 

}fulvohirta 293, 297, 208 
tgeniculata 292, 294 
Thirtipennis .. 294, 300 
+maculipenuis 294, 299 
nigripennis .. 289, 290, 293, 
297—300 

yorbicularis 292, 293, 298 
squamipennis 290, 311 
ftratlsversa .. 294, 299, 300 

vittata 289, 293, 298 
Psychodidae 38, 40, 43, 57, 
289 

Psychodinae a 38, 289 
Ptychopterinae Ac bo GS 
Pyretophorus 95; 97, 99, 104, 106, 
107, 418, 423, 

, 433 

costalis .. 99, 407 

elegans 96, 97, 99, 

106, 417, 423 

freerae ao) 2S 
jeyporiensis 96, 97, 99, 

106 

minimus Bo 283 
nigrifasciatus 096,97, 99, 

106, 424 

nursei 96, 97,99, 106, 

424 

palestinensis 99, 407 
philippinensts 430 

punctulata 96, 97, 99, 106 

watsonii ae eezd 


ix 


Q 


Page 

Quasistegomyia SPs 443, 447 

gardneri 443 

R 

Rachionotomyia ceylonensis 455 

Radioculex clavipalpus 26, 465 

Ramcia n 504, 5C5 

inepta Men SOS 

Reedomyia 458, 486, 487 
ss alboscutella 22 eee S701) 
alboscutellata 487 | 

lowisii 5 487 
niveoscutellata 487 | 
pampangensis ee 474 || 
Rhinomyza Bs 365, 369 | 

fusca e309 

Rhynchomyia 493 

Rhyphidae 259 

Rhyphus 259—263 

Tdistinctus 260, 262, 263, 264 

tdivisus 269, 263, 264 

fenestralis 259, 261, 264 

*var. indicus .. 259, 

261 

indicus so. AOE 

maculipennis 259, 260, 261 

+pulchricornis 259, 260 

punctatus 259, 262 

Runchomyia 493, 501 

philippinensis 403 

Ss 

Sabethes .. 501 

Sabethinae 405 

Sabethini 54 

Sabethoides oe SOL 

Sayomyta 317, 318, 504, 505 

Scatopse brunnescens a 205 

fnigronitida .. 281 

Sciara era 57 

Scutomyia 5 448, 452, 453 

albolineata so Lge 

nivea : 452 

notoscripta 452 

subsp. samar- 

ensis 452 

sugens 452 

treubi ees Ly, 

Silvius 365, 373 

tindianus pees 74 

vituli ae ve 3a 

Simulium 56, 282—288, 445 

taureohirtum 282, 286, 287 

tgriseifrons 282, 285 

tgrisescens 282, 283, 284 

indicum 282, 284, 286, 288 

tmetatarsalis 282, 284, 285 

yrufibasis 282, 285 

trufithorax swaneoe 

tsenilis 282, 288 
Siphona .. d 66 | 

Siphonatae 55 

Siphoniata 55 


Page 

Skeiromyia 488, 498, 500 
fusca a 500 

Skusea .. 450, 488 
culiciformis eee AST 
diurna 33, 451 
funerea Ae 45S 

var. ornata 451 
mediofasciata . ae AGI 
ypseudodiurna .. 32, 455 
pseudomediofasciata 451 
yuniformis 33,451 
fSquamomyia . 28, 489 
tinornata. 28, 489 

Stegomyia 10, 12, 439, 443, 444, 
447, 448, 453, 456 

yalbipes ‘ II, 444 
albolateralis 444 
albopictus .. 449 

amesii 444 
yannandalei IO, 444 
annulirostris 444 
argenteomaculata 444 
assamensis 12, 445 
aurostriata 445 
brevipalpis -. 445 

calopus 406, 446 

crassipes 445 

desmotes .. 445 
dissimilis .. en 445 

fasciata 8, 406, 411, 436, 

446, 449 

var. luciensis 446 
frater a= 446 

fusca 447» 491 

gavdnert 447 

gracilis 447 

imitator 447 
leucomeres 447 
medio-punctata 447 
microptera 447 
;minutissima Q;, 11; 4, 448 

nigritia 517 

nivea 5 448 
periskelata 448 

perplexa 448 
pipersalata 448 
pseudonivea 448 
punctolateralis 448 
scutellaris 8,9, 446, 448, 

449 

subsp. sama- 

rensis 449 

sexlineata .. 449 
striocrura .. 449 

thomsoni .. 450 
tripunctata 450 

w-alba : 450 
Stethomyia 95°97 106, 415 
culiciformis , 106, 423 

fragilis we 423 

pallida Ee oy ee, 
Stomoxinae ae 59, 60 
Stomoxys 56, 59. 61, 63, 66, 67, 
81, 82, 83 

aculeata +E a 69 
auvifactes .. siete OG 
bengalensis 68, 71, 79, 80 


Stomoxys brunnipes.. 


38 | 


calcitrans 59, 63, 66, 67, 
68, 73—76, 79, 
Si noes, 
chrysocephala - 69 
clavipennis .. 69 
cunctans 69 
dacnusa 81 
diva 69 
flavescens 69 
indica -59, 68, 77, 78, 79, 80 
infesta ‘ : 69 
inimica 69, 72 
tyvitans Ae 66 
? libatrix 6on72 4\ 
limbata (?) 68,77, 78, 79, 80 | 
minuta ae 69 | 
nigra ei 79 | 
toblongopunctata 67, 73 
omega : . 81 
plurinotata.. 677473 
praecox 50 69 
pulla 59, 68, 80, 81 
pungens ne 69 
pusilla 68, 80, 81 
vubrifrons : 69 
sellata : 74 
sitiens 68, 75, 76 
stimulans 3 66 
sugillatrix 69 
tesseliata : 68 
}triangularis. . 68, 77 
vulnervans ae 69 
Stygeromyia : 59, O1, 62, 82 
maculosa. . Si 82 | 
Sycorax .. 
Tr 
Tabanidae 112, 321, 401, 492 
Tabaninae af 321 
Tabanus 58, I11I—258, 321, 361, 
363, 364, 380, 401 
abazus st 240, 247 
abbreviatus Life SER Deisie 
232, 235 
abscondens 118, 161, 231, 235 
t+adjacens 247, 248 
administrans 167, 168, 171, 
231 
agricola 133, 135, 231, 248 
albilateralis .. 128, 23 
albilinea So AS 
albimedius TLS, U5L. 154, 
155, 156, 157, 231, 
233, 2345 235 
albocostatus 146, 190, 231 
talbofasciatus 116, 146, 231, 
235 
alboscutatus 122, 9h30 92S Ls 
235 
albulus 136, 231 
alexandrinus So DG, 
amaenus T1Q, dO7, 169, 231, 
235, 236 
andamantcus. . 217, 231 


Page 

| Tabanus angusticornis 236, 237 
angustus 212, 231 
annamitus 117, 149, 231, 235 
apicalis 182, 216, 231 
apricus 246, 247 
ardens 2003. 2240231. 235 
aspahanicus .. 247, 255 
assamensis 194, 231 
astur P wt 2A8 
fatrohirtus 125, 209, 231, 235 
auricinctus 22156289 
auriflamma .. 117, 143, 148, 
231, 235 

tauristriatus .. 118, 149, 161, 
231, 235 

aurotestaceus II9, 149, 163, 
231, 235 

autumunalis E55), L685 LOO, 
170, 171, 247, 253 

barbarus : Powis G2{0) 
basalis 116; 147, TOOs230% 
235 

bicallosus Il4, 229, 231, 234 
tbicinctus Mos 1A YA. TR EH ic 
234, 235 

bifarius 247, 254 
bipunctatus .. Sens 4: 
birmanicus 124, 200, 201, 
231, 235 

bituberculatus 231, 236 
borniensis 126, 216, 231, 235 
bovinus 185, 245 
bromius 246, 247 
brunneus I Zi elO7eL Os 
192, 237, 235 

;brunnipennis 158, 160, 231, 
235 

bubali 149, 166, 231, 235 
bucolicus 136, 187, 231 
buddha 236, 238, 239 
caerulescens 124, 200, 203, 
205, 225, 232, 

235 

calidus 156, 15751230 
canipalpis 246, 247 
ceylonicus 127, 129, 222, 223, 
231, 233, 235 

chinensis 152,231 
chrysurus 23612375 23951239 
cinctus mi Soe olay. 
cinevascens 215, 219, 220, 231 
clausacella 167, 171, 231, 236 
conformis : ese 
confuctus 236, 244, 245 
conicus 3 232 
consanguineus 123, 197, 232, 
235 

consocius ; 195, 196, 232 
cordiger 236, 239, 244, 247, 
254 

costalis 153, 198, 232 
crassus 122, 148, 189, 
232, 235, 236 

cyanops se 25 any, 
}cyprianus 247, 248 
decorus bo Sey, 
tdiscrepans 122, 185, 232, 235 


x1 


Page 

Tabanus tdissimilis .. 121, 180, 181, 
232, 235 

ditaeniatus .. 115, 133, 134, 
135, 232, 234, 

235, 236 

tdiversifrons 1260) 2135 204, 
2305, 232, 235 

dives 186, 206, 207, 232 
dorsiger TOS 
dorsilinea PEO Wyo) 232 
eggeri 187, 247, 249 
equestris 139, 140, 232 
erythrocephalus 7 530 
explicatus 122, I9I1, 232, 235 
factiosus T2017 Om 2325 235 
felderi [OF TSO. 230, 2325 
236 

fevvidus 6 232 
finalis : 182 
{WHAVACINCEUSE ue 1LS, 130; 232.5 
oe 234 
tflavissimus 125, 207, 232, 235 
flaviventris 126,213), 2145 
282.7235,1230 

+flavothorax 124, 201, 232, 
Ee 235 
flexilis 116, 140, 141, 143, 
ae 232, 233 
+formosiensis T2720 2325 
236 

fulvicornis .. 246, 247 
fulvimedius .. 123, 197, 198, 
232, 233, 235, 

236 

fulvissimus U2he 205.207 
232, 235 

fulvus TZ Ube sg) LGA), 
231, 232, 235, 236 

fumifer LUZ LT Ov ani7i3s 


174, 175, 177, 179, 
180, 185, 186, 208, 


217, 232, 235 


fumipennis 200, 225, 232, 235 
fuscicauda 126, 214, 217, 231 
: 232, 235 | 
fiuscicornis 116, 144, 232, 235 | 
tfuscomaculatus 122, 183 | 
geniculatus TSS 200) 2325235 
gigas ; ene 2A 
glaber 161, 247, 249 
graecus 5¢ nee 2A7. || 
gratus : 129 
guineensis SILOS 
hilaris 117, 153, 164, 232, 235 
hirticeps ae se 230 
thirtipalpis 114, 129, 232, 234 
thirtistriatus FES) 1585 232, 

; 235 
hirtus 127, 227, 232 
hoang 134, 236 
humilis be cep ets) 
hybridus 118, 149, 162, 163, 

i ; 232), 235 
ignobilis 121, 180, 181, 232, 
SAG 235 | 
imimanis 149, 166, 200, 


232, 235 | 


Tabanus incultus 


Page 
188, 199, 232, 235 


tindianus L2OpD7Ss Los. 2325 
233, 235, 236 

indicus ae 198, 232 
infuscatus 246, 247 
tinobservatus 124, 204, 205, 
226, 232, 235 

inscitus aie ee 232 
intermedius 187, 249 
tnternus V7.3 PL 775 2325 235 
ispahanicus .. 247, 255 
ixion 200.) 220), 233), 235 
japonicus : a5 ele 
javanus EL Ges on 2990) 255 
joidus 125. 147,209, 2205 
212, 233, 235 

jucundus 119, 164, 233, 235 
justorius T2a, 1925 193, 233; 
235 

khasiensis 123, 193, 2335 255 
lacvyymans sa 233 
lama Be ee {S) 
laotianus 200. 212, 220; 
233, 235 

leucocnematus II5, 130—132, 
233, 234 

fleucohirtus 125, 208, 209, 
233, 235 

leucopogon 122, 184, 188, 
233, 235 

leucopterus .. 40 wists: 
leucosparsus 1675 187, 233) 
235 

lineola 198 
longicornis .. 5. mug yS) 
lunatus ve 5 Beal 
macey A 36 233 
;malayensis L2G) D7 sy Diyos 
: 233, 235 
mandarinus 119, 168, 170, 
233, 234, 235, 

236, 237 

manilensts 150, 152, 233 
megalops 150, 152, 233 
melanognathus 200, 226, 233, 
235 

melanopygatus 197, 233 
mentitus (?) 17.17 Oneas 
minimus 221, 233 
}miyajima 247, 250 
montlifer 159, 173, 233 
monotaeniatus 118, 159, 
160, 233, 235, 401 

montanus 246, 247 
miihlfeldi ey yy, 
{negativus Dine 1375 2535 
235 

nemocallosus Trl 372535 
234 

nephodes 116, 145, 233, 
235 

nexus 121, I81', 233, 235 
nicobarensis T23\/191, 233; 
, 235 
nigrita ot an | 2A 
nigromaculatus 2335 235 
nigropictus 125, 210, 233, 235 


xii 


tsexcinctus 


115, 133, 234 | 


Page 
Tabanus nigrotectus 124, 202, 233, 235 
nitidulus 223, 233 
niveipalpis 246, 247 | 
¢non-optatus .. 116, 140, 233, 
235 
obconicus 126 2ihs 235 
235 
obscuratus .. 203, 205 | 
obsoletus ie eos 
occidentalis .. en LOS 
optatus 116,139, 140; 141, 
231, 232, 233, 234, 
3 235 
orientalis 188, 196, 198, 233, 
; 235 | 
orientis 123, 010512905 a19 7/5 
198, 228, 232, 233, 
235, 401 
oxyceratus 123, 184, 195, 
233, 235 
pagodinus 197, 233 
pallidepectoratus 125, 209, | 
é 233, 235 
palpalis 125), 2025231, 23351 
235 
par 206, 207 
partitus 150, 152, 233 
pauper 125, 207, 233, 235 
tperakiensis 124, 204, 233, 235 | 
perlinea 195, 199, 233 
tpersis 247, 251 
pictipennis TAO; 41415 143), | 
233 
polygonus .. 247, 252 
polyzonatus .. ee 
+pratti 116, 14353144, 145, | 
233, 235 | 
priscus L50,1157, 233 | 
puella os 136, 233 
pulchellus .. sreeea 7 a 
pulchriventris 236, 243 | 
pulverifer 247, 254 
pusillus 222, 223, 236, 242 
fputeus LIQ, 165, 234, 235 
pyvausta 188, 234 
pyrrhoceras . She OES 
pyrvhus 134, 234, 236 
quadrifarius .. 247, 254 
quinquevittatus «« 198 | 
yrarus 114, 128, 234, 235 | 
rectus : Ni ity GES || 
reducens Se Sa NG 
rubicundus 120, 106; 172, 
VOT), 4232, 234) 
; 235 
rubidus UES 5 1 LSi50050, 
cf. 157, 234, 235 
rubiginosus .. op cel 
vufidens ep e230 
rufiventris 167, 185, 234 
yufocallosus .. 150;2052, 234 
rusticus 112, 135 
sabuletorum 2A7 254 || 
sanguineus 123, 104,231, || 
Bs 234, 235, 236 
servillei 122, 188, 234, 235 | 


Page 
Tabanus tsiamensis 125, 212, 234, 235 
signatipennis 167, 236 
signifer 122, 167, 181, 182, 
234, 236 
tsignificans 122, 182, 234, 235, 
250 
simplissimus L27Qne2t 2246 
233, 234, 235 
stnicus 150, 152, 234, 236 
solstitialis 56 CAG 
tspeciosus TTS, 057523452355 
236 
spectabilis 247, 253,255 
speculum .. eas 
spoliatus : areal AL 
+stantoni 1205 57491 005220775 
208, 232, 234, 235 
striatus 117, 129, 149 
151—153, 155, 156, 
198, 231, 232, 233, 
234, 235 
+subcallosus 127, 227, 234 
+subcinerascens D2O=. 2G 
234, 235 
subhirtus 127 2TO 2) site 
} 334, 235 
sumatrensis so 234 
taeniatus .. ote 
taeniola St bo.) AS} 
tataricus 236, 242 
tenebrosus 124, 203, 234, 
2355 237, 243 
tenens aa 150, 152, 234 
(Therioplectes) 112, 113, 127, 
227, 236, 247 
albilatera- 

lis Beez 30 
hirticeps 236 
hirtus .. 228 

pulchri- 
ventris 236 

tsubcallo- 
SUS Eas see 27/6 
234 
twyvillei,. 229, 
234 
+tinctothorax 124, 202, 234, 
; 235 
triangularis Fa ley, 
tricolor ag oxy 
tyvigeminus 170, 171, 234, 237 
trigonus 237, 243, 244, 
245 
trilineatus .. AG. Heys 
tristis 119, 163, 234, 235 
trivittatus .. fetOS 
+tuberculatus 1275722052346 
: 235 
umbrinus ., ee 247, 
umbrosus 156,157, 234 
unifasciatus eee a7, 
funiformis 127, 218, 234, 235 


univentris 125, 186, 206, 207, 
232, 234, 235 
156, 157, 234 
116, 140, 141, 
143, 234, 235 


vagus 
vanderwulpi 


xa 


Page 
Tabanus varicolor 120, 172, 234; 235 
variegatus .. 172, 234 


virgo LTS 2 Oy 240-233), 
, 234 | 
vittatus .. 5A MUGS 
twyvillei .. W277 238A 
yao 236, 237, 244, 245 
yokoamensis 237, 246 


M223 On) 2375 
244, 246, 247 


Tabanus (Atylotus) 


‘Tachinidae be 441, 453, 502 
Taeniorhynchus 468, 478, 480, 481 
ager .. DOTA e 
argenteus 462, 478 
aureosquammatus 478 
aurites 479, 482 
brevicellulus en d7O 
confinnis eee AZO) 
conopas 35 lve) 
domesticus 55 2/0) 
tepidesmus 22s 2A AO 
lineatopennis 479 
tluteoabdominalis.. 23. 
479 

ochraceus ee AO 
pagei .. 78 450 
tenax.. 22, 480 
var. ocellata.. 480 
var. maculipes 480 

var. maculipes 

arabiensis 480 
whitmorei 5o EO) 
Teromyia yd 435, 439 
acaudata .. ee She) 
ater . 438 
funestus .. e435 
magnificus .. 438, 439 
quasiferox 55) ZIG) 
raris ve 439 
Theobaldia annulata 459 
spathipalpis e459 
Theobaldiomyia * 462, 478, 480 
argentea eee OZ 
gelidus 411, 462 
var. bipunctata 465 
var. cuneata 463 
sinensis : 463 
whitmoret Se lores 
Theobaldinella 54, 459 
Theobaldius te ser 4:59 
Therioplectes Piss bies 02770227. 
Dale 2I 22d 
albilateralis 235 
hirtus 229, 230, 235 
tsubcallosus 235 
twyvillei dy BBE 
Tipula .. wa 57, 318 
culactformis aa Sz 
flavicans ale 317 
pallida Ae 317 
plumicornts .. lay, 
punctipennis .. ee Sly 
Tipulinae as 55 
Tipulidae 55> 57 
‘Topomyia iz 185, 499 
argyropalpis -» 409 


argyroventralis 199, 500 


Topomyia decorabilis 
durbitans 
gracilis 
minor .. 
nigra Be 
rubithoracis 
‘* rubithorax ”’ 
tipuliformis 

Toxorhynchites 

amboinensis 
argenteotarsis 
brevipalpis 
gilesii. . 
immisericors 


436, 


inornatus 
javaensis 
leicesteri 
lewaldii 
marshalli 
*“ metallica ’’ 
metallicus 
minimus 
speciosus 
splendens 
subulifer 
Trichomyia sc 
Trichopronomyia annulata 
Trichoprosoponinae .. 
Trichorhynchomyia fuscus 
Trichovhynchus ay: 


U 


435, 436, 


Page 


499, 
437. 


4, 
437; 


410, 


499 
499 
499 
500 
500 
500 
500 
500 
439 
435 
435 
435 
435 


435; 


438 
436 
436 
437 
437 
435 
437 
437 
437 
435 
437 
437 

38 
477 
413 
477 
477 


Udenocera 32 GOls 408 
brunnea ee 302 
Uranotaenia 439, 453, 487, 493. 494, 
495. 495 
argyrotarsis OS 
ascidiicola Pee Gali, 
atra 495 
bicolor .. 495 
bimaculata 495 
bimaculialia 495 
caeruleocephala 
var. lateralis 495. 490 
campestris 495 
cancer 490 
ceylonica 496 
fusca 496 
lateralis... r 496 
longirostris 496 
lutescens 490 
maculipleura 490 
malayi we* 400 
maxima.. 495, 497 
mijcans aa) 407 
mIntmMa .. 497 
modesta or 407 
nitidoventer 193, 497 
NIVEA Was ne) 407 
nivipleura 497 
powelli - 497 
testacea,. 20, 497 
trilineata 498 
unilineata 498 


Uranotaenia unimaculata 
unimaculiala 
Uranotaeniinae 


Vv 


Veprius .. 

Verallina.. 

Verralina 

Verrallina 
butleri 
fragilis ob 
imitator 
indecorabilis 
malayi 
pygmaea 


xiv 


Page Page 
498 | Verrallina virilis ce a eedo2 
495 | 
AD aS W 
Worcesteria ye so LE 
| Wyeomyia ; 54. 447, 498, 501 
| aranoides .. =e SOE 
«s 373 HUET ECA = aear. see SON 
487 | greenii.. eee BON 
489 | metallica .. Se 8OR 
B90, oa nepenthicola feresOl 
49t  Wyeomyinae 3s = «) =-503 
491, 492 
491 | 
492 y 2 
491,492 Zeugnomyia 


487, 498, 499 


495 gracilis .. 498 


PREFATORY NOTE TO VOLUME IV. 


So many papers have been received and promised for the 
Records of the Indian Museum this year that it has been decided to 
publish a special volume (vol. iv) devoted to the systematic and 
geographical study of Diptera proved to be or likely to prove of 
economic or medical importance. The papers that constitute this 
volume will be issued as they are ready, concurrently with the parts 
of volume v, which will be devoted to general zoology. 

The preparation of vol. iv has been rendered possible by the 
untiring industry of Mr. E. Brunetti, by the assistance generously 
given by private collectors (notably by Lt-Col. E. A. W. Hall, 
I.M.S.) and by the liberality with which the Trustees of the Indian 
Museum have permitted their scientific officers to travel through 
India to collect and observe. These circumstances have combined 
to render our collection of Indian Diptera one of great importance 
and have enabled the Museum to resume the position as a centre of 
entomological work that it held in the days of Wood-Mason and de 
Nicéville. 

In addition to Mr. Theobald’s report on recent additions to the 
collection of mosquitoes with which it commences, this volume will 
contain a revision of the Oriental Gad-flies by Miss Ricardo, a revi- 
sion of the Oriental blood-sucking Muscide by Mr. Brunetti, an ac- 
count of the Indian Papataci flies (Phlebotomus) by Dr. Annandale, 
a revision of the Oriental Diptera Pupipara by Dr. Speiser, 
a revision of the Oriental Trypetidze (fruit-flies) by Prof. Bezzi, and 
other papers and notes. An account of the Indian Chironomidz 
(including many blood-sucking species) has been received from 
Prof. Kieffer and is now in the press, but as the group is a large one 
and has hitherto been almost completely neglected so far as Indian 
forms are concerned, this paper will be published in the Memozrs 
of the Indian Museum. 


N. ANNANDALE, 
Indian Museum, Superintendent. 
Calcutta : 
Feb. oth, 1910. 


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Po CON DERE PORT ON "THE COLLECTION 
OP SCULIChDis IN VTH ER TNDTAN MUSEUM , 
Cee Ut hre VW hee REET TONS: OF 
NEW GENERA AND SPECIES. 


By FRED. V. THEOBALD, M.A, etc. 


NOTE. 


The following report and descriptions of new species of 
Culicide is based on further collections belonging to the Indian 
Museum, Calcutta, sent me in 1908.' The types will be found 
in that Museum, but some co-types, where possible, have been 
retained for the British Museum of Natural History. Twenty 
new species are described and four new genera, also on2 new 
variety of an Anopheline. 

The series of banded-proboscis Culex provisionally placed as 
Culex tmpellens, Walker, must be specially worked out, as I find 
such wide variation that it is impossible to separate them and it is 
likely that several of the more recently described banded-proboscis 
species will have to sink as subspecies under one of the older names. 
This cannot be settled off-hand, males and females must be bred 
and the genitalia of the former and the larve examined before any 
further advance can be made in this group of Culex. 


SPECIES IN THE COLLECTION. 
Family CULICID. 
1. Anopheles lindesayi, Giles, var. maculata, var. nov. 


A very distinct variety of this marked species. The wings 
have a dense black spot of scales at the base of the long vein, a 
dense black spot of scales at the cross-veins, another at the base of 
the first fork-cell, a smaller one at the base of the second fork-cell 
and the outer edge of the costa, the first long vein and the base 
of the branches of the first fork-cell all very dark. ‘There are no 
pale spots on the wing fringe. 

Habitat Kurseong, 5,000 feet, Darjiling district, FE. Himalayas 
(Annandale). 

Time of capture 5-vii-08. 


1 The first report was published in the Records of the Indian Museum, vol. 
it, part iii, Oct. 1908, pp. 287—302. The species figured in the plates accompany- 
ing this report were described in the former report —Ep., Rec. Ind. Mus. 


2 Records of the Indian Museum. [Vor lve 


Observations.—A single perfect female. Evidently only a 
variety of Giles’s species, but the marked maculation of the wings 
gives it a very different appearance. 


2. Myzomyia rossit, Giles. 
Journ. Trop. Med., Oct., 1899. 


Additional Localities and Dates. 


Bengal.—Calcutta, ‘‘in house,” ro and 1I-viii-o8 (R. E. Lloyd) 
and 28-viii-08 (2) (2. H. Burkill) ; “‘ common below leather hoods of 
hired carriages and in public galleries of Indian Museum,” vii-o8 
(Annandale); Balliganj, near Calcutta, 9-x-08 (2) (7. Bentham) ; 
Bamungachi, Howrah, near Calcutta, 2 and 9-ix-08 (J. Caunter) ; 
Port Canning, Lower Bengal (2), 9-x-o8. Katihar, Purneah dis- 
trict, N. Bengal, 6-viii-o7 and 4—5-x-08 (2) (C. Paiva). Sur lake, 
near Puri, Orissa, 25 and 26-x-08 (2), ‘‘ biting by day when dis- 
turbed from old wells’”’ (A nnandaie). 

E. Bengal and Assam.—Chittagong, E. Bengal, 7-vili-08, 19 and 
21-ix-08 (Lt.-Col. Hall). 

Madras Presidency.—Gopkuda Id., Chilka lake, Ganjam dis- 
trict, near Bengal frontier, 7—15-viit-07 (R. Hodgart). 

Travancore State—Trivandrum, Vaikum (coastal region), 
Shencottah on the Madras frontier (e. side of W. Ghats), 
Kulattupuzha (w. base of W. Ghats): all collected by Dr. Annan- 
dale between 5 and 25-xi-08. 

The Punjab —Ferozepore (Lt.-Col. Adte) (34). 

At sea.—On board ship 10 miles off Coconada on the Madras 
coast, 17-iv-08 (C. Patva). 

NoTrEe.—One specimen from Calcutta, quite typical, with label 
stating that it was determined by Giles as Anopheles costals; he 
does not mention this in his Handbook. 


3. Myzomyra culicifacies, Giles. 
Entomologist’s Monthly Magazine, p. 197, Igor. 


Localities and Dates. 
Bengal.—Rajmahal, 31-viii-o7 (Kk. Hodgart). 
Burma.—Mandalay, 7 and @ ‘‘ in bathroom,’’ 13 and 
14-11-08 (Annandale). 
The United Provinces —Vucknow, 21-1-08 (Rk. Hodgart). 
The Punjab.—Ferozepore (Lt.-Col. Adie). 


4. Myzomyta christophersi, Theob. 
Proc. Royal Soc., \xix 91378 \ian; 1902: 


Localities and Dates. 


Benga!.—Calcutta (Lt.-Col. Alcock). 
E. Bengal and Assam.—Meenglas, Dooars, Jalpaiguri, 13-vili-07 
(C. Wallich) (1); Sylhet, Assam, 2-v-05 (Lt.-Col. Hall). 


1910. | F. V. THEOBALD: Report on Culicide. 3 


5. Nyssorhynchus janestt, Theob. 
Mono. Culictd., i, p. 134, Igor. i 
Localities and Dates. 


Bengal.—Calcutta, 5-vill-o8 (Annandale) ; Shamnagar, 3-viii-05 
(C. A. Gourlay) (3). 


6. Nyssorhynchus fuliginosus, Giles. 
Handbk. Gnats, p. 160, Ist Edit., 1900. 


Additional Localities and Dates. 
Bengal.—Calcutta (Museum premises), r5-vii-o8 and 12-viii-08, 
“at light in house,’’ 14-viii-o8, ‘‘ in bungalow at light” 
(Annandale); Balighai near Puri, Orissa, 23 and 24-x-08, ‘‘ numer- 
ous in old wells, resting by day ’’ (Annandale). 
The Punjab.—Ferozepore (20) (Li.-Col. Adte). 


7. Nyssorhynchus maculatus, Theob. 
Mono. Culicid., i, p. 171, Igor. 


Localities and Dates. 
Nepal.—Nara Ghat, Bengal frontier, 25 and 26-11-08 ; Thamas- 
pur, Bengal frontier, 18 and 20-11-08 (mus. collr.), 7 and @. 
Three rather large specimens but otherwise quite typical; the 
apical abdominal scales quite distinct. 
One o from the Punjab (?) has the outer costal border very 
dark, showing no third pale costal spot. 


8. Nyssorhynchus willmort, James-Theobald, 

Mono. Culicid., iti, p. 100, 1903. 

FE. Bengal.—Meenglas, Dooars, Jalpaiguri, 13-vii-o7 (C. Wal- 
lich). 

9g. Nyssorhynchus stephenst, Liston. 

Ind. Med, Gaz., 

Bengal.—Calcutta, ‘‘ flying by day in Museum garden,’’ 
15-vi1-08 (Annandale). 


Ravi INO. £2. TOOL. 


10. Cellia pulcherrima, Theob. 


Proc. Royal Soc., p. 369, vol. xix, 1902. 
The Punjab.—Ferozepore (Lt.-Col. Adie). 


11. Myzorhynchus barbirostris, Van der Wulp. 
Leyden Museum Notes, vi, p. 46, 1884. 


Localities and Dates. 


E.. Bengal and Assam.—15-i-07, Chittagong, 15-viii-o8 (Lt.-Col. 
Hall), 


4 Records of the Indian Museum. [VoL. IV, 


Travancore State-—Nedumangad, ten miles N.E. of Trivan- 
drum, 14-xi-o8 (Annandale). 


12. Mvyzorhynchus sinensis, Wiedemann. 
Ausser. Zweifl. Ins., p. 547, 1828. 


Additional Localities and Dates. 


Bengal.—Calcutta (Museum premises), 9-vi-08, 28-vii-o8, and 
viii-o8, ‘‘in bathroom ’’ (Annandale); Damukdia Ghat, R. Ganges, 
30-iv-08, “‘ at light on board steamer” (Annandale). 

Assam.—Sylhet, 13-i-03, 13-104, 20-i-03, 26-103, 23-11-05, 
27-v-00, 4 and 7-vi-05, 24-vii-o8 (Li-Col. Hall); Ukhrul, Manipur, 
6,400 feet, lat. 25° N., long. 94-95° E., viii-o8 (Rev. W. Pettigrew) 
(16). 

The Punjab.—Ferozepore (Lt-Co/. Adie). 

Travancore State.—-Maddathorai, w. base of W. Ghats, 18-ix-09 
(Annandale). 


13. Toxorhynchites tmmisericors, Wk. 


Journ. Proc. Linn. Soc. Lond., p. 91, 1860. 


9. Thorax clothed with metallic green and bronze scales, 
pale apple-green and pale blue at the sides in front and up to the 
base of the wings ; pleurze dense'y silvery white-scaled, dark above 
and below; head with a rich brown tinge in some lights, green in 
others, paler around the eyes; palpi and proboscis metallic purple 
and violet with mauve scales at the apex of the palpi. Abdomen 
as in the male but the tail tuft more pronounced. Fore legs 
with the metatarsals and first tarsal segments creamy white, rest 
dark: mid legs with the base of the metatarsals creamy white 
and all the first three tarsals; hind legs with the first two tarsals 
white. 

Note.—The @ is redescribed here as the specimen in the 
British Museum was too damaged to do so. 


Additional Localities and Dates. 


Bengai.—Calcutta (Museum compound), 6-i-07, 2, 16, 28 and. 
31-vil-07 (6), 2, 5, 6 and 26 vili-o7 (19) (Annandale); (Zoological 
Gardens), 9-vii and 12-vili-o8 (2) (Annandale). 

E. Bengal and Assam.—Chittagong, 3-ix-o8 (Li.-Col. Hall) ; 
Sylhet, Assam, 5-v-05 (Lt.-Col. Hall). 

The Himalayas.—Bhim Tal, 4,500 feet, Kumaon, 19 and 22-ix- 
06 (Annandale). 


14. Mucidus scataphagoides, 'Theob. 


Mono. Culictd., i, p. 277, I190l. 
N. Bengal.—Purneah, 5-viii-o7 (19 ) (C. Patva). 


15. Desvoidea obturbans, Wik. (and varieties). 


Proc. Linn. Soc. Lond., iv, p. 91, 1860. 


19:0. | F. V. THEOBALD: Report on Culicida, 5 


Additional Localities and Dates. 

Bengal.—Calcutta, 21-i-08, 13-iii-07, I6-ili-07, I5-vii-o7, 22-vii- 
07, 2-viii-07, 6-viii-07 (5), 28-i-08 (Annandale), 3 and 4-vii1-07 (2), 
15-Viii-07, 2-viii-o7, 23-viii-o7, 4, 7 and 15-xii-08, “‘ biting by day”’ 
(C. Paiva); Balliganj, near Calcutta, 30-ix, 9 and 11-x-08 (T. Ben- 
tham); Rajmahal, 30-vii-o7 (R Hodgart). 

The Himalayas.—Sukna, 500 feet, base of EK. Himalayas, 
Darjiling district, I-vii-o8 , ‘‘in deep jungle’”’ (Annandale) ; Kurseong, 
5,000 feet, Darjiling district, E. H.malayas, 5-vii-08 (Annandale). 

Travancore.—Trivandrum, 14-xi-o8 (Annandale). 

‘Numerous specimens of this species were reared in July, 
1909, by Mr. G. H. Tipper, from a glass of water left standing on a 
table in the office of the Geological Survey of India, Calcutta”’ 
(Annandale). 


16. Desvoidea panalectros, Giles. 


The co-type (Reg. No. *$3°) is nothing but an immature large 
Culex fatigans, Wied., with distinct abdominal banding. 


17. Desvoidea apicalts, sp. nov. 


Head dark brown, paler around the eyes; palpi and proboscis 
black. ‘Thorax bronzy brown, with a pale line around the front 
and sides; pleuree brown with silvery white-scaled areas. Abdomen 
black with prominent apical yellow bands and snowy white lateral 
spots. Legs black, unbanded, bases and venter of femora white. 

2. Head clothed with flat violet-black scales, some white 
ones forming a central line and small creamy curved ones forming 
a border around the eyes; a tuft of pale golden chetz projecting 
between the eyes, dark ones at the sides; clypeus black with 
small creamy spindle-shaped scales; palpi and proboscis deep 
violet-black ; antenne black, the basal segment testaceous and 
black with small creamy scales, base of the second segment bright 
ferruginous. 

Thorax black with narrow-curved almost hair-like bronzy 
scales, broader and pale ones forming a border around the front 
and sides of the mesothorax, the scales longer, broader and larger 
before the scutellum ; a dense tuft of brown cheetz over the roots 
of the wings; scutellum clothed with flat violet scales, border 
bristles rich brown; metanotum brown; pleura brown with 
patches of flat white scales. 

Abdomen black with violet reflections, all the segments but 
the first and last two with broad yellow apical bands, narrowed at 
the sides, and not passing quite to the edges; posterior border 
bristles small, brown with pale reflections ; large lateral white 
spots, most marked on the apical segments. 

Legs black, unbanded, femora pale beneath, the hind pair 
white below and at the base; chete dark (in some lights the legs 
have a brassy sheen); fore and mid ungues equal and uniserrate , 
hind equal and simple. 


6 Records of the Indian Museum. [Vor. IV, 


Wings with short fork-cells, nearly equal in length; the first 
submarginal narrower than the second posterior, its base a little 
nearer the apex of the wing, its stem nearly as long as the cell, stem 
of the second posterior as long as the cell, about one-and-a-half 
times its own length distant from the mid. Halteres with a pale 
testaceous stem, fuscous scales towards the white knob. 

Length 6 mm. 

Habitat Balighai near Puri, Orissa. 

Time of capture 24-x-08 (Annandale). 

Observations.—Described from a single perfect @. It is a 
typical Desvoidea and can be told at once by the very pronounced 
apical yellow abdominal bands. 


18. Brevirhynchus magnus, Theob. (Pls. ii and iii, wing and wing 
scales.) 


Recalnd. Musi it, p.-202, 1008. 


Additional Localities and Dates. 


The Himalayas.—Sukna, 500 feet, base of E. Himalayas, 
Darjiling district, I-vii-o8, ‘‘in thick jungle” (Annandale). 

Travancore.—Maddathorai, w. base of W. Ghats, 17-xi-08 
(Annandale) (1 @ ). 


1g. Brevirhynchus annulipalpis, sp. nov. 


Thorax black with scanty dull golden scales, arranged in 
rather obscure lines, sides and front of the mesonotum pale scaled ; 
pleuree with patches of white scales. Head black, pale in middle. 
The rather long black palpi with a snow-white band. Proboscis 
black, unbanded. Abdomen black with snowy white bands near 
the apex of some of the segments and with long thin lateral creamy 
spots. Legs dark with basal pale bands and pale cheetez. 

@. Head clothed with rather large, loose, flat dark scales, 
showing dull violet reflections, similar creamy ones forming a 
narrow median area, some creamy ones around the eyes and small 
almost white lateral flat scales, a few thick black chetz at the 
eye borders and golden ones between the eyes; clypeus dark with 
small flat pale scales; proboscis rather short, thick and uniformly 
black, metallic. Palpi about half the length of the proboscis, 
black with metallic violet reflections, a broad white band about the 
middle and a narrow creamy one nearer the base: antennz black, 
basal segment and base of the second bright testaceous, the former 
with small flat dark and creamy scales. Thorax black, clothed 
with long narrow-curved black, coppery and creamy scales, the 
latter in rather indistinct lines, sides of the mesonotum with rather 
broader pale creamy curved scales, which also pass around the 
front; the scales are larger, long and lanceolate over the roots of 
the wings, creamy and violet; dense dark brown cheetze also over 
the roots of the wings; scutellum large, clothed with loosely ap- 
plied flat violet and creamy scales and with rich brown border 


1910. | F, V. THEOBALD: Report on Culicide. 7 


bristles ; metanotum shiny black, shagreened over its surface ; pleurze 
brown with irregular patches of creamy white scales, which extend 
up to the edge of the pale areas of the mesonotum; prothoracic 
lobes with flat creamy white scales. 

Abdomen black with violet reflections, the first segment un- 
banded, with very fine pale hairs, second segment with a large 
creamy median basal spot, traces of a pale area on each side 
near the apex, but not nearly forming a band, the third to fifth 
segments with almost complete white bands towards the apical 
borders; a trace of pale scales on the sixth, similar to the 
second; apical segment with a brassy sheen and some golden 
brown chetz, a yellow lateral line on each segment; venter black 
and white. 

Legs black with white basal bands; femora black, pale 
beneath, in the hind pair white at the base and below except at 
the apex; a pale, almost white spot at the base of the fore meta- 
tarsals and first two tarsi, in the mid legs the banding is slightly 
more pronounced, especially on the metatarsi, in the hind legs all 
the tarsi have basal white bands and the chete are prominently 
pale, ungues equal and simple. 

Wings with dense brown scales; the first submarginal cell 
longer and narrower than the second posterior cell, its base nearer 
the base of the wing, its stem less than one-third the length of the 
cell; stem of the second posterior not quite as long as the cell; 
posterior cross-vein nearly twice its own length distant from the 
mid; halteres with the stem grey, then fuscous and the knob 
creamy white. 

Length 5 5 mm. 

Habitat Maddathorai, w. base of W. Ghats, Travancore 
(Annandale). 

Time of capture 16-xi-08. 

Observations.—Described from a single perfect @. At once 
told from any other known member of the genus by the banded 
palpi. The abdomen is relatively long and thin. The proboscis 
is short and thick but not so bent as in the type of the genus. 


20. Brevirhynchus apicalis, sp. nov. 


Head ochreous with a median and two lateral black patches ; 
proboscis and palpi black. Thorax deep golden brown scaled with 
a pale ochreous border around the sides and front; pleuree brown 
with creamy patches of scales; abdomen black, with apical vellow 
bands, show ng some faint traces of a median pale line, lateral 
median white spots and ochreous venter. Legs brown with nar- 
row apical pale bands. 

@. Head with flat ochreous scales, a small median dark- 
scaled patch in front, then a large and then a small dark-scaled 
lateral patch, borders of the eyes ochreous, golden cheetee project- 
ing between them, brown ones at the sides; eyes coppery red ; 
clypeus brown, palpi long and thin, black, traces of a narrow pale 


8 Records of the Indian Museum. [ VoL. IV, 


band towards the base; proboscis black ; antennz brown, basal 
segment bright ferruginous, with small flat grey and fuscous scales, 
base of the second segment ferruginous. 

Thorax black, thickly clothed with narrow-curved deep golden 
brown to bronzy scales, with a distinct ochreous hordei surround- 
ing the sides and front, some paler broader scales and golden 
brown cheete over the roots of the wings; scutellum clothed with 
flat black and ochraceous scales, mixed together ; border bristles 
golden; metanotum chestnut-brown ; pleurze brown with patches 
of small flat creamy scales. 

Abdomen black, the second to sixth segments with prominent 
broad yellow apical bands, expanded in the middle but not con- 
tinued quite to the edge of the segments, seventh witha few 
pale scales, eighth mostly ochreous scaled ; each segment with a 
median lateral pale spot; venter ochreous. Legs brown, hind 
femora pale beneath, apex of tibiz with a pale ochreous spot con- 
tinued on to the base of the metatarsus, the tibiz with narrow apt- 
cal and basal pale yellow bands; fore and mid ungues uniserrate, 
hind equal and simple. 

Wings with rather short fork-cells ; the first submarginal cell a 
little longer and narrower than the second posterior cell, its base 
slightly nearer the apex of the wing, its stem more than half the 
length of the cell, stem of the second posterior not quite as long 
as the cell ; posterior cross-vein long, twice its own length distant 
from the mid. Halteres with pale stem and fuscous knob. 

Length 8 mm. 

Habitat Sylhet, Assam (Lt.-Col. Hall). 

Time of capture 26-vi-: 5. 

Observations.— Described from a single 9. It is a large and 
distinct species easily identified by the apical and basal leg band- 
ing, apical abdominal bands and ochreous venter. ‘The length of 
the palpi and short thick proboscis agree with Brevirhynchus, but 
the head and thoracic scales, although of the same general form, are 
not so large as in the other two species. I see no reason however 
to separate it from that genus. 


21. Stegomyta fasciata, Fabricius. 


Syst. Antl., p. 36, 13, 1805. 


Additional Localities and Dates. 

Bengal.—Calcutta, 27-vii-o7, 27-viii-o7 (C. Paiva), 28-vii-08 
(I. H. Burkill), “ in Museum building,” vii-o8 (T. Bentham), July 
and August 07 (mus. colly.); Puri, Orissa coast, 18 and I9-i-09 
(Annandale). ‘* Not so common in Calcutta as S. scutellaris”’ 
(Annandale). 

FE, Bengal and Assam.—Chittagong, 21-ix-03 (Lt-Col. Hall). 

Madras Presidency,—Madras town, 31-x-08 (R. Hodgart). 

Burma.— Rangoon, “‘ in house, biting by day,” 25-ii-08 ; Man- 
dalay, 12-111-08 (Annandale). 


TQIO. | F. V. THEOBALD: Report on Culicide. 9 


At sea—On board ship, Bay of Bengal, between mouth of 
R. Hooghly and Rangoon, 22 and 23-it-08 (Annandale) ; on board 
ship, 10 miles off Coconada on the Madras Coast, 15-iv-08(C. Paiva). 
**T found.this species common on board ship all the way from 
Calcutta to’Rangoon in February ’’ (Annandale). 


22. Stegomyia scutellaris, Wik. 
Journ. Proc. Linn. Soc. Lond., iti, p. 77 (1859). 


Additional Localities and Dates. 

Bengal.—Calcutta, r1-iti-o8 (2), 15 and 23-vil-07, 6-vili-07 (1), 
19-ix-07 (6), 3I-vii-07, 5-viii-o7, (Zoological Gardens) 12-vii-08, 
2-vili-o7 (Annandale); Purneah, N. Bengal, 5-viii-o7, Bhogaon, 
Purneah district, N. Bengal, 7-x-08 (8), 9-x-08 (1) (C. Pazva) ; 
Balighai near Puri, Orissa, 23 and 26-x-08 (Annandale). ‘‘ Very 
abundant in Calcutta between March and November ”’ (Annandale). 

The Himaiayas.—Sukna, 500 feet, base of FE. Himalayas, 
Darjiling district, r and 2-vii-o8 (7), “‘only troublesome during 
day’ (Annandale); Bhim Tal, 4.500 feet, Kumaon, ‘‘ breeds in 
hollow trees in jungle,” ix-06 (Annandale). 

Madras.—Madras town, 30-x-08 (7) (R. Hodgart). 

Travancore State.—Trivandrum, 14-xi-o8 (2), Maddathorai (w. 
base of the Western Ghats), 18-x-08 (Annandale). 
. The United Provinces.—Shahjahanpur, ‘‘day biting’’ (G. M. 
Giles), one @ named by Giles Culex albopictus, Skiise. 

Burma.—Mandalay, IT-iii-o8, “‘ not uncommon in house ’ 
(Annandale). 

Ceylon.—Colombo, Victoria Gardens, 26-iv-08 (C. Patva). 


) 


23. Culex mimeticus, Noé. 


Bull. Soc. Ent. Ttal., xxxi, p. 240, 1899. 


The Himalayas.—Theog, 8,000 feet, Simla Hills, 2-v-07 (1 2 ) 
(Annandale). 

Nepal.—Thamaspur, Bengal frontier; 18 and 20-11-08 (Io), 
large specimen (mus. collr.). 


24. Stegomyia minulissima, sp. nov. 


Head black, a white border to eyes and white median spot in 
front between them ; palpi black with white apices ; proboscis black. 
Thorax deep brown with a small grey-scaied area in front, grey 
scales in front of the roots of the wings and three short creamy 
lines behind. Abdomen black with narrow white basal bands and 
white lateral spots. Legs dark brown with basal white bands ; 
mid femora with a marked median white spot and white apex: 
hind femora white at base. Very small species. 

9. Head clothed with flat black scales, a large median 
triangular white-scaled area in front and a pale border to the 
eyes ; clypeus dark brown; palpi black with snowy white apices ; 


10 Records of the Indian Museum. [VoL. IV, 


proboscis black; antennee deep brown, basal segment with white 
scales; some golden brown chet project between the eyes. 

Thorax black, the greater surface clothed with bronzy brown 
longer narrow-curved scales, a small area near the head anda 
small wedge-shaped area on each side in front of the wings with 
dull white scales, some pale creamy scales which form three indis- 
tinct lines behind in some lights; scutellum with flat black scales 
with dull violet reflections and some white ones on the lateral lobes ; 
pleure brown with flat white-scaled spots. Cheetee brown and 
black. Abdomen black with narrow basal white bands and basal 
snowy white lateral spots. Legs deep brown; the first pair with 
a basal white band to the first tarsal, the mid with a median white 
spot on the femora, the apex white and a basal white band to the 
metatarsal and first tarsal; the hind with the femora white at the 
base, and all the segments with a basal white band except the last; 
some large black cheete at the apices of the segments in the hind 
legs ; fore and mid ungues uniserrate, hind simple. 

Wings with typical large brown Stegomyian scales on the basal 
region, dense narrow linear ones on the apical areas of the veins ; 
the first submarginal cell longer and narrower than the second 
posterior, their bases nearly level. Stem of the first fork-cell 
about half the length of the cell, stem of the second posterior 
about two-thirds the length of the cell; posterior cross-vein twice 
its own length distant from the mid. 

Length 2°5 mm. 

Habitat Sukna, 500 feet, base of E. Himalayas, Darjiling 
district (Annandale). 

Time of capture 1-vii-08. 

Observations.—Described from 392’s. One of the smallest 
Stegomyi@ I have seen. It can easily be told by the thoracic orna- 
mentation and the white round spot on the femora of the mid legs. 
One specimen was taken in a bungalow. 


25. Stegomyta annandalei, sp. nov. 


Head black with a large median white patch ; palpi black with 
snowy white apices ; proboscis black. ‘Thorax black-brown with a 
snowy white patch in front and one over the roots of the wings. 

Abdomen jet-black with snowy white basal bands which gradu- 
ally swell out laterally. Legs black banded with snowy white, the 
hind legs with a band at the base of the metatarsals, first tarsals 
and the whole of the third white. 

@. Head clothed with flat black scales, with a large median 
triangular snowy white patch and a small dull white lateral patch, 
some black cheetze in front; palpi black-scaled with broad snowy 
white apices: proboscis black; antenne deep brown, basal seg- 
ment with dense snowy white scales. 

Thorax shiny black, clothed with long bronzy brown narrow- 
curved scales, except for a small snowy white patch of similar 
scales in front and a patch of flat snowy white scales over the base 


1910. | F. V. THEOBALD: Report on Culicide. i 


of the wings oneach side; supra-alar cheetze dense, black and long ; 
scutellum with the mid lobe clothed with flat black scales, the 
lateral lobes with flat white scales ; border bristles black , metano- 
tum deep brown; pleura dark with numerous white patches of 
scales. 

Abdomen black with four snowy white basal bands on the 
last four segments, which broaden out laterally, the two preced- 
ing segments with basal white lateral spots. 

Legs black, the fore and mid with basal white bands on the 
metatarsi and first tarsal ; the hind with a basal white band on the 
metatarsi and first tarsal segments, the second all dark, the third 
nearly all white, the fourth dark ; ungues apparently all simple. 

Wings with brown scales ; fork-cells short, the first submargi- 
nal longer and narrower than the second posterior, its stem a little 
nearer the base of the wing than that of the second posterior, its 
stem about two-thirds the length of the cell ; stem of the second 
posterior nearly as long as the cell; posterior cross-vein rather 
more than twice its own length distant from the mid. 

Length 2°8 mm. 

Habitat Sukna, 500 feet, base of EK. Himalayas, Darjiling 
district (Annandale). 

Time of capture vit-o8. 

Observations.—Described from a single perfect @. It resem- 
bles Stegomyia minutissima but can at once be told by the banding 
of the posterior legs and still more definitely by the flat white 
scales forming the lateral thoracic patches. The specimen was 
taken in a bungalow. 


\ 


26. Stegomyta albipes, sp. nov. 


Thorax brown, a small silvery white patch in front and a 
white-scaled line running down to the base of the wings where 
there is a large snowy white patch extending on to the dorsum 
and scutellum. Head black, white in the middle; palpi black 
with snowy apex; proboscis black. Abdomen black with narrow 
basal white bands and large snowy white lateral basal spots. 
Legs with broad basal white bands, last two hind tarsi white ; mid 
femora and tibie with a median white round spot. 

9. Head black, clothed with flat black scales at the sides, 
white in the middle; chete black; proboscis and clypeus black ; 
palpi black with snowy white apices; antenne dark drown, basal 
segment with dense flat snowy white scales. 

Thorax dark with narrow-curved deep bronzy brown scales, 
snowy white ones forming a broadish line in front and extending 
laterally towards the wings before which they spread out into a 
large white patch, which passes on to the dorsum; this is composed 
of narrow-curved scales above and flat ones below; there is also a 
white line of narrow-cu'ved scales on each side just past the roots 
of the wings and a few white scales here and there before the 
scutellum ; supra-alar cheetee black; scutellum clothed with rather 


1 Records of the Indian Museum. [VOESLW ¢ 


large flat snowy white scales; metanotum black; pleure black 
with silvery white puncta. 

Abdomen black with narrow basal snowy white bands and 
very large basal lateral white spots ; posterior border bristles pale, 
short. 

Legs black with white bands and spots; the fore femora pale 
at the base ventrally, fore tibia with a white median spot; knee 
spot white ; white bands to metatarsi and first tarsals ; mid femora 
with prominent white median spot (almost a band), a smaller one 
basally and a white apex, metatarsi and first tarsal basally white; 
hind femora white basally, and on most of the venter, apex white; 
tibiee with a white band on the apical half, metatarsi and first two 
tarsi basally snowy white, and all the last two tarsi; cheetee black ; 
ungues equal and simple. 

Wings with dark scales, base pale; first submarginal cell longer 
and narrower than the second posterior, its base nearer the base 
of the wing, its stem about half the length of the cell; stem of 
the second posterior about as long as the cell; posterior cross- 
vein rather more than twice its own length distant from the mid. 
Halteres with testaceous stem and deep fuscous knob. 

Length 3 mm. 

Habitat Maddathorai, w. base of W. Ghats, Travancore 
(Annandale). 

Time of capture 17-xi-08. 

Observations.—Desctibed from a perfect @. At once sepa- 
rated from any other S/egomyta by the last two hind tarsi being 
white and by the thoracic ornamentation, especially by the pre- 
alar white patch being composed of narrow-curved scales above, 
flat ones below. The femoral and tibial markings are also very 
distinctive. 


27. Stegomyta assamensis, Theob. 
Rec. Ind. Mus., ti, p. 290, 1908. 


Travancore.—Pallode, twenty miles N.E. of Trivandrum, 
15-x1-08 (Annandale), I small @. 


Genus PSEUDOCARROLLIA, gen. nov. 


Head clothed with flat scales and upright forked scales, a 
border of spindle-shaped scales around the eyes. Palpi of 9 
about one-fourth the length of the proboscis. ‘Thorax with nar- 
row-curved scales; scutellurn with flat scales. Abdomen with 
dense ventral scale tufts on some of the apical segments. Fork- 
cells rather short, vein-scales rather thick. 

Allied to Carrollia (1jatz) but differs in having the scutellum 
with all flat scales and in the absence of narrow-curved scales 
forming a median basal area on the head. 

The marked ventral abdominal scale tufts resemble those of 
Carrollia and Hemagogus. 


1910. | F. V. THEOBALD: Report on Culicide. 13 


28. Pseudocarrollia lophoventralis, sp. nov. 


Head black, a white border to eyes, eyes silvery above; pro- 
boscis and palpi jet-black; thorax with the front bright silvery 
white, also the scutellum, the posterior half of the mesonotum 
rich brown; pleuree with silvery white puncta. Abdomen black 
with a dull greenish tinge, basal lateral snow-white spots and 
basal white ventral bands, and ventral black scale tufts. Legs 
banded with white, base and apex of metatarsi white, also base of 
the front and hind first tarsal, base and apex of the mid; femora 
white at base and with a white apical spot on hind pair. 

9. Head black, clothed with flat black scales and a narrow 
border of spindle-shaped white ones around the eyes; narrow 
black upright forked scales and black cheetz projecting forwards ; 
clypeus, palpi, antennee and proboscis all deep black ; eyes silvery 
around the edges, black in the centre. 

Thorax black, clothed on the front half with dense long nar- 
row-curved snowy white scales, not quite extending to the pleurze 
on each side, forming more of a large round patch, remainder of 
mesonotum with rich bronzy scales except for a few snowy white 
ones over the roots of the wings; supra-alar cheetee long and black ; 
scutellum black, thickly clothed with snowy white flat scales and 
golden border bristles; pleuree rich brown, with six or seven white 
spots ; metanotum black. 

Abdomen black, with the scales black, showing in some lights 
green and peacock-blue reflections, the green mainly at the tips of 
the scales; the segments with prominent basal lateral snow-white 
spots, six on each side, last segment small, basally white; poster- 
ior border bristles pale; venter black with snow-white basal 
bands, the fifth to the seventh segments with dense black out- 
standing scales, giving a tufted appearance. Legs black with 
white bands; front legs with apex of tibize an1 metatarsi white, 
and base of first tarsal; mid legs with femora white at the base 
ventrally, a small white apical spot, a white spot at the apex of 
tibiee, a white band at base and apex of metatarsi, also base and 
apex of first tarsal and a trace at the base of the second tarsal ; 
in the hind legs the base of the femora are snowy white and there 
is a large white apical band and the base and apex of the meta- 
tarsi banded white, also base of first tarsal; leg bristles black; 
fore and mid ungues uniserrate, hind simple. 

Wings with short fork-cells, the first submarginal narrow but 
no longer than the second posterior, its stem more than half the 
length of the cell; stem of the second posterior also more than 
half the length of the cell; posterior cross-vein about twice its own 
length distant from the mid, the latter and the supernumerary are 
in a straight line; scales brown, rather dense on the branches 
of the fork-cells. Halteres with testaceous stems and fuscous 
knobs. 

Length 5°5 mm. 

Habitat Purneah, N. Bengal (C. Patva). 


14 Records of the Indian Museum. [VoL. IV, 


Time of capture 0-viii-o7, ‘‘ resting on under side of leaf of 
licht tree during day”’ (C. Paiva) (1 @ ). 

Observations.—Described from a single perfect @. It clearly 
comes in a new genus, related to Lutz’s Carrollia. The very 
marked thoracic, abdominal and leg ornamentation will at once 
separate it from any species coming in the Stegomyian group. 
The ventral abdominal tufts are very marked. 


29. Huleceteomyta trilineata, Leicester. 


The Entomologist, xxxvii, p. 163, 1904. 

FE. Himalayas.—Kurseong, 5,000 feet, Darjiling district, 3-vii- 
08 (Annandale), 1 7 and 2 9’s. 

NotEe.—These answer exactly to Leicester’s carefully described 
species, but one @ shows distinct white abdominal basal bands 
and also the ~; the second @? is quite typical. 


30. Grabhamia pulcripalpis, Rondani. 


Bull. Soc. Ent. Ital., iv, p. 31, 1872. 


Egypt.—On board ship at night, Suez Canal, 9-x-07 (Reg. Nos. 
ts and *3$°), 2 2’s (Annandale). Rather small specimens. ] 


31. Pseudograbhania maculata, Theobald. 


Journ. Bomb, Nat. Hist. Soc., xvi, p. 244, £905. 
Madras.—Madras town, 30-x-08 (R. Hodgart),1¢. 


32. Culex vagans, Wiedemann. 


Ausser. Zweifl. Ins., p. 545. 


@. Head brown with pale scales; palpi longer than probos- 
cis by about the apical segment, which is slightly shorter than the 
penultinate, deep brown, the last two segments with bright basal 
creamy bands and flaxen brown and brown hair tufts, long on the 
inner side of the penultimate segment and a long dark tuft on the 
apex of the ante-penultimate, which is swollen, two other broad 
pale bands below. Antenne brown with pale internodes and 
flaxen brown plume hairs. 

Thorax deep, rich brown with small narrow-curved golden 
brown scales; scutellum with paler narrow-curved scales, larger 
in size, with brown border bristles; metanotum brown. Abdomen 
deep brown, almost black, with basal white bands; hairy, hairs 
pale brown; basal lobes of genitalia very hairy, claspers broad- 
ish, with a large spine on the side near apex giving a bifurcate 
appearance and with dense fine hairs all along the outer edge. 
Legs deep brown, paler at the base with basal pale bands, ungues 
of fore and mid pairs unequal, uniserrate, the mid pair large 
(hind ? uniserrate). Wings with rather short fork-cells, the first 
longer and narrower than the second which is rather broad, their 


1gIo. | F. V. THEOBALD: Repori on Culicide. 15 


bases about level, the stems nearly as long as the cells; mid cross- 
vein about twice as long as the supernumerary and in a straight 
line with it; posterior cross-vein about the same length as the 
mid, about its own length distant from it. 

Length 5°2mm. 

Habitat Madras town (R. Hodgart). 

Time of capture 31-x-08. 

NotE —-This is the only @ of this species I have seen, so I 
have redescribed it. 


33. Culex impellens, Wik. (°). 
Proc. Linn. Soc. Lond., iv, 9I. 


Localites and Dates. 

Bengal.—Calcutta, 31-i-o8, 5 and 7-i1i-07, 3-vi-08, 12-vit-08, 
2i-vii-o8, 27 and 28-vii-o8, 30-vii-o7, 3I-vii-o8, 2—6-viii-07, 
10-viii-08 , 13 and 14-viii-o8, ‘‘in bungalow ’’ (Annandale), 18-vii1-08,, 
22-vili-o8, ‘‘in bathroom’’ (Annandale), 19-ix-07, 26-ix-08; Balli- 
ganj, near Calcutta, 20-vi-o8 (3) (T. Bentham); Port Canning, 
Lower Bengal, 19-x-08 (Annandale); Rajmahal, 31-vii-07 (2) 
(R. Hodgart); Purneah, N. Bengal, 6-viii-o7, Bhogaon, Purneah dis- 
trict, N. Bengal, 3-x-08 (2), 6-x-08 (1) (C. Paiva) ; Bettiah, Cham- 
paran, 7-iii-o8 (R. Hodgart); Balighai near Puri, Orissa, 25-x-08 
(3), 26-x-o8 (8), 24-x-08 (2), ‘‘in house, bites occasionally by 
day,” 23-x-08 (6), ‘‘in house, comes to light” (Annandale). 

E. Bengal and Assam.—Chittagong, E. Bengal, 26-vii-08 
(Lt.-Col. Hall). 

The Himalayas.—Sukna, 500 feet, base of FE. Himalayas, 
Darjiling district, I and 2-vii-08 (5), ‘‘ in deep jungle, biting by 
day’’ (Annandale). 

Nepal.—Thamaspur, Bengal frontier, 1o—r14-ii-o8, Sukwani, 
Bengal frontier, 15 and 16-ii-o8 (Butchu, mus. collr.). 

Burma.—Moulmein, L. Burma, 27-ii-08, ‘‘ abundant in house,” 
Rangoon, 24 and 25-ii-o8, Mandalay, 11 and 12-iii-o8 (Annandale). 

The United Provinces.—Lucknow, 21-iv-07 (1) (Annandale). 

Travancore and Cochin States.—Shencottah, Madras frontier 
(e. side of W. Ghats) (4); Pallode, twenty miles N.E. of Trivan- 
drum; Shasthancottah, twelve miles N.N.E. of Quilon; Trivan- 
drum; Nedumangad, ten miles N.E. of Trivandrum (2); Vaikum 
(coastal region); Kulattupuzha (w. base of W. Ghats); Tenmalai, 
W. Ghats (w. side); Ernakulam, Cochin State. All collected by 
Dr. Annandale between 4 and 25-xi-08. 

At sea.—At light on board ship five miles off Aleppey, 
Travancore, 4-v-08 (2) (C. Paiva). 


34. Culex pettigrewit, sp. nov. 


Head brown with pale scales; palpi brown, pale-scaled apices ; 
proboscis pale except for a small dark band at the apex and base ; 
thorax rich brown, ornamented with a paler curved line on each 


16 Records of the Indian: Museum. [VoL.; IV; 


side, another short one over the roots of the wings and two indis- 
tinct median pale lines in front, each with a central dark line. 
Abdomen deep brown, with basal grey bands, two grey median 
spots to the segments, almost joining on to the bands, sixth to 
eighth segments with many scattered ochreous scales; venter 
bright ochreous with scattered pale scales. Legs dark, minutely 
banded, apically and basally on the tarsi, the femora and tibiz 
with a pale creamy ventral line. Wangs with the subcostal vein 
pale-scaled. 

@. Head dark, clothed with rather large narrow-curved 
creamy scales and flat creamy lateral ones and dark chete in 
front; upright forked scales dark, a few pale ones in front; cly- 
peus brown with grey sheen; palpi brown with creamy scales at 
the apex and some nearer the base; proboscis dark at base and 
apex, the median creamy area with some scattered dark scales; 
antennee dark brown, base of the second segment bright testa- 
ceous. Thorax dark brown clothed with rich brown narrow-curved 
scales; a pale-scaled twice curved line on each side and a pale- 
scaled short line on each side in front and over the roots of the 
wings; in middle (in front) are traces of two parallel narrow dark 
lines with somewhat paler scales on each side; supra and pre- 
alar cheetee brown with pale apices, the supra-alar long and darker 
than the others; scutellum pale brown with narrow-curved pale 
scales and long brown posterior border bristles; the scales at the 
back of the mesonotum are similar in colour to those of the scutel- 
lum; metanotum bright brown; pleuree brown with patches of 
pale scales. 

Abdomen with the basal segment ochreous with pale scales, 
the second to fifth segments dark with grey basal bands which 
spread out in the middle and with two ochreous spots nearly touch- 
ing them, most pronounced on the fourth and fifth segments, the 
sixth with many ochreous scales dotted over the dark area, the 
seventh and eighth almost entirely ochreous; posterior border 
bristles thin and pale golden; venter bright ochreous-yellow with 
pale scales, a few dusky ones in the mid region. 

Legs rather long, dark, femora and tibiz pale below, those of 
the fore and mid legs with a pale-scaled line in addition; faint 
traces of very narrow apical and basal pale banding to the hind 
tibiae, metatarsi and tarsi, scarcely perceptible on the fore and 
mid legs; bases of the legs pallid; femora and tibize slightly spinose, 
pallid; ungues equal and simple. 

Wings with brown scales, a few creamy ones at the base of 
the costa and on all the subcostal vein and a few pale scales on the 
first long vein; first submarginal cell much longer and narrower 
than the second posterior cell, its base much nearer the base of the 
wing, its stem about one-third the length of the cell, stem of the 
second posterior nearly as long as the cell, posterior cross-vein not 
quite its own length distant from the mid cross-vein. 

Halteres ochreous with some fuscous scales on the knob. 

Length 6 mm. 


IQIO. | F. V. THEOBALD: Report on Culicide. Y7 


Habitat Ukhrul, Manipur, 6,400 feet, lat. 25° N., long. 94— 
95° E. (Rev. W. P. Pettigrew). 

Time of capture viii-08. 

Observations.—Described from a perfect ungorged 2. Three 
other °’s were fully gorged and black. The marked character of 
this species is the pale-scaled subcostal vein. The pale scales also 
occur here and there on the upper surface of the first long vein. 
The abdomen in the type is very marked but in two of the others, 
which are somewhat rubbed, the spots on the segments are not so 
distinguishable. 


35. Culex fuscocephala, Theobald. 
Mono. Culcid., iv, p. 420, 1907. 


Travancore.—Pallode, twenty miles N.E. of Trivandrum, 
15-xt-08 (Annandale), 1°. 


36. Culex fatigans, Wiedemann. 
Ausser. Zweifl. Ins., p. 10, 1828. 


Additional Localities and Dates. 

Bengal.—Calcutta, 7, 13 and 22-i-08, 31-i-08, 1-11-07, 17-11-08, 
13-iii-07 (3), 29-iii-o8, 4-iv-o8, 1-v-08 (6), 21-vi-o8, 21, 27 and 
28-vii-07 (4), 3—7-viii-o7, 25-iii and 22-vii-o8, ‘‘common in 
Museum premises’’ (Annandale), Balliganj, near Calcutta, 5 and 
6-ii-08 (12) (J. B. Richardson); Rajmahal, 31-viii-o7(3) (R. Hod- 
gart); Bettiah, Champaran, 4-iii-08 (27), 5-iii-o8 (8) (R. Hodgart) ; 
Puri, Orissa coast, 18 and 19-i-08 (7), 20 and 21-i-08 (1) (Avnan- 
dale), 2-iii-o8 (2) (C. Paiva); Balighai near Puri, Orissa, 25-x-08 
(Annandale). 

E. Bengal and Assam.—Rajshahi, 1—6-ii-07 (Annandale) ; 
Ukhrul, Manipur, 6,400 ft., lat. 25° N., long. 94—95° E., viii-o8 
(Rev. W. Pettigrew). 

The Himalayas.—Naini Tal, Kumaon, 6—7,000 feet, 1908 
(R. E. Lloyd); Dharampur, 5,000 feet, Simla hills, 13 and 14-v-08 
(Annandale); Kurseong, 5,000 feet, E, Himalayas, Darjiling dis- 
trict, 3—5-vii-o8, Sukna, 500 feet, base of E. Himalayas, Darjiling 
district, I-vil-o8 (Annanda/e). 

Nepa!.—Thamaspur, Bengal frontier, 18—20-ii-08 (2), Daha- 
wangahary Hill, near Bengal frontier, 16-ii-o8 (Butchu, collr.) ; 
Khatmandu (two @’s), Oct. 1906 (kk. Hodgart). 

Burma.—Mandalay, 6-iii-o8, 11, 12, 13 and 17-11-08 (Amnan- 
dale), Rangoon, 1-i-05 (3), 3-i-05, 23-ii-05 (8) (Brunettr), 24-11-08 (3), 
25-11-08 (3), 15-iii-o8, in house (Annandale); base of Dawna hills, 
4-iii-o8, Moulmein, 27-i-08 (11), 7-it08, 27-it08 (2), 28-11-08, 
6-ii1-08 (4), I2-1ii-08 (2) (Annandale). 

Travancore State.—Tenmalai (w. side of W. Ghats), 22-xi-o8 
(Annandale). 

The United Provinces.—Lucknow, 21 and 27-i-08 (2), 5-ii-08 
(R. Hodgart), 21-iv-o7 (4) (Annandale), 7-viii-o7 (Brunettt) ; Agra, 


18 Records of the Indian Museum. [MOL SLVs 


4-iv-05 (Brunettr) ; Dhikala, Naini Tal district, 22-iv-08 (4), 26-iv-08 
(5), Chuharwala, Naini Tal district, 15-iv-o8 (3), Patair Naini 
Tal district, 27-iv-08 (3) (Hodgart). 
The Punjab (plains)—tWahore, 8-v-08 (11) (Annandale) . 
Ferozepore (Lt.-Col. Adie). 
At sea—Bay of Bengal, on ship between mouth of R. 
Hooghly and Rangoon, 22 and 23-i1-08 (Annandale). 
Java.—Soerabaya, 16—25-vii-06 (Bruneitt). 
Philippines.—Manilla, Io—16-11i-06 (Bruneitt). 


37. Culex parascelos, sp. nov. 


Thorax golden brown, and with three paler median parallel 
lines, the areas between them darker than the sides; head, palpi 
and proboscis bright ochreous, apex of the latter black. Abdomen 
entirely clothed with ochreous scales. Legs ochreous-brown above, 
pale ochreous below with dark and ochreous linear ornamentation. 
Wings with ochreous and brown scales on the veins near costa ; 
fork-cells short ; ungues large, all uniserrate. 

@. Head brown with smal! narrow-curved pale golden scales, 
ochreous upright forked scales, somewhat darker behind; 
ochreous cheetze ; clypeus and palpi bright ochreous, almost golden 
yellow, the latter with prominent black cheetee; proboscis the 
same colour, dark at the apex, with black chete. Antenne 
brown, ochreous at base. Thorax deep brown, clothed with 
scanty small narrow-curved pale golden and rich golden brown 
scales, the former making two prominent lateral median lines and 
an indistinct median one; chetz bright golden brown; scutellum 
brown with narrow-curved pale golden scales and golden brown 
chetee; metanotum brown; pleure brown with patches of 
flat ochraceous scales. Abdomen dark, densely clothed with 
bright ochreous-yellow scales and pale border bristles; venter 
pale ochreous. 

Legs ochreous with dusky scales above, femora pale below; a 
marked yellow and black line showing on the femora and tibia ; 
ungues dark, all equal and uniserrate, rather thick. 

Wings tinged with yellow, some pale scales on the costa and 
sub-costal, remainder dark, except for a few creamy ones on the 
base of the first long vein; first submarginal cell much longer and 
narrower than the second posterior cell, its base slightly nearer 
the base of the wing, its stem about two-and-a-half times the 
length of the cell; stem of the broad second posterior as long as 
the cell; posterior cross-vein about the same length as the mid, 
not its own length distant from it. 

Length 5°5 mm. 

Habitat Madras town (R. Hodgart). 

Time of capture 30-x-08. 

Observations.—Described from two @’s. A very marked 
species easily told by the thoracic ornamentation, ochreous-scaled 
abdomen and lined legs. 


1910. | F. V. THEOBALD: Report on Culicide. 19 


38. Culex longifurcatus, sp. nov. 


Thorax uniformly fuscous and smoky; head much the same 
colour but with some paler scales ; palpi, proboscis and antennze 
smoky brown; abdomen deep brown, with basal pale bands 
spreading out in the middle to form median patches; legs uni- 
formly smoky brown; wings rather long and narrow, fork-cells 
long, the veins and scales very thin. Male palpi thin, no hair tufts. 

Q. Head dark, clothed with dull creamy narrow-curved 
scales, small flat creamy lateral ones, numerous dark upright 
forked scales, some paler ones in front; palpi, clypeus, proboscis 
and antennee brown. 

Thorax dark brown with fuscous-brown scanty narrow-curved 
scales and brown cheetee; scutellum brown with similar narrow- 
curved scales; metanotum brown and testaceous; pleure dark 
brown and grey. Abdomen deep brown, with basal, almost white 
bands which are enlarged in the middle forming more or less pro- 
nounced spots: border bristles pallid; venter all grey-scaled. 
Legs long and thin, uniformly brown, except the venter of the 
femora which is completely creamy white; ungues equal and sim- 
ple. Wings large and long; fork-cells long; first submarginal cell 
longer and narrower than the second posterior, their bases nearly 
level, its stem about one-third the length of the cell: stem of the 
second posterior less than one-third the length of the cell ; posterior 
cross-vein longer than the mid about one-and-a-half times its own 
length distant from it; lateral vein scales on the apical areas of 
the veins long and thin. 

Length 5 mm. 

@. Palpilong and thin, dark brown, acuminate, no hair tufts, 
a few dark hairs and chetee; apex of proboscis swollen. Wings 
with relatively long fork-cells ; first submarginal cell narrower and 
longer than the second posterior, its stem less than one-third the 
length of the cell; stem of the second posterior rather more than 
half the length of the cell; posterior cross-vein nearly twice its own 
length distant from the mid. Fore and mid ungues unequal, both 
uniserrate, the larger with a large tooth, hind equal and simple. 

Length 4 to 4°45 mm. 

Habitat Dahawangahary Hill, near Bengal frontier, Nepal 
(Butchu, mus. collr.). 

Time of capture 16-ii-08. 

Observations.—Described from two ?’s and one ~. A very 
marked Culex with obscure ornamentation, easily told by the long 
fork-cells, very thin veins and large wings and long legs. ‘The 
male genitalia seem very marked but there is not enough material 
to dissect them. 


39. Culex tigripes, Grandpré. 


Additional Localities and Dates. 
The Himalayas.—Kurseong, 5,000 feet, E. Himalayas, Dar- 
jiling district, 4-viii-o8 (Annandale). 
Burma.—Mandalay , 11-iii-o8 (Annandale). 


20 Records of the Indian Museum. [Von. IV, 


40. Culex concolor, Desvoidy. 
Mem. Soc. @ Hist. Nat. de Paris, iv, 405. 


Localities and Dates. 

Bengal.—Calcutta (Museum compound), 7-iii-o7, 4-vii-o7, 
5-Vit-07 (6), 15-vii-07, 23-vii-07, 31-vii-o7, 3, 4 and 5-viii-o7 (Annan- 
dale), also in June, September, October and November (Annandale) 
(Zoological Gardens), 12-vii-o8 (2) and 26-vii-o8 (6) (Annandale) ; 
Balliganj near Calcutta (T. Bentham); Port Canning, Lower 
Bengal, December; Purneah, N. Bengal, 4—6-viii-o7 (C. Paiva) 
Damukdia Ghat, R. Ganges, July. 

Assam.—Sylhet, February, April, May, December (Lt.-Col. 
Hall); Manipur, August (C. A. Gourlay). 

Travancore State—Kulattupuzha (w. base of W. Ghats), 
19-x1-08, “in bungalow” (Annandale). 


’ 


. 
, 


41. Leucomyia gelida, Theob. 
Mono. Cultcid., ii, p. 21, IgoT. 


Additional Localities and Dates. 

Bengal.—Calcutta (Museum premises), 3—4-vili-07, 5-Vviil, 
7-vili and 9-viii-o7 (Annandale); Bhogaon, Purneah district, N. 
Bengal, 30-ix-08 (5), I-x-08 (2), 3-x-08(2), 7-x-08 (C. Pavva). 

FE. Bengal.—Rajshahi, 1—6-i1-07 (.4unandale). 

Burma.—Rangoon, 25-11-08 (Annandale). 

Madras Presidency.—Madras town, 30-x-08 (2), 31-x-08 (1) 
(Hodgart). 

Travancore and Cochin States —Kulattupuzha (w. base of W. 
Ghats); 19-xi-o8, coastal region, 5-xi-o8; Ernakulam, 4-xi-08 
(Annandale). 


42. Leucomvia gelida, var. cuneata, Theob. 
Mono. Culicid., ii, p. 22, 1901. 


Localities and Dates. 


Bengal.—Balighai near Puri, Orissa, 23-x-08, “‘at light”’ 
(Annandale). 

Travancore State-—Vaikum, coastal region, 5-xi-08; Kulattu- 
puzha (w. base of W. Ghats), 19-xi-08 (Annandale). 


43. Leucomyta sinensis, Theob. 


Syn. L. gelida var. sinensis, Theob. 


Mono. Culicid., iti, p. 180, 1903. 
Bengal.—Balighai near Puri, Orissa, 23-x-08, “‘at light in 
house, becoming active after dark’ (Annandale). 


1910. | F. V. THEOBALD: Report on Culicide. 21 


44. Culicada suknaénsis, sp. nov. 


Thorax ornamented with rich brown and dull golden yellow 
scales, the latter forming a curved line on each side in front and 
two rather indistinct median lines in front, the rich brown showing 
up as four obscure spots. Head dull golden yellow with two dark 
spots. Abdomen deep brown with basal creamy bands and large 
basal creamy lateral spots. Legs brown, pale at the base, with 
narrow basal yellow bands. Wings with a brownish yellow tinge. 

9. Head dark brown, with small narrow-curved golden scales 
in the middle, dark upright forked scales behind, rich ochreous 
ones in front, a patch of dark flat scales at the sides bordered 
along the eyes with pale creamy scales and flat creamy ones placed 
still more laterally, the pale scales bordering the eyes are small 
narrow-curved ones; cheetee between the eyes bright golden brown, 
followed by some darker ones at the sides; clypeus, palpi and pro- 
boscis deep brown; antenne brown, basal segment bright golden 
yellow, with some small pale flat scales, base of second segment 
pale testaceous. 

Thorax deep rich brown, clothed with golden yellow and rich 
brown curved scales, the former as two obscure curved lateral lines 
and two broad median ones (the latter almost in one) and scattered 
irregularly over the posterior portion; a few paler ones in front of 
the roots of the wings and some short pale golden and brown chete ; 
supra-alar chetee brown; scutellum brown with small narrow- 
curved pale scales; metanotum almost black; pleuree deep brown, 
with patches of pale flat scales and tufts of golden brown cheetze. 

Abdomen deep brown, with basal yellow curved bands, and 
yellow prominent lateral spots; posterior border bristles pale 
golden; venter mostly clothed with large flat yellowish scales, 
almost white towards the base. Legs brown, femora pale ven- 
trally, metatarsi and all the tarsi with narrow pale yellow bands, 
pale knee spots, femora and tibize spinose, spines dusky and golden ; 
ungues all equal and uniserrate. 

Wings with short fork-cells; the first submarginal longer and 
narrower than the second posterior, its stem about two-thirds the 
length of the cell; its base nearly level with that of the second 
posterior cell; stem of the latter as long as the cell; posterior 
cross-vein longer than the mid, nearly twice its own length distant 
from it; the base of the subcostal and first long vein prominently 
densely scaled with black scales. Walteres with ochreous stems, 
fuscous knobs with pale creamy apical scales. 

Lengih 4to 5 mm. 

Habitat Sukna, 500 feet, Darjiling district, E. Himalayas 
(Annandale). 

Time of capture 1 and 2-vii-08. 

Observations.—Described from four perfect @’s. It comes 
near Culicada mpponit, Theobald, but can be told by the absence 
of pale apical abdominal scales and median grey scales and dark 
flat-scaled lateral cephalic patches. The abdomen in all four is 


22 Records of the Indian Museum. [Voryive 


thick and short, but this may be due to ingested blood. The 
ornamentation of the thorax varies in different lights. The dark 
thick-scaled area at the base of the wing is very characteristic. 
The specimens were taken in dense jungle and bit during the day. 


45. Mansontoides annulifera; Theob. 
Mono, Culicid., ii, p. 183, Igor. 
N. Bengal.—Bhogaon, Purneah district, 7-x-08 (9) (C. Patva). 


46. Mansonia unttformis, Theob. 

Mono. Culicid., ii, p. 180, Igot. 

Additional Localities and Dates. 

Bengal.—Calcutta, 2-vii-o7 (mus. colly.); Bhogaon, Purneah 
district, N. Bengal, 30-ix-08 (2), 1-x-08; Katihar, Purneah district, 
N. Bengal, 4—5-x-08 (4) (C. Paiva); Balighai near Puri, Orissa, 
23-x-08 (3) (Annandale). 

Burma.—Rangoon, 25-11-08 (Annandale). 

Travancore and Cochin States.—Maddathorai (w. base of W. 
Ghats), I9-xi-08; Kulattupuzha (w. base of W. Ghats), 19-xi-08 ; 
Ernakulam, Cochin State, 4-xi-08 (8), ‘‘ very common in bungalow”’ 
(Annandale). 

47. LTeaniorhynchus ager, Giles. 


The Entomologist, p. 196, July root. 


Additional Localities and Dates. 

Bengal.—Calcutta, 2-ii-o8 (Annandale); Balighai near Puri, 
Orissa, 26-x-08 (2) (Annandale). 

Burma.—Mandalay, 12-11-08 (Annandale). 

Travancore State-—Pallode, twenty miles N. E. of Trivan- 
drum, 14-xio8; Kerumadi, s. end of Vimbanad Lake, 6-xi-o8 
(Annandale). 

48. Teamniorhynchus tenax, Theob. 


Mono. Culicid., ii, p. 198, Igor. 
Bengal.—Balighai near Puri, Orissa, 23-x-08, one @ “‘at 
light’’ (Annandale). Rather smaller than the type. 


49. Teniorhynchus epidesmus, sp. nov. 


Thorax brown, mottled with creamy scales and some grey 
ones in front of the roots of the wings ; proboscis pale with a nar- 
row black base and broader black apex. Abdomen black with 
very narrow golden ochraceous basal bands and very broad apical 
ones, a dark lateral line on each side of the segments, most pro- 
nounced on the apical ones, the penultimate all golden ochreous, 
the apical with two deep black spots. Legs ochreous brown with 
apical and basal pale banding on the darker tarsi. Wings with a 


IgIo. | F. V. THEOBALD: Report on Culicide. 23 


yellowish tinge, brownish scales and a pale spot towards the tip 
on the costa. 

?. Head brown, clothed with pale creamy and ochreous nar- 
row-curved scales and broadly expanded black upright forked 
scales, paler in the middle of the head, chetze brown and pale 
ochreous; clypeus ochreous; palpi black-scaled with some 
ochreous scales and creamy scaled apices; proboscis with a very 
broad ochreous pale band, base and apex black; antenne brown 
towards the apices, testaceous towards the base. 

Thorax deep brown, clothed with creamy and brown narrow- 
curved scales, some paler before the roots of the wings, giving 
it a mot'led appearance; some short thick black chet just be- 
fore the roots of the wings, long brown and ochreous ones over 
the roots and behind them; scutellum brown, with small narrow 
pale scales and golden brown long border bristles; metanotum 
pale ochreous with a grey sheen; pleure brown and ochreous 
with patches of flat pale scales and golden hairs. 

Abdomen brown with violet reflections, basal segment golden 
ochreous with two small dark patches of scales; the remaining 
segments with natrow basal and very broad apical bright golden 
ochreous bands, except the last segment which has two promi- 
nent black spots; the other segments have more or less marked 
thin black lateral lines, except on the last two or three apical seg- 
ments where there are pale creamy lateral spots; venter ochreous- 
vellow. Legs ochreous with scattered dark scales, prominent at 
the apices of the femora and tibice and on their upper surfaces; 
tarsi slightly darker with apical and basal pale bands except on 
the last tarsal; ungues equal and simple. 

Wings with brown scales, a pale ochreous spot towards the tip 
on the costa and first long vein; the first submarginal cell longer 
and narrower than the second posterior, its base slightly nearer 
the base of the wing, its stem nearly one-fourth the length of the 
cell; stem of the second posterior not quite half the length of the 
cell; posterior cross-vein about twice its own length distant from 
the mid. MHalteres pale ochreous, with some small flat pale 
ochreous scales on the knob. 

Length 5°5 mm. 

Habitat Bhogaon, Purneah district, N. Bengal (C. Patva). 

Time of capture 2-x-o8. 

Observations.—Described from a single perfect female. The 
very bright and marked abdominal banding will at once separate 
this from other Teniorhynchi, together with the marbled thorax 


50. Teniorhynchus lutcoabdominalts, sp. nov. 


Thorax brown, marbled with bright yellowish brown and grey 
scales on the front two-thirds, the rest bright brown-scaled, the 
pale scales most prominent on the posterior edge of the brown and 
pale-scaled area, also traces of a dark curved lateral line on each 
side before the wings. Head yellowish brown, slightly darkened 


24 Records of the Indian Museun. [Vor. IV, 


at the sides, pale around the eyes; proboscis with a broad pale 
band, slightly darkened at the base and apex. Abdomen entirely 
clothed with golden ochreous scales. Legs unbanded, ochreous 
with slightly darkened tarsi. Wings with yellowish brown scales, 
the apex paler above. ’ 

?. Head brown, clothed with narrow-curved pale scales, 
especially around the eye border, bright ochreous upright forked 
scales in the middle, darker ones at the sides ; small flat pale lat- 
eral scales; clypeus brown; palpi clothed with brown scales, a few 
pale ones dotted about and many at the apex; proboscis with a 
broad pale band, the dark base and apex with a few scattered pale 
scales; antenne brown, basal half gradually becoming bright 
testaceous. Thorax brown, the front two-thirds mainly clothed 
with very pale grey narrow-curved scales, particularly prominent 
on the posterior border of this region, two ochreous-brown-scaled 
spots in front amongst the pale scales and a somewhat nude dark 
curved line on each side in front of the roots of the wings, the 
hinder region of the mesothorax with mostly golden brown narrow- 
curved scales, but some pale ones arranged in lines; two promi- 
nent lines of rich golden brown cheetze behind and similar coloured 
cheetee over the roots of the wings; scutellum pale ochreous with 
small narrow-curved pale scales and nine bright golden brown 
posterior border bristles to the mid lobe; metanotum brown; 
pleuree brown and bright ochreous with some patches of small dull 
creamy flat scales. 

Abdomen clothed with bright golden ochreous scales, the 
second, third and fourth segments with a band towards their bases 
of similar coloured scales, darker at their apices, not forming dis- 
tinct bands, last segment with two median lateral dark spots; 
venter all golden ochraceous, also the whole of the first segment 
which has many pale golden thin hairs; posterior border bristles 
pale golden. 

Legs uniformly ochreous but with some dark scattered scales 
on the femora, with very pale reflections, no trace of banding ; 
ungues equal and simple. 

Wings with yellowish brown scales, the outer part of the apex 
slightly pale-scaled ; first submarginal cell longer and narrower than 
the second posterior cell, its base nearer the base of the wing, its 
stem not quite half the length of the cell, stem of the second pos- 
terior nearly two-thirds the length of the cell; posterior cross-vein 
about twice its own length distant from the mid; halteres very 
pale ochreous, almost creamy white. 

Length 5°8 mm. 

Habitat Katihar, Purneah district, N. Bengal (C. Patva). 

Time of capture 4 or 5-x-08. 

Observations.—Described from a single perfect @. It comes 
very near Tentorhynchus epidesmus, described previously, but the 
total absence of leg banding, the more marked thoracic ornamenta- 
tion and more marked pale wing spot separate it probably from 
that species. If it is the same it is a most marked variety. 


1910. ] F. V. THEOBALD: Report on Culicide 


No 
On 


51. Chrysoconops pygmeus, Theobald. Pls. i and iii, wing and 
wing scales. 


Rec. Ind. Mus., ii, p. 300, 1908, @. 


@”. Head ochreous-brown with small narrow-curved golden 
scales and rich golden brown upright forked scales and golden 
cheetze; palpi and proboscis black, the apical segment a little 
more than half the length of the penultimate, bluntly acuminate. 
long dark hairs on the last two segments. Antenne brown with 
white internodes and dark brown plume hairs. Thorax shiny 
bright brown with narrow-curved golden scales and golden cheetze. 
Abdomen ochreous when pointed from the light, dark when to the 
light, of violet hue, the latter showing only on the apical borders 
in some lights. Genitalia with the claspers curved on themselves, 
broader and flat apically with a short, thick subapical dark spine. 
Legs uniformly dark brown with ochreous reflections, pale at their 
bases; fore ungues unequal, the larger with a small lateral basal 
tooth and a very large one near the middle, the smaller simple ; 
mid ungues unequal, the smaller simple, the larger with a large 
basal tooth and a narrower and more basal central one; hind 
ungues equal and simple. Wings with short fork-cells, the first 
longer and narrower than the second, its base nearer the apex of 
the wing, its stem as long as the cell; stem of the second posterior 
as long as the cell; posterior cross-vein as long as the mid, about 
twice its own length distant from it. 

Length 5 mm. 

Habitat Purneah, N. Bengal (C. Paiva); Rajmahal, Bengal 
(R. Hodgart) ; and Calcutta (Annandale). 

Time of capture 6-viii-o7 31-vii-07, 17-vii-07. 

Observatsons.—The 2 was described in the first report. One 
@ has been retained as a co-type for the British Museum collection. 

Dr. Annandale says the eyes of the male are iridescent 
green in life. 


52. Lophoceratomyta bicornuta, sp. nov. 


Head of male very similar to L. fraudatrix but the antennal 
organs quite distinct. Thorax rich brown. Abdomen dark brown 
unbanded, ochreous ventrally. Tegs brown unbanded, base of 
femora paler. Two prominent lateral horn-like processes on the 
basal segments of the antenne. 

». Head similar to that of L. fraudatrix. Palpi brown, a 
little longer than the proboscis, bluntly acuminate, the apical seg- 
ment slightly longer than the penultimate, the whole palpi with 
short, scanty hairs; no trace of the basal process seen in /raudatrix. 

Antenne plumose, plume hairs brown, internodes pale, basal 
segment with a large horn-like process on the inner side as in 
brevipalpus, with a series of fine hairs on the upper side; antennal 
organs on four segments. 

Thorax rich brown, showing two median parallel brighter 
brown broad lines; covered scantily with very small curved 


26 Records of the Indian Museum. [VoOL.nVs 


brown to bronzy scales; brown to deep brown chetz; scutellum 
paler with similar scales and four black border bristles to the mid 
lobe; metanotum brown. Abdomen deep brown, unbanded, pale 
hairs; venter ochreous brown. Legs brown, unbanded, fore and 
mid ungues unequal, the former uniserrate, the mid simple; hind 
small, equal and simple. 

Wings with rather short fork-cells, the first submarginal 
longer and narrower than the second posterior, its base very little 
nearer the apex of the wing than that of the latter, its stem 
slightly longer than the cell, stem of the second posterior cell 
longer than the cell; mid cross-vein longer than the supernumerary, 
the posterior longer than the mid, not quite twice its own length 
distant from it. 

Length 4°5 mm. 

Habitat Dawna Hills (base) near Kawkareik, Lower Burma. 

Time of capture 4-iii-o8 (Annandale). 

Observations.—Described from a single dissected ~. Closely 
related to L. fraudatrix, Theobald, but differs in the antennal 
organs, the absence of the accessory process at the base of the palpi 
and in wing venation. 


53. Radioculex clavipalpus, Theobald. (Pls. 1 and ii, wing and 
head.) 


Rec. Ind. Mus., iti, p. 295, 1908. 


Additional Localities and Dates. 
Bengal.—Katihar, Purneah district, N. Bengal, 4—5-x-08 
(C. Patva). 
Burma.—Rangoon, 25-i1-08 (Annandale). 
Travancore State-—Vaikum, coastal region, 5-xi-o8 (Annan- 
dale). 
54. Uvanotenia testacea, Theobald. 


Ann. Mus. Nat. Hist. Hung., iii, p. 113, 1905. 


Burma.—Base of Dawna Hills, 4-iii-08 (2 2s), “‘ in thick but 
dry jungle”’ (Annandale). 


55. Adeomyia squammipenna, Arribalzaga. 


El, Nat Ageyei 15, 3, 1078: 

Burma.—Base of Dawna Hills, L. Burma, 2-11-08, “‘in bunga- 
low at light ” (Annandale). 

At sea.—At light on board ship four miles off Tuticorin, S. 
India, 25-v-08 (C. Paiva). 


Genus PSEUDOGRAHAMIA, gen. nov 


Head clothed with flat scales; proboscis more than half the 
length of the body, curved upwards, slightly swollen at the apex; 
palpi of the @ very small and densely scaly. 


1910. | F. V. THEOBALD: Report on Culicide. 27 


Thorax densely clothed with large narrow-curved scales, 
closely appressed and with a median line of flat metallic scales, 
and a patch of flat scales on each side in front; scutellum clothed 
with flat scales, large and rounded apically; metanotum appar- 
ently nude. 

Apex of hind tibiz and base of metatarsi with rather dense 
short outstanding scales. First fork-cell slightly expanded basally 
and the veins densely scaled apically, considerably larger than the 
second posterior cell. 

A very marked genus coming near Grahamia, Theobald, but at 
once distinguished from it by the more normal form of proboscis, 


56. Pseudograhamia aureoventer, sp. nov. 


Head black, with a central silvery white spot; proboscis and 
antenne deep brown; palpi pale creamy, very small. Thorax 
black with a broad silvery white line in front, silvery white pro- 
thoracic lobes and a creamy patch in front at the sides. Abdomen 
black with large basal triangular creamy lateral patches which 
extend upwards almost to form narrow dorsal bands; venter pale 
dull golden. T,egs deep black. 

@. Head covered with flat closely appressed blackish brown 
scales with a large median triangular silvery white area with two 
long black straight cheete projecting forwards between the eves, a 
few small black ones curved inwards on each side and a large black 
one laterally, along the ocellar rim; proboscis deep black, curved 
upwards, slightly swollen apically; palpi very small, densely 
clothed with creamy scales; antennee deep brown; clypeus bright 
rich brown. 

Thorax deep rich brown, densely clothed with long narrow- 
curved closely appressed deep blackish brown scales, a- median line 
in front of flat scales, semi-transparent, with white and pale 
mauve and silvery reflections, a large area of flat dull creamy to 
ochreous scales on each side in front reaching up to the base of 
the wings; prothoracic lobes with flat silvery white scales with 
pale blue reflections in certain lights; scutellum clothed with large 
flat silvery white scales, some appearing dusky on the lateral 
lobes; metanotum deep brown; pleuree brown with flat creamy 
scales. 

Abdomen flattened laterally with large basal silvery white 
lateral patches, extending upwards to the dorsum as narrow areas ; 
base and venter of the abdomen pale dull golden, with metallic 
sheen. 

Legs deep blackish brown; coxe pale ochreous with white 
scales; femora pale ochreous below; tibiae with the scales ex- 
panded apically and with some incurved cheetze ; base of metatarsi 
with the scales slightly projecting, particularly on the hind legs; 
ungues equal and simple. 

Wings with moderately long fork-cells; the first submarginal 
much longer and about the same width as the second fork-cell, its 


28 Records of the Indian Museum. [VOnrsLyE 


base much nearer the base of the wings than that of the latter, its 
stem half the length of the cell; the cell somewhat expanded 
basally and each branch densely scaled apically, stem of the second 
fork-cell longer than the cell; supernumerary cross-vein longer 
than the mid, the mid the same length as the posterior; the latter 
about its own length distant from the mid. 

Halteres with long stem, base ochreous, apex and knob 
fuscous. 

Length 4°8 mm. 

Halttat Pallode, twenty miles N.E. of Trivandrum, Travan- 
core (Annandale). 

Time of capture 16-xi-o8. 

Observations.—Described from a single @. A very marked and 
beautiful species which cannot be confused with any other Culicid. 


Genus SQUAMOMYVIA, gen. nov. 


Head clothed with flat scales and upright forked ones behind; 
clypeus densely scaled, scales long and broad; antenne of male 
plumose; palpi small and thin, proboscis long and thin, not as 
long as the body. 

Thorax with spindle-shaped scales, small flat ones at the sides 
in front; broad spindle-shaped ones on the prothoracic lobes, 
densely covering them; scutellum with flat scales. Wings of male 
with short fork-cells. Abdomen clothed with very large flat scales, 
somewhat loosely applied. 

This forms a very distinct genus easily told by the densely 
scaled clypeus. The male genitalia also seem very marked, but 
have not been dissected. 

A male only occurs in the collection. 


57. Squamomyra tnornata, sp. nov. 


Head deep brown, a pale border around the eyes; clypeus 
prominently pale-scaled ; palpi, proboscis and antenne deep brown. 
Thorax deep brown, grey in front and at the sides in front, pleure 
white. Abdomen deep blackish brown above, white below and at 
the sides. Legs deep brown. 

@. Head clothed with flat brown scales with violet reflections, 
a border of flat white ones around the eyes, spreading out laterally , 
dull ochreous-brown upright forked scales behind; two bright 
golden chetze projecting between the eyes in front, and a few 
short dark ones at the sides; clypeus ochreous with long white 
scales, especially dense in front; proboscis thin, black, not as long 
as the whole body, curved downwards; palpi ochreous, dark- 
scaled above, thin, very short; antenne very pale brown, with 
dark plume hairs; basal segment bright ochreous except on the 
inner side where they are dark. 

Thorax black with bronzy brown spindle-shaped scales, white 
in front near the head and small flat white scales on each side in 


IQI0. | F. V. THEOBALD: Report on Culicide. 29 


front, prothoracic lobes densely clothed with white spitridle-shaped 
scales; scutellum brown with brown flat scales, those at the sides 
with pale ochreous reflections; pleurze black with flat white scales. 

Abdomen narrow, clothed with deep blackish brown scales 
with violet reflections and with white scales laterally and 
ventrally. . 

Legs deep brown with violet reflections, unbanded, coxe pale 
with white scales; venter of femora white; fore and mid ungues 
unequal; simple; hind equal and simple. 

Wings with short fork-cells: the first very little longer, but 
narrower than the second, its base nearer the apex of the wing, 
its stem more than two-thirds the length of the cell, stem of the 
second posterior two-thirds the length of the cell; posterior cross- 
vein as long as the mid and more than its own length distant 
from it. 

Length 5 mm. 

Habitat w. slopes of Dawna Hills, L. Butma, 2,000—3,000 
feet (Annandale). 

Time of capture 2 or 3-11-08. 

Observations.—Described from a single ~. It is a very marked 
Aldine at once told by the densely long-scaled clypeus. 


58. Orthopodomyia maculata, sp. nov. 


Thorax rich brown, ornamented with golden and creamy 
scales; head with pale creamy scales; proboscis with one small 
median white band; palpi of male about three-fourths the length 
of the proboscis, white at base and apex and with two median 
white bands. Abdomen black with basal white bands and two 
median white spots to some segments. Wings with dark scales 
and white spots. Fore and mid legs unbanded, hind with promi- 
nent white bands and last segment all white. 

x». Head brown with narrow-curved and upright forked 
scales of a pale creamy to almost white hue, the fork scales 
numerous and uniformly scattered over the head; antennz with 
pale creamy and brown bands, plume hairs brown to flaxen, basal 
segment black with small broad creamy scales dotted over it; the 
first few segments of the flagellum with long white and creamy 
scales; palpi black, white-scaled at the apex, a few white scales at 
the base and two median white bands, one near the apex, golden 
cheetee at the apex, about three-fourths the length of the proboscis, 
the latter black with one median narrow white band! 

Thorax black, clothed with large narrow-curved creamy scales, 
the majority golden, but some white ones in front, at the sides 
and over the roots of the wings and before the scutellum and some 
areas of dark bronzy ones; cheete long, golden and brown; scutel- 
lum black with long silvery white scales and long golden border 
bristles; metanotum black; pleure brown with patches of flat 
white scales and some long thin ones beneath the wings; pro- 
thoracic lobes with flat white scales. 


30 Records of the Indian Museum. [ Vox. IV, 


Abdomen jet-black with snowy white basal bands and on 
some of the segments two small white median spots; venter black 
with basal white bands and all the bristles golden. 

Legs black, femora and tibiz speckled with pale creamy 
scales, fore legs unbanded, but the apex of the tibiz white; mid 
legs with two pale bands involving the jcints of the metatarsal 
and first tarsal, and first and second tarsals; hind legs with a 
narrow band involving the metatarsal and first tarsal joint, a 
very broad one involving the first and second tarsal joint, a nar- 
rower one the second and third, apex of third white and all the 
fourth. 

Wings with black scales over most of the veins but with 
white spots as follows :—four large ones on the costa and three small 
basal ones; the first two spread on to the first and second veins, 
the third on to the first, second, third and fourth, the fourth on to 
the first and second, the basal ones small and irregular, there is also 
a spot on the first between the third and fourth costal spots, one 
on the base of the second fork-cell and at the tips of the branches, 
one at the apex and another near the base of the outer branch of 
the fifth and one at its base, and a small one just past the cross- 
vein on the fourth. Halteres all pale. Fork-cells rather short, 
the first narrower and about the same length as the second, its 
stem about half as long again as the cell; stem of the second as 
long as the cell; cross-veins pale, the posterior about twice its own 
length distant from the mid. 

Length 5 mm. 

Habitat Maddathorai, w. base of W. Ghats, Travancore 
(Annandale). 

Time of capture 17-x1-08. 

Observations.—Described from a single ~ , taken resting on a 
tree trunk in jungle. A very marked and beautiful species near 
O. albipes, Theobald, but easily told by having only the last hind 
tarsal white. 


59. Mimomyia minuta, Theobald. (Pls. i and iii, wing and wing 
scales.) 


Rec. Ind. Mus., ii, p. 301 (@), 1908. 

Thorax shiny brown; head pale creamy yellow; abdomen 
brown with basal pale yellow bands spreading out laterally. 
Legs with apical and basal pale banding; last hind tarsal pale 
creamy. 

2. Head clothed with flat pale creamy scales and some black 
upright forked ones at the back; two long pale cheete projecting 
forwards in the middle in front and some dark incurved ones at 
the sides; antennee deep brown, basal segment black. 

Thorax shiny black, with scattered long curved black scales 
and very long black cheete; scutellum shiny black with long 
curved black scales and black border bristles; metanotum black; 
pleuree ochreous. 


IQI0. | F. V. THEOBALD: Report on Culicida. 31 


Abdomen ochreous with deep brown scales, rather scanty 
basally giving a false banded appearance, with basal lateral areas 
of pale scales; border bristles pale. 

Legs brown to almost black; coxee and venter of femora pale ; 
apex of femora and tibie white, the remaining segments with apt 
cal and basal pale creamy bands, last hind tarsal with all pale 
scales, in the other legs pale in certain lights. 

Wings with the first fork-cell very little longer and narrower 
than the second, its stem nearly as long as the cell; stem of the 
second fork-cell not quite as long as the cell ; posterior cross-vein 
longer than the mid, rather more than its own length distant from 
it; scales large and brown. 

Length 2°5 mm. 

Habitat Calcutta (Annandale). 

Time of capture 30-vii and 3 or 4-vill-07. 

Observations.—One @ andone ~. ‘The o was described from 
Sylhet, Assam (Records Indian Museum, vol. ii, p. 301, 1908). In 
the description of the proboscis it reads as if it were long and thin 
as in Culex; this is not so, it is very swollen apically, almost club- 
shaped in some views, with a long thin stem. 


Genus BOLBODEOMYIA, gen. nov. 


Head clothed with flat scales; palpi very short in the 2 , 
slightly longer in the ~, antennz pilose in both sexes; thorax 
with spindle-shaped scales; prothoracic lobes with small flat scales, 
scutellum with flat scales; metanotum with a bunch of cheete 
posteriorly. Wings with normal venation with rather long lateral 
vein scales on the apical areas of the veins and with very promt 
nent long dense lateral vein scales on the fifth vein, along the stem 
and one branch. Complex ~ genitalia, the claspers ending in a 
bulbous process. A very marked genus. 


60. Bolbodeomyia complex, sp. nov. 


Head, thorax, legs, antennz and palpi and proboscis brown. 
Abdomen brown above, pale grey below in @ , ochreous in the o. 

@. Head clothed with flat brown scales with violet reflections 
and a pale creamy border around the eyes and at the sides; cheetze 
dark; clypeus, palpi and proboscis deep brown, palpi very small ; 
antenne deep brown, basal segment brown, pilose. 

Thorax shiny deep brown with scattered brown and paler 
spindle-shaped scales; prothoracic lobes covered with small flat 
creamy scales, scutellum ochreous-brown with flat dull scales; 
metanotum brown with a large area of pale hairs posteriorly, aris- 
ing from black points; pleuree ochreous-brown with flat white 
scales. Abdomen deep brown, traces of some pale scales at the 
bases of the apical segments; venter pale grey, the integument 
ochreous. 

Legs deep brown, coxz ochreous with white scales, venter of 
femora pale. 


NO 


Records of the Indian Museum. [VOE,; AV, 


Oo 


Wings with short fork-cells, the first longer, but slightly nar- 
rower than the second, its base slightly nearer the base of the 
wing, its stem two-thirds the length of the cell; stem of the 
second as long as the cell; posterior cross-vein about two-and-a- 
half times its own length distant from the mid. 

Male genitalia very complex ; the clasper thick, long, ending 
in a rounded knob, with a group of small curved hairs; lateral 
process at apex short, very thick and dark; a blunt long, finger- 
like process from near base of clasper; on the inner corner of the 
basal lobes four flat cheetee. 

Length 3°58 mm. 

@. Very similar to the male, but rather more white scales in 
front of the head, and the palpi smaller. 

The abdomen has some traces of basal white lateral spots and 
the venter has some narrow apical black bands. Wings with the 
first fork-cell slightly longer and about the same width as the 
second, its stem about half the length of the cell; stem of the 
second rather more than half the length of the cell; posterior 
cross-vein rather more than twice its own length distant from the 
mid, some long lateral vein scales, the fifth long vein, except on its 
upper branch, densely clothed with long thin scales. 

Length 3 mm. 

Halitat Dawna Hills, 2,000—3\000 feet, I,, Burma (7), and 
jungle at base of Dawna Hills ( ¢ ) (Annandale). 

Time of capture I and 2 or 3-11i-08. 

Observations.—Described from one ~ and one 2 , practically 
perfect, but with slightly rubbed thorax. A small obscure A‘dine, 
but at once told when examined microscopically by the complex 
male genitalia and the long lateral scales along one side of the 
fifth vein. 


61. Skusea pseudodiurna, sp. nov. 


Head dusky or pale according to the light. Thorax bright 
reddish brown. Abdomen with basal white lateral spots. Legs 
brown unbanded. 

@”. Head dark, clothed with a mixture of flat dusky and pale 
grey scales and numerous dark upright forked scales and many 
black cheetee projecting forwards; antennz with dark plume hairs, 
and pale grey internodes, dusky at the tip; palpi very small, dark 
brown; proboscis deep brown. Thorax bright deep reddish brown, 
with scanty, scattered bronzy narrow-curved scales and with black 
cheetee, especially thick over the roots of the wings; scutellum 
testaceous with narrow-curved dull creamy and bronzy scales to 
the mid lobe, dark ones to the lateral lobes; metanotum bright 
chestnut-brown. 

Abdomen black with basal white lateral spots, most promi- 
nent on the last few segments, with pale brown border bristles; 
venter dusky. Legs uniformly deep brown, except the coxee and 
venter of the femora which are pale; fore and mid ungues unequal 
and uniserrate ; hind equal and simple. Wings with short fork- 


1gIo. | F. V. THEOBALD: Report on Culicide. 33 


cells, the first longer and narrower than the second, stem of the 
first nearly as long as the cell, stem of the second longer than the 
cell; posterior cross-vein its own length distant from the mid. 

Length 4 mm. 

Habitat Sukna, base of E. Himalayas, 500 feet, Darjiling 
district (Annandale). 

Time of capture 1-vii-o8. 

Observations.—Described from a single ~. Very near Skusea 
diurna, but the bright reddish brown thorax at once separates it. 


62. Skusea uniformis, sp. nov. 


Head black with a pale border around the eyes. Thorax rich 
brown. 

Abdomen deep brown, unbanded, dull ochreous below. Legs 
uniformly deep brown, but the femora pale ochreous below. 

@. Head black with small flat black scales, except for a pale 
area around the eyes and at the sides and some dark upright 
forked scales; clypeus, palpi and proboscis black; antennze deep 
brown, basal segment black, base of segment pale ochreous. 
Thorax rich deep brown with narrow-curved bronzy brown scales 
scattered over its surface; scutellum shiny black with narrow- 
curved bronzy black scales, the surface pale in some lights, four 
dark median posterior border bristles; metanotum chestnut and 
dull brown; pleurze brown with dull flat white scales. 

Abdomen deep brown with dull brown scales, dull ochreous 
to grey-scaled venter. 

Legs uniformly deep brown except the coxe and venter of 
femora which are ochreous; ungues equal and simple. 

Wings with short fork-cells, the first slightly longer and nar- 
rower than the second posterior, their bases about level, the stem 
about two-thirds as long as the cell; stem of the second posterior 
as long as the cell ; posterior cross-vein longer than the mid, about 
one-and-a-half times its own length distant from it. 

Length 4 mm. 

Habitat Pallode, twenty miles N.E. of Trivandrum, Travan- 
core (Annandale). 

Time of capture 15-xi-08. 

Observations.—Described from a single @. The abdomen 
somewhat denuded, but there is no trace of any pale scales. 


eee et 


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Bk onL ule pordk eae UD me aN eh 
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amitiaat + 06H 08 s Abe aggre “Bs bs sand fi 
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WPS 


CHRYSOCONOPS PYGMEUS. N.SP. (9 


MIMOMYIA MINUTA. N.SP 


WINGS OF NEW INDIAN CULICIDA. 


—_—«ase> 
_ —— 


WING OF BREVIRHYNCHUS MAGNUS. N.SP, ( OE) 


HEAD OF RADIOCULEX CLAVIPALPUS. N.SP. 


WING OF LEICESTERIA API \LIs. N.SP. (92). 


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STEGOMYIA TRIPUNCTATA, N. SP. (9) LEICESTERIA APICALIS, N. SP. 


(9) CHRYSOCONOPS PYGMAEUS, N. SP. (%) 


STEGOMYIA ALBOLATERALIS, N. SP. 


ENLARGED WING SCALES. 


Pee tHE -INDIAN SPECIES. On, PAPATACI 
REVItIPHLEBOGROMUSY). 


By N. ANNANDALE, D.Sc., Superintendent, Indian Museum. 


INTRODUCTION. 


The importance of the genus Phlebotomus from a medical 
point of view has recently been demonstrated by Grassi and others 
so far as southern Europe is concerned, and there can be little 
doubt that in India too fevers of a certain type are carried by these 
flies, which, in common with several others, are commonly known 
as “ sand-flies”’ in this country. ‘The descriptions of the species 
described below are based on specimens in the collection of the 
Indian Museum, some of which have been sent me by Mr. F. M. 
Howlett, Second Imperial Zoologist; but I have been able to 
examine all the species except P. papatasi in a living condition. 
The types of all the species except P. papatasi and P. perturbans 
are preserved in the Indian Museum. My thanks are due to 
Mr. E. Brunetti for much assistance. 

The study of the material at my disposal has convinced me 
of the necessity of not relying solely on pinned specimens in 
examining minute and delicate insects such as Phlebotomus, Not 
only do certain structures shrivel and become distorted when 
dried, but it is impossible to make out the exact arrangement of 
others without proper examination under a high power of the 
microscope. So delicate, for example, is the venation of the wing 
of some species of Phlebotomus that its details cannot be seen unless 
the wing is dealt with in the manner in which any similar organ 
‘belonging to any animal but an insect would be dealt with, that is 
to say, unless it is mounted on a glass slide under a cover-slip in a 
suitable medium, after having been artificially stained. Pinned 
specimens even of Phlebotomus are valuable, as showing the natural 
colour and appearance of the insect; but specimens preserved in 
spirit are much more valuable, as enabling structural characters 
to be investigated with greater ease and surety. 

As Mr. Howlett is working at the habits of the Indian 
Phlebotomi,' I need only say that specimens are to be found in 
the corners of bathrooms during the day and round the lamp at 
night ; but a few hints as to their collection and preservation 


1 For full details regarding one Indian species (vtz., P. papatasi) see Grassi’ s 
‘* Ricerche sui Flebotomi ’’ in the memoirs of the Italian Society of Science (1907) 
and also Doerr, Franz and Taussig’s Das Pappatacifiebey (Leipzig and Vienna, 
1909). 


36 Records of the Indian Museum. [Vor.-EV; 


may be useful. The most satisfactory method, so far as my 
experience goes, is the following :—Take a small glass tube full of 
strong spirit and a fine camel’s hair brush. Dip the latter in the 
spirit and lightly touch the Phlebotomus as it rests on a wall or 
table. It will adhere to the brush and may then be washed off 
in the spirit, in which it may be permanently preserved. 

Dried specimens should be pinned in the same way as mos- 
quitoes, but, owing to their small size, this is a difficult process. 
Specimens packed loose in a pill box with tissue paper crumpled 
up not too tightly are, however, useful, provided that there is a 
considerable number of them. 

When the specimens preserved in spirit have to be examined, 
they may be carefully removed to a watch-glass and examined 
superficially under a low power of the microscope. After such 
details have been made out as can be made out in this way, a speci- 
men should be removed to a small stoppered bottle of absolute 
alcohol,’ care being taken that it is not allowed to dry up during 
the process and that the stopper is not left out of the bottle of 
absolute alcohol longer than is necessary. 

After the Phlebotomus has remained in absolute alcohol for 
about half an hour it should be removed to a watch-glass con- 
taining oil of cloves, in which it must be left until it is quite 
transparent. Important details can often be made out while it is _ 
in this watch-glass, for the point of view can be readily altered. 
Finally the specimen should be mounted in canada balsam under 
a thin cover-slip in the ordinary way. It will then be possible to 
study the general form and also such structures as the antenne, 
limbs and genitalia with comparative ease. 

To render the wings fit for detailed study, however, it is 
necessary to stain them artificially. While the specimen is in 
spirit in a watch-glass they should be snipped off at the base by 
a fine pair of scissors and carefully transferred to some preparation 
of hemalum or hematoxylin. I find Ehrlich’s acid hematoxylin, 
Delafield’s haematoxylin and Mayer’s hemalum all good for the 
purpose. Although these stains are rapid in their action, it is as 
well to leave the wing in them for about twenty-four hours. After 
this it should be washed in tap-water, removed to 70 % spirit, in 
which it need only remain for a few minutes, and then to absolute 
alcohol. Finally it should be cleared in oil of cloves and mounted 
in canada balsim. Owing to the toughness of the wings, in spite 
of their appareat delicacy, it is not necessary to pass them through 
different strengths of alcohol in the process of dehydration. 


1 In the damp climate of many parts of India absolute alcohol absorbs atmos- 
pheric moisture with great rapidity ; but this may be remedied by keeping the 
bottom of the bottle in which the absolute alcohol is stored covered with a layer 
of burnt (7.e., dehydrated) copper sulphate. The copper sulphate in this condition 
is white ; when it begins to turn green in the alcohol, it is clear that the alcohol 
is absorbing water, which is again being absorbed by the copper sulphate. In 
this case the alcohol should be poured off and freshly burnt copper sulphate placed 
in it; or the old powder may be removed, burnt again and replaced. 


IQ1o. | 


Go 


N. ANNANDALE: Sfectes of Phlebotomus. 37 


LIST OF KNOWN SPECIES OF PHLEBOTOMUS 


AND 


Europe. 


America. 


Africa. 


Asia. 


1786. 
1843. 
1864. 
LOGS. 


1905. 


1906. 


1906. 


1907. 


OF, PAPERS IN WHICH THEY ARE 
DESCRIBED. 


SPECIES. 


Phlebotomus papatast, Scopoli (1, 9) (S. Europe). 
minutus, Rondani (2) 

is mascittit, Grassi (10) (Italy) 
tupuliformis, Meunier (5, 6) (fossil in 


Pd 


» 
Baltic amber). 

Re vexator, Coquillet (8) (Maryland). 

a cruciatus | ,, (8) (Guatemala). 

7% duboscqui, Neveu-Lemaire (7) (Sou- 
dan). 

- papatast, Scopoli (15) (Northern 

| India; ? Java). 

* himaiayensts, sp. nov. (lower Hima- 
layas). 

- malabaricus, sp. nov. (Travancore, S. 
India). 

rv perturbans, Meijere (15) (Java; base 
of Eastern Himalayas). 

a babu, sp. nov. (plains of India). 

re major, sp. nov. (outer Himalayas ; 
Paresnath, W. Bengal). 

argenttpes, Annandale and Brunetti 


(11) (p'ains of India). 


LITBRATURE, 


Scopoli: Delicia faune et flore insubrice, etc. [P. 
papatasi described. | 

Rondani: Ann. Soc. ent. France (11), vol. i, p. 263. 
[P. minutus described and P. papa- 
tas! redescribed. | 

Loew: l*'auna Austriaca, Dipt. 11, p. 630. |A general 
descr.ption of the genus. | 

Kertesz: ‘‘ Psychodide”’ in Katalog der Palaarktischen 
Dipteren,i. {A bibliography of the 
genus. | 

Meunier: Ann. Mus. Hung., ili, p. 254. .[P. tepuli- 
formis described. | 


ie Naturaliste, p.103. [P.ttpuliformis desciibed 
as a type of a new genus (PiA/ebo- 
tomiella) I think unnecessarily. | 


Neveu-Lemaire: Bull. Soc. zool. France, xxxi, p. 64. 
[P. duboscqui described. | 

Coquil et: Lent. News, Philadelphia, Pa., p. 101. [P. 
vexator and P, cruciatus described. | 


38 Records of the Indian Museum. [Vor IVs 


g. 1907. Grassi: Mem. d. Soc. ttal. d. Stet. (ii), xiv, p. 353. 
_ [P. papatasi fully redescribed with 
many figures. | 

1o. 1908. Grassi: Ati. Reale Accad. Lincet (v), vol. xvii, fasc. 12, 
2nd senustre, p. 68. [P. mascittit 
described. ] 

It. 1908. Annandale: Recs Inds Mus. voll ti- peeroiees i 
argentipes described. | 

12. 1909. Doerr, Franz and Taussig: Das Pappatacifieber (Leip- 
zig and Vienna). 

13. Ig09. Austen: African Bloodsucking Flies, pl. 1, fig. 4 
(B. M., London). [P. duboscqua 
figured. | 

14. 1909. Annandale: Nature, vol. 81, p. 518. [P. papatasi 
recorded from India. ] 

15. 1909. Meijere: ‘‘ Blutsaugende Micro-Dipteren,”’ 77d. voor 
Entom., vol. lii, p. 191. [P. pertur- 
bans and P. angustipennts (synonym 
of P. papatasi ?) described. ] 


TAXONOMIC POSITION OF PHLEBOTOMUS 
AND ITS SPECIES. 


FAMILY PSYCHODIDZ. 


Small Nemocera with relatively large wings, which are 
densely covered with scales or hairs. At least six longitudinal 
veins present in addition to the costal (anterior border) and sub- 
costal ; cross-veins few in number, often practically invisible, 
absent from the distal part of the wing. 


Subfamily Phlebotomine. 


Psychodidze in which the second longitudinal vein has either 
two or three branches, which arise at a considerable distance from 
the point at which the main branch of the vein is joined to the 
third longitudinal vein. Female devoid of a horny ovipositor ; the 
external genitalia of the male consisting of at least three pairs of 
appendages in addition to an intromittent organ. Genera—Phle- 
botomus ,* Nemopalpus , Sycorax, Trichomyta, Eatonisca, Diplonema.* 


Subfamily Psychodine. 


Psychodide in which the lowest of the three branches of the 
second longitudinal vein always present is very distinct from the 
two upper ones, and joins the middle branch at a point nearer the 
base of the wing than the apex. Females with a horny ovipositor 
consisting of two longitudinal valves ; external genitalia of the 


male consisting of two pairs of appendages. (Genera—Psychoda ,* 
Pericoma.* 


The genera whose names are marked with a * are known to 
occur in India, They may be distinguished as follows :— 


1910. | N. ANNANDALE: Species of Phlebotomus. 39 


PHLEBOTOMIN 2} 


A. Wings broadly heart-shaped, densely covered 

with scales; mouth not forming a pro- 

boscis as ae .. Diplonema. 
B. Wings relatively narrow, devoid of scales 

except at the base ; mouth parts forming 

a proboscis ae Be .. Phlebotomus. 


E. ‘7 I. 
Rnd, ap 
3rd ef a | FigaA. 
j { 
4th an , 
ag satin 6th 
a 5th , 6th +th 
” 5th 


Fic. 1.—Wing of Culex (after Theobald). Frc. 2.—Wing of Phlebotomus. 


c., costal vein ; sc., subcostal; 1st to 6th, first to sixth longitudinal veins ; 
a, a’, anda’, incrassations (a’ called by Austen the 6th vein, a’ the 8th); y., 
supernumerary cross-vein ; z., mid cross-vein; P., posterior cross-veln ; A., costal 
cell; B., subcostal cell; C., marginal cell; D., first submarginal cell ; E., second 
submarginal cell; F., first posterior cell; G., second posterior cell; J., third pos- 
terior cell; K., anal cell; H., first basal cell; I., second basal cell; L., auxiliary ; 
M., spurious cell, 


PsyCHODINA}— 
A. The third longitudinal vein runs to the apex 
of the wing, dividing the wing into two 
approximately equal halves -- Psychoda, 
B. The third longitudinal vein terminates dis- 
tally at a point below the apex of the wing Pericoma, 


40 Records of the Indian Museum. [Vorzkve 


Wea Some Diagnostic Characters. 


There is some dispute as to the proper nomenclature of the 
venation of the wing of the Psychodide ; but I think it will he 
sufficient for the purposes of this paper if I compare the wing of 
Phicbotomus with that of the Culicide, for not only are the 
Psychodide (and especially Phlebotomus) close'ty allied to that 
family, but more attention has perhaps been paid to the anatomy 
of the Culicidee in recent years than to that of any other group 
of Diptera. I have therefore reproduced the figure of the wing of 
Culex (fig. I) given by Theobald in the first volume (p. 18) of his 
Monograph of the Culicide side by side with one of the wing of 
Phlebotomus (fig. 2) on which the same lettering is used. 

In accordance with Theobald’s figure and with the views of 
many diptero‘ogists, the thickened anterior or upper border of 
the wing is called the costa, while the short parallel vein running 
immediately below it is called the subcosta. The first parallel 
vein that reaches the distal margin of the wing is known as the 
first longitudinal, those posterior to it being numbered in numeri- 
cal succession.. In both genera six longitudinal veins can he 
distinguished. 

The subcosta in Culex turns upwards at the tip and joins the 
costa, while in Phlebotomus it turns downwards and joins the first 
longitudinal. ‘This is a point that will be considered later when the 
cross-veins are discussed. There is no radical difference as regards 
the first longitudinal in the two genera except that in the Psy- 
chodid it turns upwards at the tip and so assumes a superficial 
resemblance to the subcosta in Culex, in which it turns downwards. 
In Culex the second longitudinal forks once, while in Phlebotomus 
‘t forks twice. In both genera the third longitudinal is simple 
and practically straight ; it is much longer in the Psychodid than 
in Culex. The fourth longitudinal is once forked in both genera, 
but the fifth is simple in Phlebotomus and bears a branch in Culex ; 
in both genera the sixth longitudinal is simple and directed down- 
wards at the distal end. 

The cross-veins are easily detected in Culex, but in Phlebotomus 
they are very difficult to see owing to their extreme delicacy. In 
stained preparations, however, their exact position can be detected, 
at any rate in some specimens. ‘The most noticeable difference 
as regards them is that while in Culex the more important 
cross-veins are situated on the distal part of the wing, in Phlebo- 
tomus they are confined entirely to the proximal part. Consider- 
ing first the anterior half of the wing, we find in Culex a minute 
cross-vein (the ‘‘ humeral’ cross-vein, not numbered or lettered 
in the diagrams) joining the subcosta to the costa. In Culex it 
is situated near the base of the wing, while in Phlebotomus it is 
practically at the distal end of the very short subcosta, which there- 
fore appears to be forked. A careful examination shows, however, 
that the lower branch of the apparent fork is really in continuity 
with the main stem of the vein, while the cross-vein proceeds 


1910. | N. ANNANDALE: Species of Phlebotomus. 41 


from it at a distinct angle. In Phlebotomus this cross-vein is of 
extreme delicacy and, unlike the other veins, it is not provided 
with a double line of hairs. 1t can only be seen after prolonged 
staining of the wing, and even then is faint and indistinct. The 
next transverse vein that is apparent joins the base of the second 
longitudinal to the first and is marked x in the diagrams ; it is 
further from the base of the wing in Cz/ex than it is in Phlebotomus. 
Strictly speaking this is not a distinct vein but an upturned 
portion of the lower vein. The base of the third longitudinal in 
both genera ends in a small fork, one branch joining the vein to 
the second longitudinal, the other to the fourth. The former 
branch is called by Theobald the supernumerary cross-vein, the 
latter the mid cross-vein. In Phlebotomus, however, the lower 
branch appears to be in actual continuity wih the vein, while 
the upper one joins it at an angle. This is more distinct in some 
species than in others. In Culex the fourth and sixth longitudi- 
nals do not appear to be joined together at the base; but in 
Phlebotomus the fifth arises from the base of the sixth, as can be 
seen distinctly in some species, while it is connected with the 
fourth by a short cross-vein close to its base. 

The venation of Phlebotomus, taking into consideration the 
above facts, may therefore be said to resemble that of Culex in 
some essential points, but to differ from it mainly in the fact that 
the cross-veins are extremely delicate and are situated exclusively 
on the proximal part of the wing. 

From a systematic point of view, as regards specific differen- 
tiation, the most important feature in the venation is the relative 
proportions between the length of the upper branch of the second 
longitudinal, the distance between the two forks of this vein, and 
that between the proximal! fork and the point at which the vein is 
joined by the mid cross-vein to the third longitudinal. 

In Phlebotomus, but not in some of its allies (e.g., Diploneinay), 
the wing is almost devoid of scales, a few of which occur at the 
extreme base on the costal and lower borders and on those veins 
which reach the base. Each vein except the small cross-vein that 
joins the subcostal to the costal, bears a double row of fine hairs, 
and the edge of the wing is deeply fringed with longer hairs. 


External genitalia— 

The visible female genitalia in Phlebotomus are simple in 
structure, consisting of two pairs (a superior and an inferior pair) of 
compressed, more or less leaf-like appendages covered with sensory 
hairs. They offer in their shape and proportions features. of diag- 
nostic importance but become shrivelled and distorted in dried 
specimens. There is no horny ovipositor such as is found in 
Psychoda and Pericoma. 

The male genitalia are far more complicated and of much 
greater systematic importance. It is possible, moreover, to trace 
their specific differences even in specimens which have been dried, 
but even such specimens should be mounted in canada balsam for 


42 Records of the Indian Museum, [VWou-be 


a proper examination. It is difficult to homologize the different 
appendages exactly with those of other Diptera, because in all 
families great differences, often of no more than specific value, 
occur in different species. It is better, therefore, not to use 
technical terms in describing these appendages. In the male 


u. 


Fig,3. : sl. 
U. 


Diagram of the external male genitalia of Phlebotomus : u., upper or superior 
appendage; ch., chete; f., genital filament; i, intermediate appendage; P., 
intromittent organ; s.l., subgenitai lamella; L., lower or inferior appendage. 


genitalia of Phlebotomus five pairs of organs can be distinguished. 
They are shown in the accompanying diagram (fig. 3) which is a 
generalized sketch of the whole apparatus as seen from the right 
side and does not represent the organs of any one species. The 


Distal joint of superior male appendage of P. argentipes, showing the chete. 


uppermost appendage has two joints, the posterior (distal) of which 
bears a varying number of stout cheetee (fig. 4). The form, position 
and number of these cheetze are valuable diagnostic points. The 
intermediate appendage is morphologically a branch of the superior 
one but has often more than one lobe. ‘The inferior appendage 


IIo. | N. ANNANDALE: Species of Phlebotomus. 43 


never has more than one joint, but is often elbowed; it arises from 
a subgenital plate, which represents the last somite of the abdomen. 
This subgenital plate, which is highly developed in some Psychodide, 
is very inconspicuous in Phlebotomus, owing rather to its mem- 
branous nature than to its small size. In dried specimens it dis- 
appears almost completely. The intromittent organ, which lies 
between the two intermediate appendages, consists of a pair of 
slender, compressed chitinous valves, between which a pair of long, 
slender, chitinous filaments can be protruded. 


Head— 


The head in Phlebotomus is small and ‘round but the anterior 
part is produced into a more or less cylindrical rostrum, which 
overhangs the mouth parts above. ‘The eyes are large, invariably 
dark in colour and more or less emarginate opposite the base of 
the antenne. The antenne are long and filiform, having nor- 
mally sixteen joints, two of which form the scape. ‘They show 
great individual variation as regards the proportions of the joints 
and also differ as regards the exact outline of the last few distal 
joints in the two sexes. The sexual differences are, however, 
small. The palpi are generally stated to have four joints. In 
fresh specimens, however, a minute basal joint can sometimes be 
distinguished. This joint is often difficult to see and appears to 
be imperfectly separated from the others. Including it the number 
of the joints is five. 

For an account of the mouth parts Grassi may be consulted. 
They do not appear to present features of merely diagnostic 
interest and closely resemble those of Culex in general structure. 


THE INDIAN SPECIES OF PHLEBOTOMUS. 
Table of the Indian spectes of Phlebotomus. 


1. Length 2 mm. Dorsum of thorax dark, sides 
yellowish. Wing with the posterior border 
much more strongly arched than the anterior ; 
the anterior branch of the second longitu- 
dinal vein longer than either the distance 
between the two forks of the vein or that 
between the posterior fork and the mid 
cross-vein; the latter distance shorter than 
that between the two forks ee .. P. argenttpes. 

2. Length 3-375 mm. Colour an almost unt- 
form golden grey. Wing with the posterior 
border much more strongly arched than the 
anterior ; the anterior branch of the second 
vein longer than either the distance between 
the two forks of the vein or that between the 
posterior fork and the mid cross-vein; the 
distance between the two forks shorter than 


44 


6, 


P. argentipes, Annandale, Rec. Ind. Mus., ii, p. 101, 


mm. 


the first joint of the tarsus, 


Records of the Indian Museum. 


that between the posterior fork and the mid 
cross-vein 


Length 275-3 mm. Colour dark greyish brown. 


Wing with the posterior margin as a tule 
more strongly arched than the anterior ; the 
length of the anterior branch of the second 
vein, the distance between the two forks and 
that between the posterior fork and the mid 
cross-vein subequal 


purplish tinge. Wing with the posterior 
margin hardly more stronely arched than the 
anterior; the anterior branch of the second 
vein four times as long as the distance 
between the two forks, which is much shorter 
than that between the Cpe fork and the 
mid cross-vein 


Length 1°5 mm. Colour silvery “grey. Wing 


narrow, as a rule divided into two almost equal 
and similar halves by the third longitudinal 
vein; the anterior branch of the second veiu 
shorter than either the distance between the 
two forks or that between the posterior fork 
and the mid cross-vein; the two latter dis- 
tances being nearly equal 


narrow, divided into two equal and similar 
halves by the third longitudinal vein; the 
anterior branch of the second vein nearly three 
times as long as the distance between the 
two forks, twice as long as that between the 


[Vor. LV, 


P.. major. 


. P. perturbans. 
Length 2°5 mm. Colour dark brown with a 


P. malabaricus. 


; .. P, babu. 
Length 2 mm. Colour yellowish grey. Wing 


posterior fork and the mid cross-vein ..P.himalayensts. 


. Length 2-5 mm. Colour yellowish grey. Wing 


narrow, but with the posterior margin as a 
rule more strongly arched than the anterior ; 
the anterior branch of the second vein nearly 
twice as long as the distance between the two 
forks, equal to that between the posterior 
fork and the mid cross-vein 


Phlebotomus argentipes, Annandale and Brunetti. 


. P. papatasi. 


(Pisivehie: age 


wing, and pl. vi, fig. 6, male genitalia.) 


fig. (1908). 


Size and proportions. —Total Wetige in well-preserved specimens 
2mm. Length of wing 1°75 mm.; greatest breadth of wing 0°5 
Hind leg about twice as long as the head and body; the 
femur less than half as long as the tibia and distinctly shorter than 


joints together. 


which is longer than the remaining 


1gI0.| N. ANNANDALE: Sfecies of Phlebotomus. 45 


Colour.—Head (except eyes, which are black) and abdomen 
brown; dorsum of thorax dark brown or blackish; sides of thorax, 
coxe and trochanters yellowish ; legs, antenne and palpi grey ; 
the whole (especially the legs) exhibiting a strong silvery reful- 
gence. Wings smoke-grey, with a strong bluish iridescence on the 
disk and a general silvery refulgence. 

Head.—Rostrum elongate, sausage-shaped, ovoid as seen 
from above. Eyes distinctly emarginate on their internal upper 
border, narrowly separated. Antenne with 14 joints in the 
flagellum, all of which are more or less elongate and each of which 
bears three or four somewhat irregular verticels of stiff, fine hairs ; 
the proximal joint of the scape clearly separated from the head, 
rather slender, short ; distal joint subspherical, bearing a circle of 
suberect, narrow, spatulate, curved scales and (alternating with and 
below the scales) another of stiff bristles. Palpi with five joints ; 
the basal joint short, the second joint more than half as long as 
the third, which is as long as the fourth and fifth together. 

Thovax.—The dorsum is covered with long, stiff, upright, 
curved hairs 

Abdomen slender and cylindrical in both sexes, considerably 
shorter than the wings, covered with rather slender curved hairs 
of irregular length but never very long or stout. 

Wings bluntly pointed; the posterior border much more 
strongly curved than the anterior. ‘The anterior branch of the 
second longitudinal vein much longer than the distance between 
its base and the second fork; the stem of the vein between the 
second fork and the mid cross-vein slightly longer than the 
distance between the two forks; the posterior fork nearly on a level 
with the fork of the fourth vein. 

External genittata.—(@) The superior appendages somewhat 
widely separated from the inferior, which are small, pointed and 
claw-shaped. (@) All the appendages long and apparent. ‘The 
inferior appendages somewhat slender, laterally compressed, 
longer than the basal joint of the superior appendages, obliquely 
truncated, devoid of spines, covered with stout hairs, which form 
a dense brush at the tip of the appendage ; a distinct elbow not 
far from the base of the appendage ; the intermediate appendages 
small, consisting of a stout, bluntly pointed dorsal lobe and a 
small ventral lobe which is a minute, pointed process bearing a 
bunch of bristles at the tip and attached to the ventral margin of 
the dorsal lobe. The superior appendage with the distal joint 
considerably shorter than the proximal one, bearing five stout, 
pointed chete, each nearly as long as the joint, arranged as 
follows :--a single cheta at a point about a third of the length of 
the joint from the base, a pair of slightly unequal cheetee near the 
middle of the joint, and a terminal equal pair. ‘The proximal 
joint slightly longer than the last apparent segment of the abdo- 
men. Intromittent organs relatively long, ensiform, consisting of 
two slender, pointed lateral valves, between which a pair of long 
filaments can be protruded. 


46 Kecords of the Indian Museum. [Vor. TV; 


This species is easily distinguished, except in the case of 
females distended with blood, from all other Indian forms by the 
strong contrast in colour between the dorsal surface and the sides of 
the thorax. It is a very common species in Calcutta at all times of 
the year, but only occasionally. Often it seems to disappear for a 
time (a few weeks or days) and then to reappear again in large 
numbers. It is commonly found in dark corners of bathrooms 
and basements in houses and round the lamp at night, and has the 
habit of biting people’s ankles under the dinner table. 

Distribution.—Probably all over the plains of India. I have 
examined specimens from Calcutta (all times of year), Rajmahal 
(July, February), Asansol (Paiva, February) and Pusa (Howlett) in 
Bengal, and from several localities in the coastal districts of Tra- 
vancore (November). 


Phlebotomus major, sp. nov. (PI. v, fig. 4, wing, and pl. vi, 
fig. 4, male genitalia.) 


Phlebotomus sp., Giles, Gnats or Mosquitoes (and. ed.), p. 5, fig. 2 

(1902). 

Size and proportions.—Total length 3-3°75 mm. Length of 
wing 2°75 mm., greatest breadth 1 mm. Hind leg two-and-a-half 
times as long as the head and body; its femur less than half as 
long as the tibia and considerably shorter than the first joint of 
the tarsus, which is longer than the remaining joints together. 

Colour.—A uniform golden grey with very strong silvery lights ; 
the disk of the wings with a bluish iridescence; eyes black; legs 
sometimes darker than abdomen. 

Head.—Rostrum somewhat conical, short, truncated in front. 
Eyes widely separated, emarginate above. Antenne with the two 
joints of the scape subspherical; those of the flagellum elongate, 
slender, densely covered with soft hairs. Palpi with five joints; 
the basal joint very short, the second, third and fourth subequal, 
shorter than the fifth. 

Thorax much as in P. argentipes. 

Abdomen long and slender, clothed with long recumbent hairs 
and with tufts of longer, upright ones on the dorsal surface. 

Wings rather narrower than in P. argentipfes, pointed, with the 
posterior border much more strongly arched than the anterior. 
The anterior branch of the second longitudinal vein much longer 
than the distance between the two forks, which is considerably 
shorter than the distance between the second fork and the mid 
cross-vein ; the second fork slightly nearer the base of the wing 
than that of the fourth longitudinal vein. 

External genttaha—(o) Superior and inferior appendages 
similar to those of P. argentipes except that they are more slender 
and elongate, the basal joint of the superior appendage being nearly 
twice as long as the last apparent joint of the abdomen; inter- 
mediate appendages devoid of a ventral lobe; valves of the intro- 
mittent organs slender and blunt ; genital filaments well developed. 


1910. | N. ANNANDALE: Sfectes of Phlebotomus. 47 


This species is closely allied to P. argentipes, from which it is 
easily distinguished by its larger size, uniform golden colour, more 
elongate tarsi and male genitalia. 

Distribution.—Outer Himalayas from the base up to 7,000 
or 8,000 feet. Paresnath Hill, Western Bengal. I have examined 
specimens from Naini Tal and Bowali (Kumaon), Kurseong (Dar- 
jiling district), the Nepal Terai, and Paresnath Hill (Chota Nag- 
pur). The fauna of this hill, which is separated by about 180 
miles from the foot-hills of Nepal, the nearest part of the Hima- 
layas, has a distinctly Himalayan facies, and I think that P. 
major may be regarded as the Himalayan representative of P. 
argentipes. All the specimens of the former species that I have 
taken myself have been taken at light in the evening. The only 
one I saw on Paresnath (a male) was taken at an altitude of 
4,300 feet in April. The species appears to be common at Naini 
Tal and Kurseong and probably also at Simla in May and June. 
Dr. A. D. Imms took specimens at Bowali in July. 


Phlebotomus perturbans, Meijere. (Pl. v, fig. 3, wing, and pl. vi, 
figs. 5, 5a, 50, male genitalia.) 


P. perturbans, Metjere, Tijdschr. v. Ent., vol. lii, p. 201, pl. 12, 

fig. 13 (1909). 

Size and proporiions.—Total length 2°5-3 mm. Length of wing 
3°25 mm.; greatest breadth of wing 07 mm. Hind leg 2} times 
as long as head and thorax; the femur a little more than half as 
long as the tibia, slightly longer than the first joint of the tarsus, 
which is distinctly shorter than the other joints together. 

Colour.—Thorax, abdomen, femora, tibiz, tarsi and antenne 
brown, with the usual silvery lights; head (including palpi and 
proboscis), coxee and trochanters yellowish; eyes black. 

Head.—Rostrum short and thick, rounded at the tip. An- 
tennee with the basal joint of the flagellum not much longer than the 
second; the second joint of the scape bearing a circle of curved 
scales. Palpi long, with five joints; the fifth joint the longest ; the 
second and third much longer than the fourth; the first very short. 

Thorax much as in P. argentipes, except that the hairs are 
shorter. 

Abdomen relatively shorter than in P. major, clothed with 
recumbent hairs and with a few upright curved ones on both the 
dorsal and the ventral surfaces. 

Wing bluntly pointed, with the posterior border not much 
more strongly arched than the anterior. The length of the 
anterior branch of the second longitudinal vein, the distance 
between the two forks of this vein and that between the posterior 
fork and the mid cross-vein subequal; the fork of the fourth 
longitudinal distinctly nearer the tip of the wing than the pos- 
terior fork of the second longitudinal. 

Genitalia (2) The superior appendage considerably larger 
than the inferior, both leaf-like and rounded at the tip. 


48 Records of the Indian Museum. [VOLE 


(@) Inferior appendage shorter than the basal joint of the superior 
appendage, slender, bearing at the tip a bunch of long, stout 
bristles. Intermediate appendage without a ventral lobe, slender, 
pointed, with the tip curved. Superior appendage very long, the 
distal joint much shorter than the proximal, bearing four cheete, 
which are less than half as long as the joint and are arranged in 
two pairs—an equal pair at the tip and one in which the two 
cheetze are not equal on the inner surface about half way along 
the joint; the proximal chzetee are blunt, the distal ones pointed. 
Intromittent organ very slender, the valves rounded at the tip; 
the genital filaments well developed. 

This species is easily distinguished from all the other Indian 
forms I have seen except P. malabaricus (to which it bears a close 
superficial resemblance) by its-dark brownish colour. It is abund- 
ant in the jungles at the base of the Eastern Himalayas at the 
beginning of the rains. Large numbers flew to my light in the 
forest bungalow at Sukna (alt. 500 feet) near the Darjiling Hima- 
layan Railway on the evenings of July ist, 2nd and 3rd, 1908. I 
have not seen it elsewhere. 

Distribution.—Base of the Eastern Himalayas (Darjiling dis- 
trict): Java: 


Phlebotomus malabaricus, sp. nov. (Pl. v, fig. 1, head, fig. 2, wing ; 
and pl. vi, fig. 1, male genitalia.) 


Size and proportions.—Total length 2.5 mm. Length of wing 
2 mm., greatest breadth 075 mm. Hind leg 1} times as long as 
the thorax and abdomen; the femur nearly ? the length of the 
tibia, 1% that of the first joint of the tarsus, which is } that of the 
remaining joints together. 

Colour.—Thorax, abdomen and legs (except coxe and 
trochanters) brown with a tinge of purple and with the usual 
silvery lights; wings purplish, strongly iridescent; head, coxe 
and trochanters pale brown; eyes black. 

Head. —Rostrum short, rounded in front. Antenne normal; 
the second joint of the scape pear-shaped, with several circles of 
hairs and scales; the first joint of the flagellum much longer than 
the second. Palpi long, with five joints; the basal joint very 
short, the second shorter than the third or fourth; third and- 
fourth joints subequal, together shorter than the fifth. 

Thorax densely covered with upright curved hairs. 

Abdomen covered with recumbent hairs. 

Wings pointed; the posterior border not much more strongly 
arched than the anterior. ‘The anterior branch of the second longi- 
tudinal vein very long, about four times as long as the distance 
between the two forks of the vein, which is much shorter than that 
between the posterior fork and the mid cross-vein; the fork of the 
fourth longitudinal almost level with the posterior fork of the second. 

Genitalia —(?) Much as in P. perturbans. (o) Inferior 
appendage slender, cylindrical, slightly shorter than the basal 


1910. | N. ANNANDALE: Spfecies of Phlebotomus 49 


joint of the superior appendage, bearing at its tip (in addition to a 
bunch of fine bristles) seven minute, conical, chitinous spines. 
Intermediate appendage without a ventral branch; the dorsal 
branch resembling that o° P. perturbans. Superior appendage with 
the distal joint less than half as long as the proximal one, bearing 
four cheetee, each of which is as long as the joint; one pair of 
cheetze is situated at the tip of the appendage and the other a 
short distance nearer the base of the joint. Intromittent organ 
long and slender; the apices of the valves club-shaped; genital 
filaments well developed. 

This species is closely related to P. ferturbans, the place of 
which it takes in the jungles at the base of the Western Ghats in 
Travancore, where I took numerous specimens at light in Novem- 
ber, 1908. ‘The anterior branch of the second longitudinal vein, 
however, is much longer than itis in P. perturbans and the legs are 
shorter, while the chetz of the male genitalia are much better 
developed. 

Distribution.—Travancore, S. India, below the western slopes 
of the Western Ghats. I took specimens at Nedumangad, Pallode 
and Maddathorai in November. 


Phlebotomus babu, sp. nov. (PI. iv, fig. 1, wing, and pl. vi, figs. 3, 
3a, male genitalia.) 


Phlebotomus sf., Howlett, in Maxwell-Lefroy’s Indian Insect Life, 

P. 559, fig. 358 (1909). 

Size and proporttons.—Total length 2°5 mm. Length of wing 
1°75 mm., greatest breadth of wing 0°25 mm. Hind leg rather 
less than twice as long as the thorax and abdomen: the femur 
slightly shorter than the tibia, which is more than twice as long as 
the first joint of the tarsus; the first joint of the tarsus less than 
half as long as the femur, shorter than the remaining joints 
together by half of its own length. 

Colour.—Silvery grey; the fringes on the wings, the thorax 
and abdomen appear in certain lights to be darker than the wings 
themselves and the limbs: the usual silvery lights present. 

Head.—Rostrum prominent, somewhat arched. Antenne 
with the second joint of the scape bearing several rows of flat 
scales; the joints of the flagellum normal. Palpi with four joints; 
the first three subequal, short; the fourth nearly as long as these 
three together. 

Thorax less tumid above than in P. argenttpes. 

Abdomen rather short, clothed in close-set upright hairs of 
different lengths and with a dense fringe of upwardly curved hairs 
running along each side of the abdomen near the ventral surface. 

Wings narrow, pointed, the posterior border hardly more 
strongly arched than the anterior, the third longitudinal vein 
dividing the wing into two nearly equal and similar halves. ‘The 
anterior branch of the second vein not much shorter than the 
second branch, shorter than the distance between the two forks of 


50 Records of the Indian Museum. [Vor. IV, 


the vein or the distance between the posterior fork and the mid 
cross-vein; the two latter distances nearly equal; the posterior 
fork nearer the base of the wing than the fork of the fourth 
vein. 

Genitalia.—(@) The superior appendage long and narrow, 
somewhat curved; the inferior appendage less than half as long, 
straight. (#7) The inferior appendage club-shaped, distinctly 
elbowed, about as long as the basal joint of the superior append- 
age, bearing (in addition to slender bristles) several chitinous 
spines at the tip and on the ventral surface. Intermediate 
appendage short, blunt, with the ventral lobe represented by a 
short process on its ventral margin. The superior appendage with 
the basal joint stout, about twice as long as the distal joint, which 
bears four pointed equal or subequal chetze; the chete rather 
longer than the joint, situated two at the tip and two a short 
distance from it. 

This species is the smallest with which Iam acquainted. It 
is frequently found together with P. argentipes, from which it is 
easily distinguished by its smaller size, narrower wings and 
uniform grey colour; I have taken it also with P. malabaricus. In 
habits it resembles P. argentipes, than which it appears to be some- 
what more common. 

Distribution.—Probably all over the plains of India. I have 
examined specimens from Rawalpindi (C. W. Mason, September) ; 
from Allahabad (A. D. Imms, October) ; from Rajmahal on the 
Ganges (July) ; from Asansol (February, Paiva); from Purneah and 
Pusa (Howlett) in Northern Bengal ; Calcutta (all times of year) and 
Port Canning, Lower Bengal ; Puri, Orissa ; Rambha in the north- 
east of Madras (March); Trivandrum, Pallode and Maddathorai, 
Travancore (November), and Igatpuri in the Western Ghats, 
Bombay (November). 


Phlebotomus himalayensis, sp. nov. (PI. iv, fig. 2, wing, and pl. vi, 
fig. 7, male genitalia.) 


Size.—Total length 2-255 mm. Length of wing 1I°5 mm. 
greatest breadth of wing 0o'4 mm. 

Colour.—Yellowish grey with the usual silvery lights ; the dor- 
sum of the thorax slightly darker than the sides. 

Head.—Rostrum rounded at the tip, stout and elongated. 
Antennz normal, with two circles of flat scales and one of flattened 
hairs on the second joint of scape. Palpi with five joints ; the 
fifth the longest. 

Thorax and abdomen as in P. babu. 

Wing narrow, obtusely pointed, the posterior margin slightly 
more strongly arched than the anterior ; the anterior branch 
of the second longitudinal vein not much shorter than the second 
branch, at least four times as long as the distance between the two 
forks, which is about half as long as that between the posterior 
fork and the mid cross-vein ; the posterior fork and the fork of 


1910. | N. ANNANDALE: Sfectes of Phlebotomus. 51 


the fourth longitudinal almost level, the former being slightly 
nearer the tip of the wing than the latter. 

Genttalia.—( ? ) Appendages short and broad, the superior not 
much longer than the inferior. (@) Inferior appendage slender, 
distinctly elbowed, rounded and slightly clubbed at the tip, nearly 
as long as basal joint of superior appendages, devoid of chitinous 
spines. Intermediate appendages simple, pointed, curved down- 
wards at the tip. Superior appendages with the distal joint about 
half as long as the proximal, bearing at the tip three pointed 
cheetee, each of which is slightly shorter than the joint. Intromit- 
tent organ with two elongate, slender valves, which are truncated 
at the tip. 

This species resembles P. babu in general appearance but is 
somewhat larger and yellower. The wing is easily distinguished 
from that of P. babu by the great length of the anterior branch 
of the second longitudinal vein. P. himalayensis frequently occurs 
together with P. major, just as P. babu does with P. argentipes. 

Distribution.—The Himalayas, between 4,000 and 7,000 feet. 
I have examined specimens from Naini Tal and Bowali in Kumaon 
(A. D. Imms, R. E. Llovd) and Kurseong (Darjiling district). It is 
apparently common in both districts in May, June and July. 


Phlebotomus papatasi, Scopoli. (Pl. iv, fig. 4, wing, and pl. vi, fig. 
2, male genitalia.) 


P. papatasi, Grassi, Mem. d. soc. ital. d. Sct. (111), xiv, p. 353 (1907). 
Phlebotomus sf. (o genitalia), Howlett, in Maxwell-Lefroy’s Indian 

Insect Life, p. 560, fig. 359 (1909). 
>? P. angustipennis, Metjere, Tijdschr. v. Ent., vol. lil, p. 202, 

pl. 12, fig. 14 (1909). 

Size and proportions.—Total length 2°55 mm. Length of wing 
2°25 mm.; greatest breadth 0-4 mm. Hind leg more than twice 
as long as thorax and abdomen; the femur a little less than ? 
the length of the tibia, which is nearly twice the length of the first 
joint of the tarsus ; the first joint of the tarsus in the anterior 
legs distinctly shorter than the remaining joints together. 

Colour pale yellowish grey. 

Head.—Rostrum somewhat slender. Antenne normal ; the 
basal joint slender, the second joint of the scape subspherical, 
with a circle of scales and two of flattened hairs. 

Palpi with (?) five joints; the basal joint indistinct, the 
second, third and fifth subequal, each longer than the fourth. 

Thorax and abdomen offering no apparent peculiarities. 

Wings narrow, pointed, with the posterior border markedly 
more arched than the anterior. ‘The anterior branch of the second 
longitudinal vein much shorter than the second branch; the former 
longer than the distance between the two forks, shorter than that 
between the posterior fork and the mid cross-vein ; the posterior 
fork slightly nearer the base of the wing than the fork of the 
fourth longitudinal. 


52 Records of the Indian Museum. [Vot. IV, 1910.] 


Genitalia.—( 2) The superior and inferior appendages short, 
subequal. (@) Inferior appendages slender, shorter than the 
basal joint of the superior appendages, bearing about six short 
chitinous spines at the tip. Intermediate appendages with three 
lobes, the proximal lobe slender, curved, filiform ; the middle lobe 
longer than either of the other two, slender, arched, with a fringe 
of hairs on the concave posterior margin ; the distal lobe concave 
above, pointed, clothed with fine hairs. ‘The superior appendage 
with the distal joint rather longer than the proximal one, bearing 
five short, blunt cheetze, three at the tip (the middle one being 
shorter than that at either side of it) and two, one nearer the base 
than the other, on the middle part of the joint. The intromittent 
organ very short, pointed, the genital filaments well developed. 

By the kindness of Dr. J. H. Ashworth, I have been able to 
compare Indian specimens with specimens from S. Europe (see 
Nature, vol. 81, p. 518, 1909). 

Meijere’s P. angustipennis, so far as is shown by his figure of 
the wing and his brief description, only differs from P. papatasi as 
regards the outline of the posterior border of the wing and the 
longitudinal vein adjacent to it. In some species (e.g., P. babu) 
this is a variable character.! 

Distribution.—Southern Europe and Northern India (plains) ; 
? N. Africa. I have seen a number of specimens from Rawalpindi 
(collected by Mr. C. W. Mason in September) and a single male 
from Pusa, Bihar, Northern Bengal (Howlett). The Rawalpindi 
specimens were accompanied by specimens of P. babu and were for 
the most part infected by a parasitic fungus. 

The importance of this species from a medical point of view is 
expounded in Doerr, Franz and Taussig’s Das Pappatacifieber 
(1909). 


1 It would perhaps be more correct to say that this part of the wing is 
peculiarly liable to distortion in preserved specimens. A comparison of the differ- 
ent figures of the wing of P. papatass that have been published proves very consi- 
derable apparent variation in this respect. 


, 
. . 
« . , 
hi 7 . 
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, . 
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o : 
: ’ = 2 
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all 
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7. : : > Ligh “SS gt fs : 
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EXPLANATION OF PLATE. 1Ve 


WINGS OF INDIAN PHLEBOTOMI. 


Fic. 1.—Phlebotomus babu. 


2.— a himalayensts. 
{— A argenti pes. 


Ali 3 papatast (Rawalpindi). 


Rec. Ind. Mus.,Vol.1V,1910 


3x 72. 


EXPLANATION OF PLATE V. 


WINGS AND HEAD OF INDIAN PHLEBOTOMI. 


Fic. 1.—Head of Phlebotomus malabaricus, from in front 


33 


2.—Wing of ¥ 5 
perturbans. 


major. 


ae ” ” ”) 


gs ace) ” ”? 


Rec. Ind.Mus.,Vol. IV, 1910. - Plate V. 


BiG 


Bo 6 S. 


EXPLANATION OF PLATE VI. 
EXTERNAL MALE GENITALIA OF INDIAN PHLEBOTOMI. 


(The sub-genital lamella is omitted in all the figures and the append- 
ages have been denuded of fine hatrs.) 


Fic. 1.—Male genitalia of Phlebotomus malabartcus. 


Pie Dasa gers 5 a papatast. 

en Bemete Ss; be re babu (cheetee removed). 

Jo omy 1555 3 5 a ,, (extremity of dis- 
tal joint, showing 
the base of the 
cheetze). 

” 4.— ” ” ”) ” MAJOY. 

Be SS BS aA perturbans. 

Oat Ae AG 5 argentipes. 


02S OM a a s himalayensis. 


-Rec. Ind. Mus.,Vol. IV, 1910. Plate VI. 


a A 


— 


Q 


3ax 380. 


Da. 


5 pais. 


7x 240 


NUL ONO MLE VATU RS ge N CULILCID 2. 
E. BRUNETTI. 


Since the connection between mosquitoes and malaria was 
definitely established a few years ago, numerous writers (medical, 
anatomical, -biological, economical and so on,—practically every- 
thing but dipterological) have appeared, probably attracted by the 
economic aspect of the question. It is to be.regretted that hardly 
any of these have possessed any general entomological knowledge, 
and the natural consequence has arisen; the differences between 
the various forms have been so grossly exaggerated taxonomically 
that a few years more progress at the same rate might see the Culi- 
cidee elevated to the rank of a Sub-kingdom, with classes, sub- 
classes, orders and so on. The absurdity of even the present 
state of things is so self-evident to everyone with a general know- 
ledge of Diptera combined with some acquaintance with taxonomic 
values in other orders also, that a serious protest seems imperative. 

The critical observations contained herein are not intended 
as any attack on what our American contemporaries amiably 
designate ‘‘ mosquito sharps,’’ but as a real and earnest protest 
against the utter disregard to zoological value and stability exhi- 
bited in the creation, on characters often of quite minor importance, 
of hosts of new ‘‘ genera’’ and even ‘‘ sub-families,’’ to say 
nothing of species, the latter being a point on which opinion may 
for the present be reserved. 

Prof. Williston ‘is, so far as I am aware, the only systematic 
dipterologist to endeavour to stem the tide of irresponsible additions 
to the literature of this order,! but apparently with little success, 
and it seems-incumbent on me to support dipterological opinion, 
having associated myself, at least clerically, with the Culicide 
of the Orient. 

His most instructive paper should be carefully read by all, 
because the whole subject is presented with the greatest moderation 
yet with absolute conciseness, in spite of the present condition 
of things in this family being calculated to cause considerable irmi- 
tation to the systematist. 

To quote a few of his more pertinent observations, it is im- 
portant that of the present-day writers on mosquitoes nearly all are 
non-dipterologists, ‘‘ some indeed, whose only papers on ento- 
mology have been those proposing new subfamilies ! 

‘“ Their ignorance of related Diptera has more than once been 
deplorably shown by writers on the Culicide ’’ and he observes 
‘“no one is competent to discuss philosophically the classification 
of any group of animal life, who is not well grounded in the principles 
of taxonomy as applied to related animals,’’ for ‘‘ the mosquitoes 


Can. Ent., xxxviii, 384. 


” 


! «¢ The Classification of the Culicide, 


56 Records of the Indian Museum. [VoL. IV, 


Corethring ; but in my.compilatory work, the Anopheline and 
“Edeomyine were accorded sub-family rank, merely in deference 
to specialists (as was distinctly stated in my reply to Messrs. Dyer 
and Knab’s criticism of my Oriental Catalogue, Canviint «xis, 127). 

These groups were presumably differentiated as follows : : 
Anopheline, palpi longin @ and 2; Culicine, palpi long in @ short 
in 9; Ldeomyine, palpi short in @ and 2; Corethrine, proboscis 
short and not formed for biting, which separates this sub-family 
from the other three. 

No doubt the genera of intermediate nature efface the dividing 
lines between these sub-families, which only supports the plea that 
no true separation exists except between the Culicine and Coreth- 
yin@é. Miss Mitchell echoes writers on mosquitoes only, in asking 

‘why should the Corethrid: #@ be placed in Culicide,” and in referring 
to both groups as ‘‘ heterogeneous elements.’ 

Replying to this it may be remarked that Corethra, to the 
systematic dipterologist, has always been in Culicide, where, I main- 
tain, it must most certainly still remain, as in its natural affinity. 
To compare it with Szmulium is quite wrong, the two genera having 
nothing incommon. Most of the workers in mosquitoes forget that 
venation is one of the soundest characters in classification, and that 
Corethra has a venation identical with Culex. 

The absence in the former genus of the stiff proboscis formed 
for piercing, characteristic of the Culicin@, does not necessitate 
the elimination of the genus from Culicide. 

Stomoxys and its allies have a stiff piercing proboscis, yet they 
have been admitted till recently merely as a group of genera in the 
sub-family Muscine.. In Girschner’s rearrangement of groups in 
Muscide s. lato the Muscine sub-family is suppressed but Stomoxvs, 
etc., are none the less afforded generic rank only, and incidentally 
it may be noted that. with this new sequence of genera in Muscidee I 
cannot possibly agree. Drymeia has also a_ stiff proboscis, yet 
is merely an exceptionally structured genus of Anthomyide. 

Other genera in other families could also be cited. 

The methods of depositing the eggs, the anatomy of the 
stomach, the minor characters: of the larve, and in fact, all the 
features emphasized by Miss Mitchell, would be regarded by sys- 
tematic dipterologists as subservient to, for instance, venation, and 
any bodily structure in the adult of a much higher nature than the 
variation of organs known to be subject to the greatest differen- 
tiations. 

To borrow again from Williston’s article, ‘‘ The three or four 
new families that have been proposed in recent years, all of them 
with more distinctive characters than the Corethrine possess, have 
been unanimously rejected by dipterologists.’’ 

Besides, many of the points urged by Miss Mitchell are flatly 
denied shortly afterwards by Mr. F. Knab, in the same Journal 
(Can. Ent., xxxix, 340). 

Mr. Knab’s reply to Miss Mitchell’s article calls for little 
comment here as it mainly consists of refutations or doubts of 


1910. | E. BRUNETTI: Taxonomic values in Culicide. 57 


the statements made by the latter author concerning the life history 
of various Culicide as compared with species of Chironomide, 
Psychodide and Dixa. As Iam personally unacquainted with the 
earlier stages of any of these genera, it would be presumption to 
decide ‘‘ when doctors disagree,’’ but I have seen no reply to Mr. 
Knab on the subject, and his facts appear well founded. 

Regarding Dixa, it has always been regarded as somewhat 
abnormal; to me personally it appears intermediate between 
Tipulide and Culicidee. 

The excuse for the erection of such a number of new genera, 
and the splitting up of the Culicide into several sub-families, is 
usually the unwieldiness of the genera, which otherwise would 
contain such a large number of species in each. 

As it is well known to be a common thing for students to deter- 
mine the species first, and discover its genus afterwards, the in- 
stability of most of the genera is surely emphasized, as the present 
writer never heard of this method of determination being followed 
in any other group. 

Moreover it is entirely wrong, scientifically, to consider that 
the presence of a large number of species in any genus is sufficient 
justification for dividing it into several others and according them 
equal value. A genus, as Prof. Williston truthfully says, should 
be something more than a second name for a species, or a cogno- 
men established for convenience’ sake only. ‘‘ A genus is a concept ”’ 
is written in the late Baron Osten Sacken’s handwriting, inside 
the covers of his hand-copy of Aldrich’s Catalogue of North American 
Diptera (now in the possession of the Indian Museum), and it should 
have a real zoological value and significance ; all genera throughout 
the animal kingdom being theoretically more or less on the same 
plane of systematic value. Personally, I object to all classification 
which is not as nearly as possible a natural one, and purely artificial 
groups should at the very outside attain only the rank of sub- 
genera,! 

The plea of the unwieldiness of extensive genera cannot be 
upheld, as the systematist is quite accustomed to such genera. 
In the first five volumes of the Catalogue of Diptera now in process 
of publication by Prof. Kertesz, are to be found numerous such 
genera, with approximately the following number of species each: 
Mycetophila 190, Sciara 460, Chironomus 320, Culex 182 (up to 1920), 
Cecidomyia 180, Dasyneura 160, Tipula 310, Odontomyia 160, 
Chrysops 150, Pangonia 180, Anthrax 460, Exoprosopa 230, Bomby- 
lius 240, Asilus 260, Promachus 150, Laphria 230, in addition to 
nearly twenty others with a little over or a little under 100 species 
in each ; the whole triumphantly capped by the gigantic genus 


1 An objection to ‘‘ preliminary descriptions,’’ often of a few lines or words 
only, may suitably be here recorded. It seems doubtful whether priority can 
justly be claimed in such cases, the full descriptions being often long delayed ; 
though I recognise the awkwardness of the situation in deciding whether a ‘ pre- 
liminary ’’ description is of sufficient length to stand good or not. 


58 Records of the Indian Museum. [Vou. IV, 19f0.] 


Tabanus with 912, a genus in which over and over again abortive 
attempts have been made to dismember it successfully. 

The real cause of the undue inflation into genera of what 
should be merely groups of species, and the elevation of a few of 
such so-called genera into sub-family rank isthe general absence of 
knowledge of the other families in Diptera in the present-day writers 
on Culicide. Williston has already been quoted on this subject. 

Therefore, in his remark that ‘‘ triviality has reached its limits 
(referring to the slender characters on which so many genera have in 
recent years been established), I cannot but entirely concur: this 
without any individual reproach to workers in mosquitoes, many 
of whom have been most courteous to me personally. 

It is almost certain that a wider knowledge of the accepted 
zoological value of such terms as ‘“‘ family,’’ ‘‘ sub-family,’’ 
‘“* genus,’’ etc., would convince them of the grossly exaggerated 
value attributed by them to what the systematic dipterologist 
would term quite secondary characters. 

In short, if any culicidologist would have the patience, before 
making new genera and sub-families, to read up some of the syste- 
matic dipterological literature of the last 60 or 80 years, more 
especially the tables of genera in the various families of Diptera, 
contained in Schiner’s Fauna Austriaca (than which no better 
standard work on the order has ever been issued)! he would find the 
greatest varieties of forms, not only in antenne, palpi, genital organs, 
exterior covering (whether scales, hairs, bristles, spines or other- 
wise), proportionate parts of the body and so on, but in venation 
also; all this in the same family, yet in spite of the hundreds of 
new species erected yearly, all attempts to create new families and 
sub-families on slender characters meet with strenuous opposition. 

It may be remarked here, although the subject will be treated 
more fully in the forthcoming Supplement to my Catalogue, that 
recent writers on this family appear to depart deliberately from 
biological precedence in the methods of presenting the results of 
their studies to others, with the result that the consultation of their 
writings is unnecessarily rendered materially more difficult. This 
is chiefly in their method of quoting from other authors; in the 
indices, and the undue prominence given to the @ . 

In the present paper however the only object has been to call 
attention to the instability of the great bulk of the generic and 
higher divisions recently proposed in this family. 


>” 


1 The examination of these tables alone forms, perhaps, the most compres 
hensive yet concise method of obtaining a rapid insight into the principles of 
classification in this order. They may be with advantage supplemented by the 
equivalent tables relating to North American genera, contained in Prof. Williston’s 
admirable manual ‘* North American Diptera,’’ 2nd Ed. 


Me wer VES ONe oO bt HE OR PRN? AL BIT, OOD - 
Seer G MUS CID A -(SFONtOX FN Z 
PHiLE aw MAEOM Y TA. Suse. AND 
PRES TREY NCH OMA [A . 

GEN. NOV.). 


By E. BRUNETTI. 


In the present paper, sixteen species are regarded as probably 
distinct, belonging to the group Stomoxine. I have also estab- 
lished a new genus Pristivhynchomyia, with one species, allied to 
Mr. Austen’s recently created genus Philematomyza. 

These latter two genera certainly belong to the group of genera 
immediately approximate to Musca, whatever this group may 
be called. The arrangement of genera in Muscide in the 3rd 
volume of the Palzearctic Catalogue contains several associations 
with which I can hardly concur ; especially the placing of the genus 
Musca itself in Anthomyide, and the abolition of Oestridee as a 
family. 

Two genera not yet found in the Orient are included amongst 
these descriptions, that they may be recognised should they occur 
here. 

One (He@matobosca, Bezzi) is European, and may occur in the 
Himalayas or in North-West India ; the other (Slygeromyia, Aust.) 
is African, and might easily be imported through the agency of 
cattle traffic. 

That the species are generally closely related to each other is 
easily realised by the number of synonyms even amongst the few 
European species. Stomoxys calcitrans, 1,., has been described seven- 
teen or eighteen times, Hematobia stiinulans, Mg., possessing six 
synonyms and Lyperosia irritans, I,., a comparatively uncommon 
species, two. 

I have little doubt of the specific distinctness of all the forms 
described in this paper, with the exception of the genus S/omoxys, 
of which a considerable amount of further study seems imperative. 

At times it has even occurred to me that all the so-called species 
from the East could almost be comprised under three typical 
forms, which may be termed calcitrans, L.., indica, Picard, and a 
species with a distinctively formed fore tarsus in the @ (the only 
sex known)—/udlla, Aust. 

For this reason I cannot feel satisfied with the table of species 
in Stomoxys, and in this genus the present article must be regarded 
as more preparatory than conclusive. 


60 Records of the Indian Museum. [VoE. IV2 


It may be noted here that some confusion may possibly arise, 
and erroneous deductions arrived at by a misunderstanding, as to 
whether the width of the frons is measured across the vertex or 
across the middle. 

Mr. Austen, in his paper dealing with these genera measures 
along the vertex, but I have always adopted the second course, 
taking its average width, ignoring the widening at the vertex and 
towards the frontal triangle ; or in other words I have regarded its 
width as the distance between two perpendicular lines drawn so as 
to touch the greatest length of eye margins. 

My thanks are due to Prof. Bezzi for some valuable notes on 
Lyperosia and Hematobosca and especially to Mr. E. E. Austen for 
information, respecting Lyperosia minuta and L. exigua, enabling 
me to avoid describing both species again as new. ‘The former 
I knew only from four specimens (African) in indifferent condition ; 
the latter from the description only, in which, be it noted, no men- 
tion is made of the conspicuous long hairs on the hind tarsi in theo’. 
As my descriptions of both species have been drawn up from a 
good series of both sexes of each species, I allow them to remain 
as redescriptions of minuta and exigua; the original descriptions 
of these two species not always being accessible. 

In connection with studies on this group, the following papers 
may be consulted :— 


rt. E. E. Austen. A monograph of the Tsetse flies (Glossina). 
London, British Museum (1903). 

2. K. Griinberg. ‘‘ Uber blutsaugende Musciden,”’ Zoologis- 
cher Anzeiger, xxx, 78 (1906). 

3. M. Bezzi. ‘‘ Die Gattungen der blutsaugenden Musciden,’’ 
Zeits. Hymen. v. Dipt., 1907, p. 413. 

4. M. Bezzi. ‘‘Mosche ematofaghe,’’ Rend. Istit. lomb. di 
Scie lett... 1Q07-) Pp: 1453- 

5. E. E. Austen. ‘‘ New genera and species of blood-sucking 
Muscide from the Ethiopian and Oriental Regions, 
in the British Museum,’’ Ann. Mag. Nat. Hist. (8), ii, 
285 (1909). 

6. KE. E. Austen. Illustrations of African blood-sucking 
flies, other than mosquitoes and tsetse flies. London, 
British Museum (1909). 


Table of genera in STOMOXINE. 


A Arista plumose above and below. 
I. Ist and 3rd longitudinal veins at base bare. 

Apical part of 4th vein quite straight. 

Ist posterior cell rather narrowly open. 

Body generally more slender (Lyperosza-like). 

Palpi comparatively less spatulute. 

Arista with comparatively fewer hairs below H@matobosca,' 
Bezzi. 


| Not yet recorded from the Orient, but occurs in Italy. 


IgI0. | 


AA 


Ce 


2. 


Oo 


E. Brunetti: Blood-sucking Muscide. 61 


Ist and 3rd longitudinal veins at base bare. 

Apical part of 4th vein not straight, but less curved than in 
Hematobia. 

Ist posterior cell broadly open. 

Body generally stouter (Stomoxys-like). 

Palpi comparatively more spatulate at tips. 

Arista with comparatively more hairs below, Ldellolarynx, 

Aust. 

Ist and 3rd longitudinal veins with a few minute bristles 
at their bases. 

Apical part of 4th vein much curved (as in Stomoxys). 

Ist posterior cell broadly open. 

Body generally stouter (Stomoxys-like). 

Palpi comparatively less spatulate at tips. 

Arista with comparatively less hairs below, Hematobia, 


Rob. Desv. 
Arista plumose above only. 
Palpi much shorter than proboscis. 
(3rd longitudinal vein bristly at base) Stomoxys, Geoff. 


Palpi always approximately as long as proboscis. 

Apical part of 4th vein quite straight. 

3rd longitudinal vein with minute bristles at base. 

Ist posterior cell rather narrowly open. 

Proboscis not tapering ; palpi clavate at tips. 

Body chiefly Stomoxys-like. 

Sternopleural bristles black, Stygeromyia,! Aust. 


Apical part of 4th vein with distinct curve, as in Stomoxys 
and Hematobia. 

3rd longitudinal vein at base bare. 

Ist posterior cell rather broadly open. 

Proboscis gently tapering from base to tip; palpi not 
clavate at tips. 

Sternopleural bristles distinctly yellow, Lyperosia, Rond. 


HAMATOBOSCA, Bezzi, 1907. 
Zeits. Hym. Dipt., 414. 


This genus may be characterised thus: Body more slender 
than in Hematobia; palpi as long as proboscis ; arista with some 
hairs below; 3rd longitudinal vein bare at base; 4th, as a whole, 
considerably curved (in comparison with that of Lyferosia) ; Ist 
posterior cell rather narrowly open. 

I add a description of this genus to my paper, although it is 
not yet known from the East, because it may easily occur in the 
northern part of India. 


1 Not yet recorded from the Orient, but occurs in Arabia. 


62 Records of the Indian Museum. [Vor.EVs 


The typical (and only) species is atripalpis, Bezzi, Bull. So. 
Ent. Ital., xxvii, 60 (1895) (H@matobia), described from Italy. 

Austen’s recently erected genus bdellolarynx is very closely 
allied. 


BDELLOLARYNX, Austen, 1900. 
Ann. Mag. Nat. Hist. (8), iii, 290. 


‘““Small, stoutly built, thick-set flies, closely allied to 
Hematobia, R.D., but distinguishable as follows :—No sexual colour- 
dimorphism (unless it be in wings); front, facial angles, and 
anterior margin of buccal cavity less prominent ; jowls descending 
considerably behind, instead of nearly horizontal, but basi- 
occipital region much less protuberant ; bristles below facial angles 
small and fine, instead of relatively coarse; in @ front wider 
with upper inner margins of eyes more nearly parallel, less ap- 
proximate in middle of front ; arista feathered above with consider- 
ably longer hairs, and below with about six fairly long hairs ; Ist 
and 3rd longitudinal veins entirely bare, without bristles at base.’’ 
The cheetotaxy is as follows: Humeral bristles 2; post-humeral 
I; notopleural 2; presutural 1; supra-alar I; intra-alar 1; post- 
alar 2; dorso-central 5 (2 in front of and 3 behind suture, may be 
difficult to distinguish, especially in @ ) ; inner dorso-central I (some- 
times 2 in @); scutellar 4 (1 preebasal, 1 basal, 1 discal, situated 
close to lateral margin, I apical); mesopleural about Io or 12; 
sternopleural I : I. 

‘* Bdellolarynx is distinguished from Hematobosca, Bezzi, 
by the palpi being much more spatulate at the tips, by the arista 
having a greater number of hairs below, and, in the wing, by the 
shape of the Ist posterior cell and the course of the terminal portion 
of the 4th longitudinal vein (in Hematobosca the latter details are 
the same as in Stygeromyia, Aust.). Typical species, B. sanguino- 
lentus, sp. nov.’’ 


Bdellolarynx sanguinolentus, Aust., 1909. 
ocx cits, 200. 


o 2. India, Ceylon, Assam. Long.o 43-5 mm. ; 2? 33-44 mm. 


‘“ Mouse-grey or slate-grey, clothed with short black hair, 
bristles also black ; palpi buff; dorsum of thorax with a pair of 
extremely narrow, widely separated, parallel, longitudinal ad- 
median blackish stripes, a somewhat triangular blackish mark 
extending backwards from inner end of each humeral callus, and 
an elongate blackish streak in a line with latter mark behind 
transverse suture (two outer marks less distinct in ¢ ); dorsum of 
abdomen with a narrow, interrupted, longitudinal, median, clove- 
brown stripe, and, on 2nd and 3rd segments, paired transverse, 
roughly triangular, clove-brown blotches, the pair on 2nd segment 
especially large. Wings hyaline or tinged with tawny olive ing, 
hyaline in 2 ; femora greyish clove-brown, extreme tips ochraceous 


IgI0. | E. Brunetti: Blood-sucking Muscide. 63 


buff, one or more pairs sometimes more or less brownish except at 
base, tarsi dark brown.’’ 

Types in the British Museum; @ captured by me at Calcutta, 
8-11-09; @ from Mussoorie, ix-o6 [Howlett]. Other specimens in 
the British Museum from Allahabad, 6-x-05 [Howlett]; Sylhet, 
13-iv-05 [Hall]; and Henaratgova and Haldumulla (both Ceylon), 
7-iiand I4-vi-92 respectively [| Yerbury]. Inthe Indian Museum are 
_3 @¢@ from Tharbani, Nepal, 27-ii-o8; and from the base of 

the Dawna Hills, Lower Burma, 2-iii-o8 [Annandale]; 37 ~ from 
Bandra (Bombay Presidency) ‘‘ hovering in slaughter house,”’ 
28-xi-09 [Hodgart]; and one 2 from Maddathorai (base of Western 
Ghats, Travancore, 17-xi-o8 [Annandale]. I have a #7 in my own 
collection taken by me in Calcutta, 18-ii-o5. Both Dr. Annandale 
and the Museum Collector Mr. Hodgart have noticed the apparent 
habit in the males of this species of hovering, an unusual practice 
of the members of this subfamily, although I have occasionally 
noticed it in Stomoxys calcitrans. 


N.B.—Although the specimen forming the type of this genus 
was taken by me I was unaware of its generic difference from 
Stomoxys at the time of capture. 


HAMATOBIA, Rob. Desv., 1830. 
Essai sur les Myod., 388. 


Original description.—** Palpi elongati, ultra epistoma porrecti, 
interdum apice dilatati ; apud quasdam species, tarsi postici externé 
serrati.’’ 

This genus was first split off from Stomoxys by Robineau 
Desvoidy for those species in which the palpi are as long as the pro- 
boscis, in contradistinction to calcitvans with its very short slender 
palpi. The author added that the palpi are sometimes a little 
dilated at the tip, and that in some species the hind tarsi (‘* tarses 
posterieurs ’’) were serrated. He only included four species, all 
from France. Of these, two (ferox and geniculata) are synonymous 
with stimulans, Mg., and another (serrata) is synonymous with 
ivvitans, 1,., a species for which Rondani erected the genus Lyferosia, 
The remaining species, ¢zbialis, still stands good as distinct and 
remains in Hematobia. 

From the more restricted point of view, the characters of 
Hematobia are: (1) the presence of minute but distinct (though 
easily broken off) bristles at the base of both the Ist and 3rd longitu- 
dinal veins, this being the only genus in which these bristles 
appear on the Ist longitudinal vein ; (2) the arista being plumose 
below as well as above ; (3) the palpi, which are practically as long 
as the proboscis; (4) the well-curved apical part of the 4th longitu- 
dinal vein (as in Stomoxys, to which genus the present species bear 
considerable resemblance in size and shape) ; (5) the broadly open 
Ist posterior cell. Lesser characters may be gleaned from my table 
of genera. 


64 Records of the Indian Museum. [VoL2 EV, 


Table of Oriental species. 


Legs all black, except base of tibize 


pale a .. sanguisugens, Aust.,o. 
Posterior femora wholly pale =<. i74ipcs-sp. now. 


Hematobia sanguisugens, Aust., 1909. 
Ann. Mag. } Nat. Hist. (8), iti, 288. 


gy. Himalayas. Long. 5h 4-6 mm. 


‘* Olivaceous-grey, or brownish grey, with clove-brown mark- 
ings ; dorsum of thorax with 2 pairs of clove-brown longitudinal 
stripes (outer stripes broader and widely interrupted at transverse 
suture), anda less sharply defined median stripe ; dorsum of abdo- 
men with a clove-brown median longitudinal stripe, extending 
from front margin of 2nd to beyond middle of 4th segment, narrowly 
interrupted before hind margins of 2nd and 3rd segments, a pair of 
transversely elongate dusky blotches on Ist segment, a pair of large 
clove-brown spots on 2nd segment and a pair of similar but smaller 
spots on 3rd segment ; wings slightly infuscated, light sepia coloured ; 
legs ploy brow n, bases of tibize ochraceous- buff. 2 

The types of this species are in the British Museum taken 
at Kasauli, Punjab (W. Himalayas), ‘‘on cows’’ [Lt.-Col. F. 
Wyville Thomson}. ‘* The flies sucked the animals, and their abdo- 
mens became distended with blood. I have never noticed them 
biting man.’’ Mr. Austen compares his species to the European 
H. stimulans of Meigen, distinguishing it by ‘‘ its usually somewhat 
smaller size (average length of 6 #7 @, 5°75 mm.), by the median 
dark dorsal stripe on the abdomen being practically continuous 
throughout its extent instead of widely interrupted before reaching 
the hind margins of the 2nd and 3rd segments, by the 4th abdominal 
segment in the ~ being always without a pair of dorsal spots, 
and by the rst longitudinal vein being either entirely bare or having 
at most one or two minute bristles, instead of a row of bristles con- 


spicuous under a strong lens when viewed at a low angle from the 
direction of the hind margin of the wing.’’ 


N.B.—One o& in the Indian Museum, taken 10-vili-og by 
Mr. Paiva at Darjiling (7,000 ft.), agrees almost perfectly with 
Mr. Austen’s description. Incidentally, I may note that, previous 
to reading of his species, I had marked this specimen as a variety 
of stimulans. Mr. Austen however mentions six specimens that 
show consistency in the markings. The lesser size I do not value 
as a specific character as I have noticed that Oriental specimens of 
very common European species appear to be generally undersized. 
The intrinsic value of the minute spines on the bases of the Ist 
and 3rd longitudinal veins, when present, is also, to my thinking, 
much less than some writers have considered it. 


Ig10.] E. Brunetti: Blood-sucking Muscide. 65 


H. stimulans may quite possibly occur in Northern India ; 
this species, if found, should be recognisable by the differences from 
sanguisugens given above. 


Hematobia rufipes, mihi, sp. nov. 


9. Himalayas, Nepal. Tong. 4-5 mm. 


Head. Frons one-third width of head, yellowish grey, with a 
broad dull bare stripe. Ocellar triangle with two pairs of small 
bristles above and a pair of strong ones below. 

Fronto-orbital bristles 9, the upper 4 placed rather out of the 
straight line, slightly irregularly, the lower 5 normal and well curved 
inwards. 

Face whitish grey, with the usual bristles about the mouth. 
Antenne blackish, arista with 7 long hairs above, and 5 below. 
Palpi reddish yellow, with black bristles, which are more numerous 
at the tip. Proboscis wholly shining black, slightly longer than 
palpi. Back of head grey, occipital orbit normally fringed with 
bristles. 

Thorax. Dorsum yellowish brown, with a broad median grey 
stripe from anterior margin to beyond the middle ; this stripe edged 
with a dark brown but not sharply-defined line. "Towards the margin 
of the dorsum the colour is much browner. Humeri grey. The 
median grey stripe appears blackish, if viewed from behind. The 
whole of the sides of the thorax uniformly blackish grey, with minute 
sparse black pubescence. Scutellum brownish yellow, with minute 
pubescence, two strong spiny bristles towards each side. The two 
apical bristles are broken off, but appear to be strong. Chetotaxy. 
Humeral bristles 3 (one or more sometimes weaker) ; post- 
humeral 1; presutural 1; noto-pleural 2; supra-alar 2; intra- 
alar 1; post-alar 1; meso-pleural 10, the uppermost placed just 
below the hindermost noto-pleural ; sterno-pleural 1: 1. The dorso- 
central bristles not very conspicuous, apparently I before, 3 or 4 
behind suture and a hindermost very strong one. 

Abdomen. Yellowish grey ; a narrow median black stripe ; 
each segment mainly occupied by a pair of large blackish spots 
of irregular shape. Whole dorsum covered with soft hairs. Belly 
grey. 

Legs. Reddish yellow. Fore femora (except extreme tips), 
tips of hind femora, and the tarsi, blackish. Fore femora with 
rows of long stiff hairs on outer and lower sides ; middle femora 
with 2 curved hairs above, at tip ; hind femora with a few longish 
hairs above at tip, one below. Middle tibiae with 2 or 3 bristles 
at tip, hind tibize with one bristle behind, at tip. Legs minutely 
pubescent. 

Wings clear, outer cross-vein nearly straight. Tegule yellow- 
ish ; halteres yellow. 

Described from 2 2 2 in the Indian Museum. Darjiling 


(6,000 ft.), 29-ix-08 [Brunetti , tvpe| ; and Noalpur, Nepal, 23-11-08. 


66 Records of the Indian Museum. { Vor. ave 


STOMOXYS, Geoff., 1764. 


Hust: Desens. s11yc538: 
See also Sch. F. Aust. (Dipt.), i, 577 (1862). 


The genus Stomoxys, Geoff., as understood by the older authors, 
and as described by Schiner, included both stimulans, Mg. (now 
referred to Hematobia), and trritans, I,. (now placed in Lyferosia). 
Both these latter species were primarily split off from Stomoxvs, 
under the name H@matobia, Rob. Desv., on the strength of the palpi 
being as long as the proboscis. 

Some discussion has arisen as to whether the name Stomoxys, 
Geoff., or Szphona, Mg., should stand for the present genus. ‘Two 
recent authorities,—Speiser (Zeits. f. wiss. Insek., i, 461) and 
Bezzi, both in his ‘* Mosche ematofaghe ’’ (Rend. Istit. lomb. (2), 
xl, 17) and in the 3rd volume of the new Palearctic Catalogue ,— 
have replaced Stomoxys by Siphona, Mg. 

I am glad to see that Mr. Austen (Ann. Mag. Nat. Hist. (7), 
x1x, 445) has deprecated the change, giving full and satisfactory 
reasons for supporting the continuance of the name Stomoxys for the 
present genus. 

The following generic description is founded on that of Schiner, 
with the elimination of such characters as only apply to the more 
recently established genera, and the addition of other distinctive 


characters which appear to be, in the restricted sense of the genus, 
generically consistent. 


STOMOXYS, Geoff. 
(Redescription.) 


Medium-sized, grey flies, one species (calcitrans) being one 
of the common ‘‘ house flies.’’ 

Head semi-circular, somewhat flattened in front; epistome 
nearly straight, nearly bare ; buccal cavity hardly prominent, with 
a row of strong spines each side ; cheeks rather narrow, descending 
only a little below the eyes. 

Front in o narrow, in @ about one-third the width of the 
head, the exact dimensions in both sexes varying in different species, 
and (within narrow limits) sometimes in the same species. In both 
sexes a row of about 7 tronto-orbital bristles, which in the @ are 
augmented by smaller ones on the upper part of the frons placed 
nearer to the eyes. Eyes bare. 

Antennee recumbent, moderate in size, Ist joint very small, 
2nd distinct, with a few short bristles, 3rd about three times the 
length of the 2nd. Arista plumose on upper side only, with about 
8 to 10 long, well separated hairs of (after the first one or two) 
gradually decreasing length nearly to the tip, these hairs being 
very slightly curved, but seldom bisinuate. Proboscis horizontal, 
very prominent, of hard chitinous material, considerably thicker 
on basal part, narrowing gradually to tip ; labella very small. Palpi 


IQIO. | E. BRUNETTI: Blood-sucking Muscide. 67 


cylindrical, narrow, small, much shorter than proboscis, often 
inconspicuous through lying flat against the epistome. 

Thorax a little longer than broad, of the average Muscid type, 
with macrochetee,! and minute bristly hairs. 

Abdomen ovato-conical, moderately arched, generally marked 
with infuscated spots, a dorsal stripe, or transverse darker bands. 
Legs of moderate size and length, minutely pubescent, with a 
few stronger bristles ; inner side of middle tibiae bare. Wings of 
the typical Muscid venation, resembling that of Cyrtoneura stabu- 
lans, F. (also a ‘‘ common house fly ’’’), except that the 4th longi- 
tudinal vein is bent upwards in a sharper curve after passing the 
inner transverse vein, thus leaving the Ist posterior cell rather 
broadly open ; 3rd longitudinal vein with some distinct but minute 
bristles at its base. 

The metamorphoses occur in decaying vegetable matter, horse- 
dung and similar substances. Both sexes in the adult suck blood, 
including, occasionally, that of man, but they are especially irri- 
tating to horses and cattle. The genus is probably world-wide, or 
at least wherever civilised man is found. 


Table of Oriental species of STOMOXYS. 


A Fore tarsus normal. 

B Abdominal segments without 
bands, but with two distinct 
(often ill-defined) spots on 
at-“least the -2nd :and> 3rd 
segments. Tibize pale at 
base only. 

Frons in @ 4 width of head ; 
in 9 4. 

D Thorax with 4 well-defined 
(but variable) stripes, never 
divided. Width of frons 
jing din 9.. : 

DD Thorax with 4 oblong large 
spots, each more or _ less 
divided. Width of  frons 
(presumably) neither + nor 
+ as sex iS given as 
doubtful oF ..  plurinotata, Big. 

CC Frons in @ + (9 unknown). 

Tibize with basal half pale, 
middle pair apparently 
wholly so abe .. oblongopunctata, sp. nov., @. 


J} The chetotaxy of S. calcitvans is fully described under the heading of 
that species rather than under that of Stomovys, as I do not wish to suggest 
that that particular arrangement of bristles is necessarily generic, not being 
acquainted with the chetotaxy of species from other parts of the world, that 
is to say of species which must obviously remain in this genus. The chetotavy of 
at any rate all the species I have seen is uniform. 


, 


calcitrans,L., 7 @. 


68 Records of the Indian Museum. [VoL.. EV; 


CCC Frons in 7 ito ¢ (2? 2 un- 
known). 
EK Frons shining white ; abdom- 
inal marks as in calcitrans 
(2? Oriental) sitvens, Rond., o. 
EE Frons quite dark; 2nd, 3rd, 
and 4th abdominal segments 
each with two black marks 
in the shape of elongated 
triangles a .. tmangularis, sp. nov., o&. 
BB Abdominal segments with a 
distinct black band on, 
at least, hind margins. 
F Frons comparatively wide, 14 
to rather wider than +. 
G Frons 4; tibiz pale at base 
only ; transverse dark band 
at base of 2nd and 3rd 
segments as well as on hind 
margins .. bengalensis, Picard,@ 2. 
GG Frons_ rather wider than +; 
anterior tibie and tarsi pale ; 
no band at bases of segments ; 
2nd segment wholly clove- 
brown or nearly so (2 un- 


known) Aa .. pusilla, Aust., 7 
FF Frons narrow, +} to ¢; tibie 
wholly pale ae n 2 \undica, Picard, o'9.. 


AA Fore tarsus in o with Ist joint 
fringed on inside with a 
row of hairs of equal length 
(2 unknown) .. pula. AuStic 


N.b.—The above table of species is by no means satisfactory 
to me as sufficiently distinguishing them, but in the present un- 
certain knowledge as to the limits of each, it is the best that 
I can offer. 


Stomoxys calcitrans, I,., 1761. 
Fauna Suec., Ed. ti, 467 (Conops). 


N.6.—For the numerous descriptions of this species under 
the specific name calcitrans, see the recent Catalogue of Palearctic 
Diptera, vol. iii, p. 611. 

A list of the various synonyms of calcitrans is appended. 

pungens, DeGeer, 1776, Ins., vi, 78 (Musca 1d.). 

tessellata, Bo T7045 Bat Sys iV, 395- 

1d., 1805. ae Ant 32 81 


1 As noted under the debe of this species, I cannot specifically distin- 
guish /imbata, Austen, from indica, Picard. 


IQI0. | E. Brunetti: Blood-sucking Muscide. 69 


aculeata, Rob. Desv., 1830, Myod., 386. 


d 


diva, id., ta. 387. 

infesta, id., ibe 387. 

minuca, td., tae 387. 

pungens, va., 1d. 386. 

sugillatrix , Tas. id., 386. 

aurifactes , 1d. , 1863, Hist. Nat.d? Dipt.4u, 607; 9. 
chrysocephala, 1d., tae, 604. 
claripenms , td., id., 604, &. 
cunctans , id., td., OG75 oP 
flavescens , td., TC a 605, o. 
minuta , ad., Tas, 606, 2. 
precox, id., td., 608, @. 
vubrifrons , vd. ad., 606, o. 
vulnerans , 1d. d., 605. 725 
aculeata , Mcq., 1835, Suites a Buff., 11, 242. 

pungens, dd., Las, 242. 

pungens, Mg., 1838, Sys. Besch., iv, 170. 

> libatrix, Rob. Desv., 1830, Myod., 387. 


(Redescription.) 


Head. o. Frons one-fourth width of head; grey, yellowish 
grey or slightly bluish grey. A very wide frontal stripe appearing 
from above almost velvet-black, thus leaving only very narrow eye- 
margins. Generally, this black stripe has a greyish V-shaped piece cut 
out of its upper end, which lighter space encloses the ocellar triangle, 
which latter is small, and touches the absolute vertex ; and bears, 
just above the lower ocellus, two pairs of bristles (the lower pair 
the larger). There are also 4 other pairs close together (all 6 pairs 
forming one double row) on the upper part of the triangle, the 
uppermost pair being on the extreme vertex. Seen from below, the 
black frontal stripe is invisible, the whole frons and face appearing 
white, or yellowish white. Fronto-orbital bristles, about 15, the 
upper part of the row not always placed uniformly. ‘wo pairs of 
vertical bristles as usual. Eyes black, absolutely bare. Antenne 
blackish, 3rd joint more or less vellowish grey, 2nd joint with 
several small bristles and 1 or 2 long ones ; arista with 8-9 bristles 
and minute pubescence on basal half above ; thickened at base, where 
it is often yellowish. Face white or yellowish white, covered with 
close, microscopic tomentum. Mouth opening similar, with the 
usual strong bristles on each side. Palpi small, reddish yellow, with 
a few short black bristles. Proboscis shining black, quite bare, 
tapering from base to tip. Back of head generally more or less 
bluish grey, with very short pubescence. 

In the ? the fronto-orbital bristles are less strong and less 
regular, generally rather less in number, and on the upper part 
of the frons there is a short extra row each side, placed nearer the 
eye margin, sometimes also with additional irregularly placed small 
bristles. In all other respects the 2 head is similar to that of the ~. 


70 Records of the Indian Museum. [VoL. IV, 


Thorax. Ground colour varying from yellowish grey to whitish 
grey. Seen from in front, a dorsal pair of moderately narrow, 
well separated stripes, which are normally mummy-brown, but 
vary sometimes to the extent of being nearly blackish. These 
stripes extend from the anterior margin nearly to the posterior 
one. On the outer side of each stripe is a similar stripe, distinctly 
interrupted at the suture, in front of which it becomes sometimes of 
a thickened cigar shape, placed diagonally, pointing to the humerus, 
sometimes it retains its direct direction forwards as a continuation 
of the portion behind the suture. It often also takes the form of an 
irregularly shaped spot. At the posterior ends of these dorsal 
stripes are two irregular black patches, immediately in front of the 
scutellum. Towards the side of the dorsum, just above the wing, 
is a concolorous brown streak, sometimes distinct enough to form 
a short third stripe. 

Viewed from behind, the dorsal space between the two median 
lines appears much lighter, varying from light grey to even quite 
whitish. Scutellum concolorous with thoracic dorsum, generally 
with an irregular mummy-brown triangle at base. Chetotaxy. 
Humeral bristles 3, post-humeral 1 (weak), presutural 1, noto- 
pleural 2, dorso-central with only the hindermost in each row really 
strong ; a distinctly less strong one in front of it, whilst the remainder 
are barely larger than the small bristles dispersed generally over the 
whole dorsum, but there is usually a single moderately strong one 
just in front of the suture. Two moderate sized inner dorso-central 
bristles. Of the side bristles, there are 4 placed above the wing 
more or less in the shape of a diamond, all of which I should be dis- 
posed to term supra-alar, or possibly 3 supra-alar and I postalar. 

The mesopleura bears a hinder row of about 12, diminishing 
in size from above downwards except that the strongest ones are 
generally the 2nd and 3rd, or the 3rd and 4th. Some long stiff hairs 
on front part of mesopleura. Sternopleura with a strong bristle on 
upper hind corner, a strong one on lower hind corner, in company 
with some shorter comparatively strong ones, but which are much 
longer than the very small bristles covering the whole sternopleura 
and, to a greater or less extent, the whole of the side of the thorax. 

Scutellum with a strong basal bristle each side, near margin, 
with a less strong preebasal one in front, and the usual apical pair 
which are very strong. Occasionally a pair of bristles correspond- 
ing to dorsal macrochete stand out a little stronger than the 
microchetze covering dorsum of thorax and scutellum, but in most 
specimens they are by no means obvious. 

Abdomen. Normally (and generally) yellowish grey ; some- 
times much more yellowish, at least on the dorsum ; sometimes 
approaching slate-grey, either wholly or towards the sides of the 
segments ; seen from behind at a low angle, sometimes quite whitish. 
The markings, although following a general plan, are very variable. 
In what may be regarded as the typical form, there are on both the 
2nd and 3rd segments, a basal mark in the shape of a moderately 
wide short dorsal stripe, which reaches to about the middle of the 


1gIo. | E. Brunetti: Blood-sucking Muscide. 71 


segment, where it is sometimes cut off sharply and squarely, some- 
times drawn out to a distinct point, sometimes fading away 
indistinctly. There is a pair of more or less circular spots of 
moderate size placed on the posterior margins, each midway be- 
tween the middle line and the side border. On the 4th segment is 
a single, more or less triangular basal mark, sometimes small, or 
indistinct, or occasionally absent. All these marks are vandyke- 
brown in colour, occasionally approaching clove-brown. These prin- 
cipal markings can be seen most distinctly from above, or a little 
in front, often appearing blurred or almost invisible if viewed 
from behind. What I should call the secondary markings, are 
irregular, almost Sarcophaga-like reflections, often invisible when 
viewed from certain directions; of a pale greenish grey colour. 
They take the form of an irregular elongated spot each side, often 
connected by the colour extending along the posterior margin of the 
segment ; and a pair of comparatively large, more or less circular 
spots on the 4th segment, these latter spots being present in the 
majority of examples. 

The whole abdomen is shortly pubescent with short soft black 
hairs, which are little thicker on the shoulders, and a little longer 
and stronger on the posterior margins, and over the dorsum of the 
4th segment. 

When it comes to describe varieties, the difficulties as to the 
limits of the species are very apparent. Practically I regard as 
probable varieties of this species, all fotms which partake of the 
above pattern or any reasonable variation of it, this variation being 
considered to be of very wide limits. 

The short dorsal stripes on segments 2 and 3 occasionally form 
an unbroken stripe, extending to the base of the 4th segment. In 
one form they are very short, and the usual circular spots become 
quite elongated and narrow ; whilst in another variety there is a 
pair of additional similar brown nearly circular spots in front of the 
main pair, and placed on the base of segments 2 and 3. ‘here is 
often a pair of very narrow spots on the Ist segment, sometimes 
united into a transverse narrow line. In some examples the ordi- 
nary brown circular spots on segments 2 and 3 are united by the col- 
our extending narrowly along the posterior margin; sometimes 
bordering the basal margin of these two segments also. In another 
variety, well marked in its more pronounced form, the dorsal stripe 
is expanded on both base and posterior margin, on which latter 
it is often considerably widened and joined to the two spots, thus 
somewhat approaching bengalensis, Picard. 

It must be remembered that the extreme base of each segment 
is shining black, but that this is only visible when the segments 
are abnormally drawn out, and that in a normal state, the black 
colour is covered by the posterior margin of the preceding segment. 

Belly generally concolorous, with short, somewhat sparse black 
pubescence, without conspicuous markings. 

Legs. Blackish, minutely pubescent ; femora, and sometimes 
tibia also, more or less with greyish reflections. Fore femora 


72 Records of the Indian Museum. [VOL EYs 


with a row of bristles above from middle to tip, diminishing in size, 
a row on outer side on basal half, replaced towards the tip by 
moderately soft, rather long hairs; a row of about 16 to 18 com- 
paratively strong bristles on under side, extending the whole length 
of the femur, and equalin size. Middle femora with some bristles, 
lying rather flat, on front side; two contiguous, moderately strong, 
slightly curved bristles above, near tip, and a row of bristly hairs 
on under side. Hind pair with a row of bristles on front side, a 
single curved bristle below near tip. Knees tawny, the colour ex- 
tending sometimes over the whole basal half of some of the tibie ; 
tips of tibie with 2 or 3 short spines. ‘Tarsi blackish, under side 
with a little gold-yellow pubescence, which is sometimes lightly 
present on inner side of anterior tibiz. Pulvilli dirty white. 

Wings. Practically clear, but in the o» sometimes slightly 
tinged with brown on basal and front parts; Ist posterior cell 
widely open, the 4th vein forming a well rounded loop at the bend 
upwards ; apical portion gently curved. Outer cross-vein normally 
with one gentle inward curve, variable, at times nearly straight, 
or nearly bisinuate. Tegule vellowish white. 

Described from a long series of specimens of both sexes in the 
Indian Museum collection and my own, and also from European 
specimens. It is practically one of the world-wide species that 
occurs wherever civilised man is found, and in at least the East 
it is apparently present all the year round in every locality. Actual 
dates would therefore be of little significance. It is in the Indian 
Museum collection from Gangtok (Sikhim), Sylhet, Kurseong 
(Darjiling), Mussoorie, Simla, Calcutta, Port Canning, Puri, Berham- 
pur, Bhogaon, Travancore State (S. India), Rajmahal (all India) ; 
Colombo, Rangoon, Kawkareik (Lower Burma), Mandalay, and 
from on board ship, at light, off Tuticorin. Personally I have 
met with it in many parts of the Punjab, Mussoorie, at Rangoon 
Singapore, Penang, and other places further East, and have it from 
Java, Ceylon and the Philippines. 


N.B.—The above lengthy description has appeared advisable 
from the variability of this very common species. Further varia- 
tions and combinations of the differences (mostly in the abdominal 
markings) above stated, occur in individual examples, and as the 
species has already been described under no less than 17 different 
names, I hesitate to regard as distinct any form which appears to 
fall within its possible range of variation; especially as I have seen 
no - distiict'- “variety,” “races = Horm)’ 7 or *\ subspecies! 7 
peculiar to any special locality and illustrated by any number of 
consistently marked specimens. 


‘* Stomoxys libatrix,’’ Rob. Des., 1830. 
Ess. Myod., 387. 


The author’s very short description runs as follows :— 
' Very near inimica, R. D. (I.c., p. 387, from North America) : 
sides of the face a little more yellowish ; abdominal segments. 


y 


IQIo. | E. Brunetti: Blood-sucking Muscide. 73 


especially the 4th, bear a small transverse line at their bases ; the 
emarginations are blacker ; tegule brownish ; wings a little clearer.’’ 


N.B.—Prof. Bezzi (‘‘ Mosche ematofaghe, Rend. Istit. lomb., 
1907) says that, although admitted as distinct in Van der Wulp’s 
catalogue, it is probably only another synonym of calcitrans, L.., 
an opinion in which I cannot but concur, and, therefore, failing evi- 
dence to the contrary, I propose to sink the name as synonymous 
with calcitrans. 


Stomoxys plurinotata, Big., 1887. 
Bull. Zool. Soc. Fr., xii, 593. 


“Long. 54mm. Antennis nigris ; facie et fronte albescentibus, 
vitta frontali, bifida, nigra; palpis fulvis ; thorace albido cineras- 
cente, maculis quatuor latis, elongatis, fuscis, duplicatis, pleuris 
cinereis ; calyptris albis, halteribus pallide flavidis ; scutello ciner- 
ascente ; abdomine cinereo, segmentis I-3 superne univittatis, 
2° et 3°, utrinque, in medio, punctis duobus fuscis, rotundatis, 
notatis, cum vitta mediana aliquoties junctis; pedibus nigris, 
tibiis parum fulvo tinctis; alis hyalinis. Ceylan 3 specimens.’’ 

I append atranslation of Bigot’s more extended French descrip- 
tion :— 

Antenne black, tip of znd segment pale; vertex greyish, 
frons and face dirty white, frontal stripe wide, bifid above ; palpi 
pale, proboscis black. Thorax grey, with 4 large oblong elongated 
spots, nearly. contiguous, blackish, each divided by a not very 
distinct greyish line. Scutellum greyish; sides of thorax grey ; 
sometimes the spots mentioned appear as 4 longitudinal stripes 
interrupted at the suture. Tegule white, halteres yellowish white. 
Abdomen grey, with a dorsal black band on segments I to 3, some- 
times terminating at the base of the 3rd ; the 2nd and 3rd having 
_ each two roundish blackish spots, which are often united to the 
median stripe ; sometimes the Ist and 2nd segments have 2 large 
irregular blackish spots: also the sides of the abdomen are generally 
marked with elongated spots of irregular shape and of the same 
shade ; legs black, base of tibize reddish, wing clear: sex ° 

N.B.—No one seems to have met with this species since its 
original description, yet J include it as distinct, although it may 
quite possibly be another variation of calcitrans. The type is in 
Bigot’s collection: he does not mention the width of the frons, but 
from his doubt as to the sex of his species I am inclined to regard 
it as probably distinct from calcitrans, of which presumably he 
would be quite conversant with the respective width of the frons in 
each sex. 


Stomoxys oblongopunctata, mihi, sp. nov. 


Assam. Long. 6 mm. 

Differs from calcitrans by the wholly reddish yellow antennee 
and arista, and the pair of elongo-triangulate brown spots on the 
2nd and 3rd abdominal segments, placed on the hind margins 


74 Records of the Indian Museum. [Vorclve 


their bases on the sides of the segments, and their apices meeting 
the moderately broad concolorous dorsal stripe, which does not 
reach the posterior border of the 2nd segment. The extreme bases 
of these two segments are very narrowly brown, the 4th segment 
has a very indistinct trace of the commonly present blackish grey 
spots. Thoracic dorsum lightly yellowish grey, that of the abdo- 
men slightly bluish grey. Basal half of tibiee pale, in certain lights 
the whole of the middle pair appears pale. Frons $ width of head ; 
with face, whitish grey. Anterior ends of outer pair of thoracic 
stripes rather deeply black, enlarged into a spot of considerable 
size just above the shoulders. 


N.B.-—-This specimen is evidently quite distinct from calcitrans, 
yet Ican compare it for purpdéses of description with no other. It 
was sent to Mr. Austen, who returned it, marked, ‘‘ ? brunnipes 
Grunb. or sp. nov. closely allied.’’ It appears to me, however, 
that it can hardly be brunnipes, Griinb. (an African species, but 
which, of course, may quite easily occur in the Orient as well), 
as that species is said by its author to have a frons only § the width 
of the head, whereas in the present example the frons measures 1. 

From Grtitnberg’s description I should expect a considerably 
larger proportion of the tibize and tarsi to be brown than is the case 
in this specimen. I therefore venture to describe it as new. 

In the Indian Museum, from Sylhet, Assam, taken by Lt.-Col. 
Hall, 31-1-05. 

For purposes of comparison, in the event of my being in error 
as to its specific difference from brunnipes, I append a translation 
of both sexes of Grtinberg’s species. ; 

Originally described from German East Africa, only the ~ was 
mentioned, but immediately following this description was that of a 
supposed different species, sellata, this being now accepted as the 
2 of brunntpes. 


Stomoxys brunnipes, Griinb., 1g06. 


Zool. Anzeig., xxx, 7, 89; @ (S. sellata, Grunb.), go. 


(Description of o@ - translation.) 


Black, with brownish yellow, partly greyer dusted. 

Frons in @ one-eighth width of head, with narrow velvet- 
black stripe ; sides of frons, and the cheeks, shining yellow grey 
dusted. Antenne black, with brownish yellow arista. Palpi 
brownish yellow. 

Thorax yellowish grey dusted, grey on shoulders and between 
the black stripeson fore border. The black stripes converge in front 
of the suture, but they are sometimes separated behind it, by the 
presence of narrow intermediate spaces. 

Abdomen brownish yellow dusted, the segments with broad 
blackish band on posterior borders. When viewed from behind, 
a black dorsal line can be distinctly seen, which is expanded on the 
fore borders of the segments. 


1910. | E. Brunetti: Blood-sucking Muscide. 75 


Coxe, trochanters and femora black, extreme femora tips, tibize 
and tarsi brown; wings with a little blackish tinge; tegule also ; 
halteres yellow. 

Long. 6-7 mm. Only the ~. ‘The Camaroons, German East 
Africa. 


(Description of 2 - translation.) 


Frons in 2 not quite one-third width of head, with broad, deep 
velvet-black stripe, and broad (one-half of the width of the middle 
stripe), whitish grey, slightly glistening side borders. Under side 
of head also whitish grey, slightly glistening, Ist and 2nd joints of 
antenne black, 3rd dark brown, grey dusted, arista and palpi pale 
brown. 

Thorax above grey, with broad black stripes on each side, 
nearly confluent and nearly reaching the scutellum. 

Thorax black above, with a broad grey middle stripe: shoulders 
lightly slate-grey ; pleuree a little less grey ; scutellum black, with 
greyish brown shimmer, distinctly dusted at the tip. 

Abdomen black, rather shining, dark brown dusted on upper 
side and shimmering grey at the sides. Viewed from behind, 
elongated brown side spots become visible on the 2nd and 3rd seg- 
ments, reaching from the fore border to the middle of each segment ; 
last segment dark greyish brown dusted, unmarked. 

Coxee, trochanters and femora black ; extreme tips of latter, 
also the tibiz and tarsi, brown. Wings nearly clear, with slight 
brownish tinge, tegulae white; halteres yellowish white. Long. 
74mm. Camaroons, German East Africa. Only the @. 


Stomoxys sitiens, Rond., 1873. 
Ann. Mus. Civ. Stor. Genova, iv, 288, ° . 


This species, of which apparently the ¢ is still unknown, has 
not been definitely recorded from the East, but specimens in the 
Indian Museum collection, sent to Mr. Austen for examination 
being returned as possibly sztiens, causes me to include a descrip- 
tion of the species in this paper and to add some comments. Ron- 
dani’s original description is as follows :— 

‘““Long.5mm., 7. Similis calcitranti europe, sed minor, et 
distincta praesertim: Facie et orbitis albissimis, non sub-luteis, 
et oculis in fronte sat proximis, orbitis mediocribus vitta nigra 
angusta sejunctis, et sic oculorum distantia triplo circiter minor 
latitudine singuli oculi, non ut in mare calcitrantis trons lata fusca, 
et parum angustior singulo oculo. Praeterea in sztvente, abdominis 
grisei segmenta tria basi paulo canescentia praesertim ad latera ; 
caetera ut in sp. europea. Keren (Abyssinia).”’ 


(Tvanslation.) 


Near S. calcitrans of Europe, but smaller, and differing as 
follows: Face and eye margins extremely white, not yellowish, 


76 Records of the Indian Museum. [Vou Slys 


and the eyes in front considerably approximate, the moderately 
wide eye margins separated by a narrow black stripe, and thus the 
distance between the eyes less by about a third than the breadth 
of a single eye, not as in the @ of calcitrans, frons broad, fuscous, 
and hardly narrower than a single eye. Moreover, in siztiens the 
pubescence of the three basal grey abdominal segments is greyer, 
especially towards the sides ; the rest as in calcitrans. 

The four specimens in the Indian Museum collection tentatively 
regarded by Mr. Austen as possibly szttens were collected by me 
at Calcutta, 18-ii-05, Penang, 8-viii-o6, and Rangoon, 23-xii-04 to 
3-1-05, but they are all, unfortunately, in poor condition, and exact 
identification is, perhaps, impossible. Other specimens in my own 
collection taken by me in the same localities are in no better state 
of preservation. 

Now, Bezzi states (“‘ Mosche ematofaghe,’’ Rend. Istit. lomb., 
1907, p. 14) that he has seen a considerable number of specimens 
of satvens from Eritrea and notes that the principal difference from 
calcitvans is its narrower frons ; adding that he believes Rondani 
was in error in stating the frons to be splendid shining white, as 
he finds it has an even greater tendency to yellow than in cal- 
cutrans. 

Secondly, he, in his table of species, divides brunnipes, Griinb. , 
from sztiens, Rond., by the former having ‘‘ at least the four anterior 
tibiae all pale on the outside,’’ whilst of sitiens he says ‘‘ tibia 
at the base narrowly pale,’’ calcitrans also, be it noted, falling in the 
same subdivision in his table. The four Museum specimens being 
all ¢ @ the test of the width of the frons does not apply. The frons 
is certainly more whitish than in average calcitrans, but not more 
so than in occasional specimens of it. The specimens are in too bad 
condition for accurate description, but it may be noted that the abdo- 
minal spots are more elongate than in normal calcitrans ; a dorsal 
stripe is present on the 2nd segment, extending over part of the 
3rd ; all the tibiz are dull reddish brown (much brighter in one 
example) ; the 2nd joint of the antenne is more or less reddish. 

A second specimen in the Indian Museum from Calcutta, I4-iv- 
08, agrees with sitzens with the exception of all the tibiz being red- 
dish brown, the middle pair lighter still, almost yellowish. The 
frons is glistening shining white, one-seventh in width, and the 
abdominal spots similar to calcitrans. Rondani mentions that 
the pubescence of the abdomen is grey toward the sides, but in the 
present specimen it is uniformly black. 

It is possible it may be an Oriental form of the African species, 
but personally I have found that the wholly brownish tibie, when 
present as a character, are always consistently so. 

Likewise, in calcitrans and those species which have only the 
base of the tibize pale, the colour rarely, if ever, extends over more 
than the middle of the limb. 

That sitiens is a good species, distinct from calcitrans by the 
narrow frons, with the shining white frons as a secondary character, 
seems clear, but I doubt if any specimen of it has come before me. 


TgI0.] E. BRUNETTI: Blood-sucking Muscide. 77 


Stomoxys triangularis, mihi, sp. nov. 


@. South India. Long. 44 mm. 
Head. Frons 1, dark grey; face and antenne blackish grey ; 


5) 
proboscis rather long ; palpi pale. 

Thorax. Rather deep yellowish grey. The four deep black 
stripes narrow and separate on anterior margin, but united soon 
afterwards into two broad stripes, which are continued thus to 
posterior margin of dorsum. A black lateral stripe in front of wing. 
Sides of thorax greyish anteriorly, blackish grey behind. Scu- 
tellum very dark grey. Chetotaxy apparently normal. 

Abdomen blackish grey. Ist segment with posterior border 
somewhat darkened, 2nd with a small triangular basal spot and 
two large triangular spots occupying nearly all the segment, their 
bases almost meeting in middle of segment, their apices directed 
towards the posterior corners, 3rd segment with two similar irregu- 
larly triangular spots placed more obliquely ; 4th segment with 
two sub-triangular spots, their apices pointing hindwards. Belly 
black. 

Legs black ; knees and tibize narrowly at base reddish yellow. 

Wings and tegule pale brown, slightly deeper on anterior half. 

Described from 2 7 @ in the Indian Museum collection taken 
by Dr. Annandale in the Travancore State, South India (Madda- 
thorai, 17-xi-08, type, and Pallode, 16-xi-08). 

N.B.—An obviously distinct and quite dark species, compared 
with all others I know from the East. The abdominal markings are 
distinct (when viewed from the side) in one of the specimens, but 
the abdomen of the other is best described as blackish grey with 
irregular blackish patches, though it is certainly of the same species. 


Stomoxys indica, Picard, 1908. 
Bull. Soc. Ent. Fr. 20. 
? S. limbata, Aust. 


A translation of Picard’s description is appended :— 


Long. 44-5 mm. o. Interocular space very straight, equal 
to one-eighth the width of the head. Frons and face with more 
golden pubescence than in the @. [ines on thorax darker. Wings, 
tegule and halteres strongly smoky. lLegs testaceous, more 
brownish than in 9. 

@. Interocular space equals one-third width of head. Frons 
and face with silvery grey pubescence. Antenne black, 3rd joint 
white dusted ; arista testaceous, black at tip. Palpi testaceous, 
not reaching margin of epistome. Thorax grey ‘‘ ternate "’ 
with two moderately wide rather dark brown stripes on each side 
of middle, the outer pair interrupted at the suture. Scutellum 
wholly grey ; wings clear; veins light brown ; tegule transparent ; 
halteres white. Legs testaceous; coxe black, femora brownish, 
anterior ones lighter on inner side, darker on outer side. Abdomen 


78 Records of the Indian Museum. [Vorseive 


grey, slightly bluer than rest of body; the first 3 segments furnished 
with a straight black dorsal line and a wide black band on posterior 
margins, that on the 2nd segment covering a larger space than the 
grey part. 


N.B.—Previous to reading Mr. Austen’s description of his 
new species limbata, or seeing specimens of it determined by him, 
I had identified a number of examples of both sexes in both the 
Indian Museum and my own collection, as certainly imdica, Picard. 
On the Museum receiving back from Mr. Austen 2 7 @ and I 2? 
sent to that dipterologist some time previously, I found that he had 
referred them to his imbata. 


Apart from minor details, the only serious discrepancy in the 
descriptions of the two species appears to be in the width of the 
frons (which, however, I regard as one of the most valuable charac- 
ters) ; this being quoted by Picard as one-eighth the width of the 
head in the @ and by Austen as one-fourth to one-fifth. Now, 
although the width at the vertex is undoubtedly wider, the width 
across the frons about midway between the vertex and the base of the 
antenne is between + and 4 in the 2 7 wof the Museum collection 
determined by Mr. Austen as paratypes of his limbata; and 
in the @. All the ~ o I have seen have the frons from 1 to } in 
width (and, incidentally, ‘‘ slightly constricted in the middle,’’ 
as Austen says of limbata, although Picard mentions the frons of his 
species as being ‘“ very straight ’’). 

I therefore feel constrained to regard all these specimens ex- 
amined by me as Picard’s imdica, and Mr. Austen’s limbata as a 
probable synonym of it. A noteworthy feature of similarity is that 
both authors mention the extra width of the marginal black band 
on the 2nd segment, and a final coincidence is that both authors 
describe their species from the same three localities, India, Ceylon 
and Assam. 

S. mdica occurs in the- Indian Museum collection from 
Calcutta, 18-vi-08 [Annandale] ; Rajmahal, Bengal, 7-vii-o9 [A nnan- 
dale|; Bhogaon, N. Bengal, 3-x-og [Paiva]; Allahabad, I5-viii-09 
[Lord]; Maddathorai, Travancore State, 16-xi-o8 [Ammnandale] ; 
Victoria Gardens, Colombo, 26-iv-08 [Paiva] ; Rangoon, 24—26-ii-08 
[Annandale]; Singapore, 1I-viii-o6 [Brunetti]. In my collection 
I have it from Calcutta, 24-vii and 14-viii-o4, and from Rangoon, 
18-viii-o6, taken by myself. 


Cole 


Stomoxys limbata, Aust., 1900. 
Ann. Mag. Nat. Hist. (8), iii, 292. 


7 2. India, Ceylon, Assam. 


~”. ‘‘ Smoke-grey: face and sides of front bright yellowish- 
silvery, front narrow; dorsum of thorax with usual clove-brown 
longitudinal stripes, admedian stripes narrow and wide apart ; 
dorsum of abdomen with deep clove-brown or blackish transverse 
band on hind border of each of first three segments ; wings with a 


IgI0. | E. BRuNETII: blood-sucking Muscide. 79 


brownish tinge ; femora dark clove-brown, their extreme tips and 
base of hind tibiz ochraceous-buff, front and middle tibiae and 
tarsi mummy-brown, hind tarsi and hind tibiz except base sepia- 
coloured.’’ 

@. ‘* Apart from usual sexual differences, agreeing essentially 
with @ , except that dark markings on dorsum of thorax and abdo- 
men are paler (olive-brown instead of clove-brown or blackish), 
that the wings although brownish are less noticeably infuscated, 
that the squamee are paler (whitish or yellowish white), and that 
the front and middle tibie are also brighter in hue.’’ Type in 
British Museum from Sylhet, r1-ii-o5 [Lt.-Col. Hall|; also a 
specimen from Peradeniya, Ceylon, 22-v-92 [ Yerbury]. 

‘* S. imbata is closely allied to S. nigra, Mcq., but is distinguished 
by the front in the ~ being somewhat narrower, with the frontal 
stripe slightly constricted in the middle, instead of having its sides 
parallel ; and in the @ by the front and middle tibiz being mainly 
ochraceous instead of for the most part clove-brown or blackish.’’ 

The length of this species is from a little under 5 mm. 
to 54 mm. ; width of head 7 16 to 2 mm., 2? 1'8to2mm.; width 


d 
of front at vertex 7 ,0'4mm., 2 just over 0°5 mm. 


N.6b.—Specimens seen by Mr. Austen and identified by him as 
his /imbata are in the Indian Museum collection from Port Canning, 
Lower Bengal, 21-vii-o7 [Annandale]; Calcutta, 20-viii-o7; and 
Sylhet, 31-i-05 [Lit-Col. Hall]; Mr. Austen. also refers to this 
species a ¢ from Peradeniya, Ceylon, 22-v-g2 [Lt.-Col. Yerbury}. 

N.B.—As noted under S. zndica, Pic., I am strongly inclined 
to regard /imbata as synonymous with that species. 


Stomoxys bengalensis, Pic. 
(Translation.) 


@. Interocular space one-fifth width of head. Frons and 
face with whitish pubescence, lightly gilded. Antenne black, arista 
testaceous, tip black ; palpi pale yellow. 

Thorax grey, with two parallel black stripes on each side of a 
very indistinct median line. Scutellum rather dark grey. Wings 
a little smoky, veins blackish brown ; tegule and halteres dark 
brown. Legs black, base of tibiz pale brown. Abdomen light 
slaty brown, with black dorsal line on first three segments ; each 
segment with a narrow basal transverse band, and a wide one on 
posterior margin, especially that on the 2nd segment ; 4th segment 
wholly grey (in fresh specimens), long. 7} mm. ? unknown. 
Sent by Col. Raymond from Calcutta. 

N.B.—S. bengalensis should, by the description, be a good 
species, and I had noted three @ in the Indian Museum collection 
(from Calcutta) as being probably this species. They were, however, 
returned after examination by Mr. Austen as calcitrans, and closer 
study both of them and subsequently acquired specimens lead me 
to suppose they fall within the range of the commoner species. 


80 Records of the Indian Museum. [ VoL." EV, 


If so, however, the approximation to the description of bengalensis 
is considerable, as the abdominal spots are sometimes spread out, 
so as to form a sort of band on the posterior margin, and there are 
in some examples a narrow dark band at the base of at least the 
2nd and 3rd segments. 


Stomoxys pusilla, Aust., 1909. 
Ann. Mag. Nat. Hist. (8), iii, 293. 


@. India. Long. 4:2 to 4°5 mm.; width of front at vertex 
o'4 mm. 

‘“ Dorsum of thorax mouse-grey or yellowish grey, with usual 
clove-brown longitudinal stripes ; dorsum of abdomen olive-grey, 
first segment, except a small ill-defined area in centre (not reaching 
hind margin), second segment, either entirely or with exception of 
extreme front margin, and a median longitudinal stripe and fairly 
deep posterior transverse band on third segment, clove-brown ; sides 
of front and sides of face, when viewed from above, light maize- 
yellow ; wings brownish; femora clove-brown, front and middle 
tibie and tarsi and extreme tips of front and middle femora buff, 
hind tibiz and tarsi light mummy-brown, base of tibize paler.’’ 

Type in British Museum, Allahabad, x-05 [Howlett]. 

““S. pusilla differs from S. limbata in its smaller size, in the 
yellower colour of the sides of the front and face, in the dorsum 
of the second abdominal segment being entirely clove-brown or practi- 
cally so, instead of having a clove-brown posterior transverse band 
and median longitudinal stripe, and in the pale tibize and tarsi of the 
front and middle legs.’’ 


N.B.—This species and the next I have never seen. The 
present one must approximate very closely to indica, Pic.; pulla 
is, of course, well defined by its fore tarsus. 


Stomoxys pulla, Aust., 1g09. 
Ann. Mag. Nat. Hist. (8), ili, 204. 


@. India. Long. 4:2 to 4.4 mm. 


‘“ Very dark species ; when viewed from above, body appearing 
almost uniformly clove-brown, extreme front margin of thorax 
mouse-grey, with commencement of usual clove-brown longitudinal 
stripes ; when abdomen is viewed from behind, at a very low angle, 
terminal segment appears olive-grey, while second and third segments 
may appear more or less mouse-grey, with clove-brown transverse 
blotches, and perhaps a trace of a narrow median longitudinal 
stripe ; front narrow, occupied for most part by frontal stripe, 
sides of front not noticeable except anteriorly ; wings brownish ; 
legs clove-brown, tibiae ochraceous-buff at extreme base, first joint 
of front tarsus fringed on inside with a row of hairs of equal length, 
claws black.’’ 


IgIo.| E. BRuNETI1: Blood-sucking Muscide. SI 


Type in British Museum, from Mussoorie, N. W. India, x-06 
| Howlett). 

‘“ This is a very distinct species, which, while resembling the 
foregoing (pusilla) in size, is at once distinguishable by the 
dusky coloration of the body and legs, by the sides of the front 
being scarcely visible except anteriorly, and by the remarkable 
row of hairs on the inside of the first joint of the front tarsus. 
Owing to the latter character S. fulla o~ presents some slight ap- 
proximation to the ~ of the African S. omega, Neust., in which, 
however, the row of hairs on the inside of the front tarsus extends 
to the end of the second joint, while the hairs themselves are much 
longer and conspicuously curled.’’ 


Stomoxys, sp. 


A single ~ in the Indian Museum, apparently immature, 
certainly belongs toa distinct species from all others given in this 
paper. It has a frons which narrows rapidly from the vertex 
(where it is about } the width of the head) downwards, being 
only between 4 and + at a short distance above the antennz, where 
it widens as usual. ‘The outer side of the middle tibize are wholly 
moderately light brown, and the abdomen is apparently unmarked, 
but being in a somewhat crumpled state it is impossible to describe 
it more precisely. In other respects it resembles a slightly under- 
sized calcitrans. From Calcutta, 18-vii-07. 


Stomoxys dacnusa, Speis., 1908. 
Zool. Anz., xxxiii, 666. 


‘“ Orichalceo griseo, vittis duabus brunneoviolaceis, pedibus 
luteis, coxis atque femoribus preter ultimam tertiam partem 
brunneo piceis, alis basi et nervatura validiora luteis squamis lutes- 
centibus. Annam.’’ 

2. Long. corp. 5°5 mm. ; alarum 5°5 mm. 

Further notes from his more lengthy description in German 
may be added as follows :— 

Yellowish grey, with a brassy tinge. From the anterior border 
of the thoracic dorsum, a moderately broad brownish black, with a 
violet tinge, stripe towards each side. 

Unfortunately the hinder part of the thorax, the pleure, chest, 
and first segments of the abdomen cannot be correctly described, 
owing to the blood which has exuded over them. ‘Tip of scutellum 
and of abdomen uniformly grey. Legs pale brownish yellow ; only 
the coxee, and basal two-thirds of the femora, blackish brown or 
nearly black. 

Head blackish brown, only the base of the antennz paler, 
yellowish brown, the antennz, with the arista, tawny brown. 

Frons, almost exactly one-third width of head at vertex, 
widening in front. Chetotaxy normal. Palpi very little spindle- 


82 Records of the Indian Museum. [Vor.-1v 


shaped at tip, tawny. Fore femora with the brown colour nearly 
reaching the tip, under side with a row of longer bristles, in addition 
to the normal hairs. Middle femora above, towards the tips, 
with some small ones, hind femora with rather larger, distinct 
bristles, below with some preapical ones. Hind tibize with two 
bristles near the middle. Hind metatarsus with some stronger 
bristles below. Wing as in calcitvans; Ist longitudinal vein 
(subcosta) ends exactly above the small cross-vein ; 2nd (radius) 
rather nearer the wing tip than in calcitvans. 


N.B.—Being described from a single damaged specimen, 
the specific identity of this species must be admitted with caution; 
I therefore place it Jast of all. 


Description of STYGEROMYIA, Aust., 1907 (non-Oriental). 
Ann. Mag. Nat. Hist. (7), xix, 445. 


‘“In some respects intermediate between Stomoxys, Geoft., 
Hematobia, Rob. Desv., and Lyperosia, Rond. (sensu Bezzi). 
In general appearance and form of body similar to Stomoxvs, but 
in shape of proboscis and palpi resembling H@matobia, though with 
arista feathered only on upper side, as in Stomoxys and Lyperosia. 
Head somewhat flattened from front to rear, with basi-occipital 
region only slightly swollen; proboscis short, stout and shining, 
of uniform thickness throughout, not tapering to the tip, chitin- 
ous, but terminated by a pair of small fleshy labella ; palpi equal 
to proboscis in length, large, clavate towards the tips, curving up- 
wards, and with stout bristles on the outer side at the distal 
extremity. 


‘“* Bristles of thorax : Humeral 3 ; post-humeral 1 ; notopleural 
2; presutural 1; supra-alar 1; intra-alar 1; post-alar 2; dorso- 
central 6 (1 in front of and 5 behind the suture) ; inner dorso-cen- 
tral 1; scutellar 4 (1 preebasal, 1 basal, 1 discal, I apical); meso- 
pleural 9 or 10, wider apart than in Stomoxys or Hamatobia ; sterno- 
pleural 1 (posterior, as in Stomoxys, instead of 1: I, as in Hema- 
tobia). 

“Wings with 1st posterior cell narrowly open at the tip, the 
width of the opening being precisely that seen in the same cell in 
the wing of Musca corvina, Fab., and less than half of that exhibited 
by the ist posterior cell in the case of Stomoxys calcitrans, 1. : apical 
portion of 4th vein beyond the bend Perfectly straight, not bent in- 
wards at the extremity, as in Sfomoxys and Hematobia. Typical 
species, Stygeromyia maculosa, sp. nov.” 


N.B.—This is not an Oriental genus, being described from 
Africa, but since, owing to the traffic in cattle between one con- 
tinent and another, it may easily be imported, I have thought it 
best to include at least the generic description to facilitate identi- 
fication should it occur in the East. 


1910. | E. Brunetti: Blood-sucking Muscidae. 83 


LY PEROSIA, Rond., 1862. 
Dipt. Hal. Prod., v, 230. 
Loc sctt: | 1; 93 (nom? in tabs): 


In splitting off this genus from Hematobia, R. Des., Rondani 
in a table of genera gave the following diagnosis of it: ‘‘ Arista 
hairy only on upper side, the hairs wavy; 2nd longitudinal vein 
reaching the costal opposite the small cross-vein.’’ In this genus 
Rondani placed two species, ‘‘ ivritans, I,.,’’ and ‘‘ serrata, R. D.”; 
Prof. Bezzi very kindly supplied me with notes on this genus from 
Rondani’s work, and informs me that that author was mistaken in 
his identification of ivvitans, I,., having another species before him 
at the time, and for this species my correspondent proposes the 
name of titillans (Rend. Istit. lomb. (2), xl, 24). 

Moreover, Rondani’s “* serrata, R. D.’’ was the true zvritans, L,. 


LY PEROSIA, Rond. 
(Revised description.) 


Body more elongate than in Stomoxys ; abdomen about twice 
as long as broad. Eyes bare, approximate in @ , frons in @ about 
2 width of head. Antennze according to Rondani inserted at the 
middle of the eyes, with the 3rd joint a little longer than the 2nd, 
but I should describe them as inserted very slightly lower on the 
face than in Stomoxys. Antennee plumose on upper side only, the 
hairs undulating. Proboscis as in Stomoxys ; palpi comparatively 
stout and broad, as long as proboscis, less bristly than in Hematobia. 
Apical part of 4th longitudinal vein curved rather less than in 
Stomoxys and Hematobia, the whole vein being considerably 
straighter, and without the definite ‘‘ elbow’’ so usual in the 
higher Muscine; 3rd longitudinal vein bare at base ; Ist posterior 
cell comparatively broadly open; sternopleural bristles yellow. 


Table of species. 
Hind tarsi not dilated. 
Abdomen without a dorsal stripe .. muinuta, Bezzi, 7 2. 
Abdomen with a dorsal stripe (some- 
times indistinct). 
Hair on body brown .. exigua, Meij., 2. 
Hair on body yellow .. flavolurta, sp. nov., 2. 


Hind tarsi dilated. 


Hair on body brown its .. exigua, Meij.,o. 
Hair on body yellow om .. flavolurta, sp. nov., o. 


84 Records of the Indian Museum. [VoE. EVs 


Lyperosia minuta, Bezzi, 1892. 
Ann. Mus. Civ. Genova (2), xiii, 192. 
¢ @. India, Ceylon (originally described from Somaliland). 


‘‘Cinerea, lutescenti-pilosa, pedibus luteis; antennarum 
articulo secundo rufescente ; oculis etiam in mare a fronte latius 
cula sejunctis. Long. mm. 7 1'8—2°3, 2 3—3'8 Obbia.’’ 


(Redescription.) 


Head. o&. Eyessub-contiguous ; the frons for a short distance 
being extremely narrow, the actual width varying slightly in 
different specimens. Vertical triangle only moderately wider ; 
the frons below its narrowest part, rapidly widening. Normal 
colour of frons dark grey or blackish grey in the middle and lighter 
grey along the edges of the eyes, which latter are without any dis- 
tinct eye margin. For a considerable space above the base of the 
antenne the frons is occasionally quite reddish or reddish brown. 
Lower part of the head, including oral cavity, yellowish grey, 
varying lighter or darker according to the individual. Ocellar 
triangle (isosceles) distinctly raised above the surface, concolorous, 
ocelli distinct. Antenne grey, sometimes with a slight reddish 
brown tint, microscopically pubescent ; 2nd joint sometimes much 
brighter, always with a short distinct bristle ; 3rd joint more or less 
grey dusted. Arista black considerably thickened at the base, 
where it is normally black also. The arista bears normally at least 
6 long equidistant, bisinuate hairs ; with often an additional one 
placed just before the first, and also a very small one near the tip. 
There are invariably several very minute bristles at the base. 
Palpi elongate, black or very dark blackish brown, gradually 
thickened towards tip, with short stiff hairs ; practically as long as 
the proboscis, which is dark brownish yellow, shining. Back 
of head moderately dark grey, with microscopic hairs. 

Chetotaxy. Seven fronto-orbital bristles; mostly curved, 
and often pointing in different directions. Vertical triangle with 
2 stronger longer bristles in front, crossing one another, and 2 
smaller ones behind, which also cross each other, with 2 other 
similar ones practically on the vertical margin of the head, with 
I pair each side of these, of true vertical bristles. 

@. Frons at vertex, one-fourth width of head, widening 
only slightly, as at the level of the antenne it is only one-third the 
entire width. 

Fronto-orbital bristles normally arranged in two rows, an upper 
row of four, placed closer to the margins of the eyes, the bristles 
directed forwards or slightly to either side ; and a lower row (these 
would be the lower fronto-orbital bristles of Osten Sacken), of either 
three or four, which is placed nearer the middle of the frons, and in 
which the bristles always curve inwards. The rows sometimes 
almost form one linear row, or occasionally overlap, and sometimes 
the upper row consists of three bristles only ; in some individuals 


1910. | E. Brunetti: blood-sucking Muscide. 85 


the number of bristles on one side of the face is different to that on 
the other side, in fact this occurrence is not uncommon. 

Thorax. Variable, within certain limits. Dorsum rather 
dark acuous grey, with a more or less distinct but nearly always 
obvious, broad light grey stripe beginning in the middle of the 
anterior border and fading away at or before the transverse suture. 
Humeral calli light grey ; the transverse suture, and more or less 
of the scutellar dorsum, also. In other examples, the dorsum is 
best described as being yellowish grey, with two broad, well-separa- 
ted, well-defined stripes from the anterior border nearly to the 
scutellum ; with sometimes an additional narrow intermediate stripe 
behind the suture. 

The sides of the thorax are lighter, ash-grey or yellowish grey. 
Metanotum dark blackish grey, practically bare. 

Chetotaxy. The principal bristles are nearly always uniform 
in perfect specimens, but are very easily broken off. They are 
often smaller than their normal size, and in that case sometimes are 
not conspicuously larger than the small bristly hairs with which the 
thorax is covered. Allthe bristles are brownish yellow and generally 
reclinate. Humeral bristles 2; post-humeral I; presutural Ir ; 
notopleural 2; supra-alar 1; intra-alar 1; post-alar 1. A_ pair 
of inner dorso-central bristles, and a row on each side of compara- 
tively small but distinct dorso-central bristles, the hindermost 
being as usual the largest ; whilst outside this row is a second row of 
microscopic ones which only extends from the transverse suture to 
the posterior border. The normal row of dorso-central bristles is 
continued to the anterior border. Along the median line of the 
thoracic dorsum are numerous microscopic bristly hairs, of which 
those in front of the suture are arranged more or less in four rows, 
while those behind it are more irregularly placed. 

On the scutellum there are two basal bristles, placed near the 
edge, a post-median row of four, of which the inner ones are less 
strong ; and two strong long apical ones. Over the whole of the 
thoracic dorsum and the scutellum are microscopic bristly hairs, 
which are more or less irregularly placed. 

On each side of the thorax there is a row of 6 (occasionally 7, 
or even 8) on the hinder side of the mesopleura, and 3 very distinct 
sternopleural ones, placed 1: 2. Smaller bristly hairs occur irregu- 
larly, being much more numerous in some specimens than in others. 

Abdomen. Rather dark grey (in some examples much lighter) ; 
with short, brownish yellow hairs, curved backwards ; and a row 
of long bristly curved hairs towards the posterior border of each 
segment. Belly dark grey, sometimes reddish (probably in the 
case of specimens captured soon after having sucked blood) ; the 
dorsal bristly hairs continued over the sides to a considerable 
extent. 

Genital organs in the ~ apparently consisting of a telescopic 
arrangement of three pieces, brown, with blackish marks and some 
hairs; the tip hairy. Ovipositor in the @ conically elongated, 
concolorous, protruded, pubescent. 


86 Records of the Indian Museum. [Vor nbs 


Legs. Coxe, rather bluish grey dusted, reddish yellow towards 
tips, with a row of 5 or 6 bristles. Femora rather brownish or 
blue-grey dusted, reddish yellow at base and tip; with rows of 
short bristly hairs. Fore femora generally curved, and with a row 
of comparatively long bristles on outer and inner sides, the latter 
row being the longer ; the rest of the fore femora being clothed in 
rows of bristly hairs. Middle femora with rows of very short 
bristles, and some longer ones near the base on under side, and two 
on upper side near tip. Hind femora, sometimes somewhat curved, 
with rows of small bristles and a row of long comparatively weak 
ones on under side, and a strong one on outer side near tip. 

All the bristles on the legs brownish yellow. 

Tibiz brownish yellow or yellowish, closely pubescent, with 
very short stiff hairs. Tarsi yellowish, blackish towards tips, micro- 
scopically pubescent. Pulvilli greyish white. 

Wings. Practically clear, iridescent in certain lights ; micro- 
scopic bristles along costa to beyond end of 2nd longitudinal vein ; 
very soft hairs on remainder of wing border. Inner cross-vein ex- 
actly opposite end of 1st longitudinal and placed above centre of 
discal cell. Third and 4th veins converge normally towards tips. 
Halteres and tegule yellowish grey. 

Described from about 12 specimens of each sex in the Indian 
Museum, mostly from the Victoria Gardens, Colombo, taken by 
Mr. Paiva, 26-iv-08. Other specimens are from Chittagong, 27-v11-08, 
8-ix-o8 [Lt.-Col. Hall|; Mandalay, 12-11-08 [Annandale]; Sukna 
(500 ft.), 3-vii-o8 [Annandale], where it was abundant on cattle ; 
Calcutta, 6—3I-vili-09. 

N.B.—Dr. Annandale noticed that the fly rests with its wings 
folded together, as contrasted with exigua, which when at rest, 
extends its wings horizontally. 


Lyperosia exigua, Meijere. 
(Tvanslation.) 


Hematobia exigua, Meij., in P. Schat, “‘ Verdere Mededeelingen 
over Surra,’’ Mededeel. Proefstation Oost-Java, 3e. Ser., No. 44 
(1903). 

Hematobia exigua, Meij., Bijd. tot se Dierk. Natura Artis 
Magistra, xvili, p. 104 (1904). 

Front in @ very narrow, with very narrow white eye margins, 
between which a narrow black line is present. Frontal triangle 
black. In the @ the white eye margins separated by a black 
frontal stripe of uniform width. Under side of head white. An- 
tennee brownish yellow, especially the somewhat thickened 2nd 
joint, which is thereby conspicuous ; the 3rd joint darkened at tip. 
All the joints with whitish shimmer. Palpi as long as the pro- 
boscis, brownish yellow. Proboscis brownish yellow, darker above. 

Thorax dark grey, whitish dusted, in front with two narrow 
blackish longitudinal stripes. The sides of the dorsum, viewed 
sideways, broadly brownish tinged. Sides of thorax, humeral calli, 


1910. | E. Brunetti: Blood-sucking Muscide. 87 


transverse suture and a cross-band on the scutellum, whitish 
shimmered in certain lights. 

Abdomen dark grey, with a blacker (ending at the tip of the 
3rd segment) dorsal stripe, which is only wholly visible when 
viewed from certain directions. Sometimes it appears in the form 
of darker spots. 

Legs brownish yellow, on the outer side more or less darkened ; 
sometimes the hind pair, with the exception of the outer basal 
part of the femora and tibiz, quite dark ; in the fore legs, with the 
femora, nearly to the tips; the middle third of the tibize and the 
tarsi tips, sometimes darker. 

Veins as in izvritans, I,., the small cross-vein straight, placed 
beyond the tip of the 1st longitudinal vein. Halteres and tegule 
whitish. 

Length of body 4 mm.; wing length 4 mm. 

According to Herr Schat, this species, together with Stomoxys 
calcitrans, I,., may be capable of transmitting disease. 


(Redescription.) 


Head. Frons in ~, 1 to % width of head; in 9 between i 
and +. Ground colour almost wholly ash-grey, except the black 
frontal stripe, which is very narrow, yet quite distinct, in the o, 
and forms in the 2 one-fourth of the width between the eyes at 
the level of the ocellar triangle, gradually widening to one-third at 
the level of the base of the antenne. 

The frons has inthe ~ a row, normally of nine, distinct, well- 
developed brownish vellow or yellow bristles, arranged practically 
in a single row, mostly curved inwards towards those of the op- 
posite row, especially the lower ones. 

In the ¢@ they are more irregularly placed, forming normally 
an upper frontal row of 3 or 4, situated rather nearer the eyes, and 
pointing outwards or forwards ; and a lower row (the lower fronto- 
orbital bristles of Osten Sacken) of 4, 5 or 6, situated nearer to the 
frontal stripe, also placed rather closer together, and always curved 
inwards. ‘These two rows, the upper and the lower one, are not 
always placed in the same relation to each other. Those in the 
upper row are more liable to be placed out of line, the lowermost 
bristle being often on a level with, or even below, the uppermost 
bristle of the lower row. All the bristles are always brownish yellow 
or yellow. 

Antenne yellow, 3rd joint very slightly white dusted ; arista 
often distinctly yellow at extreme base. Palpi yellow, with a few 
haits. Proboscis dark brownish or brownish yellow. — Bristles of 
cheeks yellowish, or whitish yellow; lower part of head with some 
pale hairs ; back of head with numerous very short bristly hairs. 

Thorax. Dorsum normally moderately dark grey ; often with 
either a bluish, brownish or sub-aénous tinge; sometimes almost 
ash grey, especially in the median portion. ‘There is generally a 
fairly distinct pair of narrow well-separated fuscous stripes, a 


88 Records of the Indian Museum. [Vors LV: 


sub-triangular brownish spot towards each side in front of the suture, 
and a similar but larger and longer spot behind the suture each 
side. The macrocheetze of the thorax are practically the same as in 
minuta, but the very small bristles covering nearly the whole surface 
are relatively larger than in the previous species. ‘The large bristles 
are brownish yellow, the smaller ones blackish. Sides of thorax ash- 
grey or with a bluish tinge, mesopleura with a row of long yellow 
bristles on hinder side, as in minuta, and smaller ones in front ; 
sternopleura with numerous bristly hairs, and a row of curved 
yellow long hairs below humeral! calli. 

Scutellum concolorous with thorax, macrochete as in minuta, 
but the smaller bristly hairs more numerous and much stronger. 
Metanotum blackish grey, bare. 

Abdomen. Rather dark grey, lighter at base, where a distinct 
blackish dorsal narrow stripe extends over the Ist segment to some 
portion of the 2nd. The whole surface of the abdomen is covered 
with short hairs, and there is a row of well-separated long curved ones 
towards the posterior border of each segment, and a fan-shaped 
bunch of yellow ones towards each side of the Ist segment ; all the 
other hairs and bristles on the abdomen being distinctly brownish 
or brownish yellow. Belly concolorous, pubescent. 

Legs. Mainly yellow, coxze bluish ash-grey or yellowish grey, 
-femora except base and tip often more or less lightly violet-grey 
tinted, sometimes almost wholly yellow. All the femora bear 
rows of rather long soft yellow hairs, the fore pair have a row of 
longer bristly hairs on upper side and a row of still longer ones on 
under side ; middle pair with two stronger bristles on upper side, 
near tip; hind pair with three stronger ones below, near tip. 
Tibize, which are sometimes a little brownish, closely pubescent, with 
comparatively short bristly, yellow hairs, a long bristle in front at 
tip of fore pair, some short stiff bristles at tip of middle pair, and a 
long bristle on hinder side near tip of hind pair which are darker 
than the others. Tarsi mainly blackish, bristly, pubescent ; meta- 
tarsus and bases and tips of the remaining joints more or less 
yellowish. Second and 3rd tarsal joints (ino ) distinctly dilated at 
tip, each with 3 or 4 long curved hairs on under side, and generally 
all the joints with one straight hair and one longer curved one on 
the upper side at the tip. Pulvilli whitish, grey fringed claws black. 

Wings. Asin minuta. 

Described from a good series of both sexes taken in the slaughter- 
house in Calcutta, 6—3I-vili-og. Also from several of each sex 
from Travancore State, South India, captured by Dr. Annandale 
as follows: Nedumangad 14-xi-08 ; Quilon 10-xi-o8 ; Maddathorai 
17-x1-08 ; Pallode 15-xi-o08, nearly all on cattle and buffaloes. Also 
from specimens from Sukna (500 ft.}, 3-vili-o8 [Annandale]; Chitta- 
gong (E. Bengal), 2-viii-o8 and 3-ix-08 [Lt.-Col. Hall]. Allthe above 
specimens are in the Indian Museum collection. Two? ¢? inthe Pusa 
collection were taken at Bombay on cattle, 29-v and 16-x-05, and 
I have a considerable number of both sexes in my own collection 
from Calcutta and West Java. 


IQIO. } E. BRUNETTI: Blood-sucking Muscide. 89 


N.B.—Dr. Annandale has noticed that this species when at 
rest, sits with its wings spread out horizontally. 


Lyperosia flavohirta, mihi, sp. nov. 


@” @. Lower Burma. Long. 23-3 mm. 


This species is very near exigua, the differences all being minor 
ones, vet I believe, by taking them in the aggregate, a specific 
separation for this form is warranted. 

1. The lighter coloured parts of the head and thorax are more 
whitish than in exigua. 

2. The small bristly hairs covering the thorax, scutellum and 
abdomen are distinctly more yellow than brownish yellow. 

3. The dorsal stripe on the abdomen is generally continuous 
almost to the tip, and is more uniformly wide ; whereas in exzgua 
it often terminates on the posterior border of the 2nd segment, 
sometimes appearing on the Ist segment only, and often (when 
nearly complete in length) takes the form of elongated triangles, 
bases wppermost. 

4. The legs are paler yellow and the whole body is paler. 

Described from 6 @@ and 2 9 2 taken by Dr. Annandale 
on cattle at Kawkareik, Lower Burma, I-111-08. 


N.B.—Dr. Annandale noticed this species in great abundance 
on cattle in the interior of the Amherst District in Tenasserim, 
March 1909. He remarked that when at rest it stood higher on its 
legs than other Indian species of the genus, thus approximating to 
the appearance of a Dolichopodid. 


Lyperosia irritans, L,. 


As this paleearctic species may quite possibly be found in 
Northern India, a brief description is appended, drawn up from 
the only two specimens before me, 2 @” @ in my own collection, 
from Roumania. 

Dark yellowish grey, 3} to 4 mm. long with 4 not very well- 
defined thoracic blackish stripes, of which the middle pair attain 
the anterior margin. Abdomen with blackish dorsal stripe on 2nd 
segment. Palpi distinctly spatulate at tip, flattened, nearly as 
long as the shining black proboscis. Frontal space } width of head, 
mainly occupied by whitish eye margins, but leaving a very narrow 
black stripe. Legs black, knees pale, hind femora with a single 
hair below, just beyond middle, and another at tip; hind tibize 
with some comparatively long hairs on inner sides. Wing clear, 
Ist posterior cell very narrowly open ; outer cross-vein long, nearly 
straight. 


PHILAMATOMYIA, Aust., 
Ann. Mag. Nat. Hist. (8), 111, 295. 


1909. 


‘“ Greyish flies, not unlike Musca domestica, Linn., in general 
appearance, but distinguishable at once, owing to the remarkable 


go Records of the Indian Museum. [Voy. IV, 


proboscis. Front in @ narrow, its width in centre being from 
one-eleventh to one-fifteenth of total width of head ; width of front 
in @ at vertex, one-third of total width of head ; proximal portion 
of proboscis (mentum) a strongly swollen chitinous bulb, distal 
portion soft and fleshy, folded back under distal end of bulb when 
not in use, but when in use extended, its terminal section consisting 
of a ‘tubular extension’ which is protruded from between the 
labella, and is surrounded at the distal extremity with a circlet 
of stout chitinous teeth ; venation generally as in Musca domestica.’’ 

Mr. Austen proceeds to give a more detailed generic description 
and adds the following chetotaxy :— 

‘“ Thoracic bristles: humeral 3 (in co sometimes 4); post- 
humeral I ; notopleural 2; presutural 1; supra-alar I; intra-alar 
I; post-alar 3; dorso-central 4 or 5 (the large and conspicuous 
bristles alone included—z or 3 in front of, 2 behind suture) ; inner 
dorso-central 1; scutellar 4 (1 preebasal, I basal, I discal, I apical) ; 
mesopleural, normally 6 (space between uppermost two greater 
than that between any other two) ; sternopleural 1: 2.’’ 


Philematomyia insignis, Austen, 1909. 
Loc. cit. ante, 208. 


@7 @. India, Ceylon, Sokotra, Cyprus, Senegal and Congo 
Free State. 

Length of 17 7 @ 3—5'8 mm. ; of 25 2? 9 4—64 mm. 

‘* Smoke-grey to isabelline-grey, or yellowish grey ; dorsum of 
thorax with 4 dark longitudinal stripes, broader and clove-brown 
or blackish in & , narrower and dark grey or brownishin 2 ; often 
with an additional broad median stripe, most distinct in front and 
behind; dorsum of abdomen with a tranversely elongate dark 
blotch on Ist segment, not reaching hind margin, and a small clove- 
brown median triangle at base of 2nd and 3rd segments, in some 
specimens also with a similar but smaller and usually lighter mark 
at base of 4th segment ; all hair and bristles on head, body and legs 
black ; palpi ochraceous-buff ; wings hyaline or with a slight ochra- 
ceous tinge, veins in proximal half buff or ochraceous-buff ; legs 
black, femora grevish pollinose.’’ Types of ~ and ? from India 
in the British Museum. Mr. Austen records it from the East from 
Calcutta, 23-vi-o7, ‘‘on draught bullocks, burrowing into the 
hair ’’ ; Mussoorie, ix-0o6 ; Allahabad, 11-x-05 [Howlett]; Bentota, 
Ceylon, 13-vi-g1 [Yerbury]; Daukotuwa, Ceylon. ‘The above data 
derived from specimens in the British Museum. 


N.B.—This species (identified by its author in the case of some 
of the specimens examined) figures in the Indian Museum from 
Calcutta (where it is common also from May to September), 15-ii-08 
[Annandale] and 5-iii-05 [Brunetti]; Chittagong, 14-vii-o8 [ Lt.-Col. 
Hall|; Mandalay, 13-iii-o8; Kawkareik (North ‘Tenasserim), 
5-11-08, on cattle ; Rangoon, 25-ii-o8 ; and Port Canning (30 miles 
south of Calcutta), 21-vii-o7, and Quilon, S. India, 9-xi-08 [all 


IQI0. | E. Brunetti: Blood-sucking Muscide. gl 


Annandale]; Puri; Bengal, 2-11i-08, Purneah, N. Bengal, 7-viii-07, 
Madhupur, Bengal, 15-x-0g, and Victoria Gardens, Colombo, 26-iv-08 
fall Paiva}; Cawnpore, xii-04 [brunetti|; Allahabad, 12-viii-o9 
[Lord]; Purulia, Bengal, 13-x-og. I have it in my own collection 
from Rangoon (both sexes not uncommon), 23-x1i-04 to I-111-05, and 
18-viii-06 ; Calcutta, 24-vii-o4 and 22-viii-o4 ; Jubbulpore, India, 
15-xi-07, and Telok-Ayer, N. Borneo, 29-vii-06, all the specimens 
taken by me. 


PRISTIRHYNCHOMYIA, mihi, gen. nov. 


With the exception of an important modification of the pro- 
boscis, identical with Philematomyia, Aust., the general characters, 
the venation and cheetotaxy agreeing exactly. 

The two parts of the proboscis, however, are structurally re- 
versed, the wide basal part being fleshy and flexible, the second 
part (of about equal length) being sub-cylindrical, black and 
distinctly chitinized, possibly retractile to the extent of its with- 
drawal partly or wholly within the fleshy basal portion. At the 
end of the chitinous portion is a soft fleshy tip, the terminal 
orifice being of the shape of a triangle with a rounded base (the 
edges being thickened somewhat by a rim bearing the teeth). At 
the apex of the triangle is a single black tooth, whilst arranged 
around the orifice above are three pairs of similar black teeth. 

Under high microscopic power the apparent ‘‘rim”’ of the 
orifice is seen to be the base of each tooth extended considerably 
on each side, so that the ‘‘ rim’ is not continuous. 

The new genus is intermediate between Philematomyia and 
Musca, but the presence of the teeth suggests that it can hardly 
be other than a “‘ biting fly.” 


Pristirhynchomyia lineata, mihi, sp. nov. 


@. Calcutta. Long. 4-5 mm. 

Head. Yellowish grey, whitish grey when viewed from below. 
Frons at level of antenne } width of head, slightly narrower at 
vertex ; with a broad dull black median stripe (widest in the middle). 
Vertical triangle blackish, distinctly raised, with 3 pairs (rarely 4) 
of bristles, the upper pair small, the lower pair strong and long. 
Normally 7 strong fronto-orbital bristles placed along the edge of 
the black centre stripe, above these are two very minute ones, with 
a powerful one above these, this latter one being level with the 
centre of the ocellar triangle. ‘The upper strong bristle, and the 
7 large fronto-orbital ones are always present. 

Outside this row is an irregular row of very small but distinct 
bristles. T'wo minute vertical bristles, and a pair of strong ones on 
vertex near the upper corner of each eye. Antenne blackish grey, 
3rd joint lighter ; 2nd joint with several short stout bristles and 
one much stronger one, Arista considerably thickened at base ; 


) 


92 Records of the Indian Museum. [Voy. IV 


7 or 8 nearly straight long hairs on upper side and 4 or 5 on lower 


Fic. 1.—Proboscis of Pristivhynchomyia lineata; 
dorsal view of specimen mounted in canada balsam 
for microscopic examination. 


side, with some very 
minute additional 
ones on basal half 
(both above and 
below). Buccal 
opening blackish, 
with a row of strong 
spiny bristles each 
side, the uppermost 
one being the strong- 
est; a number of 
very small bristly 
hairs on under side 
of epistome. Palpi 
blackish grey, cy- 
lindrical, narrowed 
towards base, with 
very short irregular 
bristles. 

Proboscis consist- 
ing of a sub-conical 
thickened fleshy 
basal portion, fol- 
lowed by a subcy- 
lindrical, chitinous 
black portion (both 
portions about equal 
in length); at the 


end of the latter a softer tip bearing 7 black teeth in accordance 


with the generic description. 


Fic. 2.—Proboscis of Pristivhynchomyia lineata; profile view of dried specimen. 


—————— eee er lee 


IgI0.] E. Brunetti: Blood-sucking Muscide. 93 


Thorax. Varies from dark ash-grey (sometimes with a bluish 
tinge), to distinctly yellowish grey. Normally, the markings 
of the dorsum appear as four stripes, the inner pair barely inter- 
rupted at the suture but slightly widened behind it ; the outer 
pair less regular, distinctly interrupted at the suture ; in front of 
which the direction is changed, pointing to, and nearly reaching, 
the shoulders ; and behind which, the stripes, seen from behind, 
appear widened near the bases of the wings; thence narrowing 
and disappearing before attaining the posterior border of the 
dorsum. Scutellum and sides of thorax concolorous. 

Chetotaxy as in Philematomyia insignis, Aust. The whole 
surface of the dorsum is covered with minute bristles. Of the dorso- 
central rows of bristles, the intermediate smaller ones are larger in 
size than in that species, but 5 or 6 macrocheete are usually evident 
behind the suture and 2 (with a lesser, intermediate but by no 
means minute one) in front of it. 

Abdomen. Ist segment blackish, remainder yellowish grey ; 
2nd and 3rd narrowly black at the base and on posterior border. 
A rather narrow dorsal black line extends over both segments; 
4th segment with two small elongated blackish dorsal spots. 

The whole abdomen with short stiff hairs; no macrochete, 
but a row of rather stronger bristly hair near posterior border of 
each segment ; these being somewhat stronger and longer towards 
the sides and on the 4th segment. Belly concolorous, with short, stiff 
hairs. Ovipositor capable of complete withdrawal; when _pro- 
truded, a conspicuous, yellowish grey, nearly bare, cylindrical organ 
with a strong, black, hard apical knob. 

Legs. Black, with a little greyish reflection in certain lights ; 
minutely spinose. Fore femora with several longitudinal rows of 
strong bristles on outer side, a stronger spiny row on under side ; 
middle femora with 2 or 3 short, stiff bristles on hinder side near 
tip ; hind femora with 2 or 3 similar bristles below, near tip. 

Anterior tibiae with 2 distinct spines at tip, shorter ones on 
hind pair ; middle tibiz with 3 or 4 bristles on hinder side ; hind 
tibiae with apparently an irregular row of similar bristles on outer 
side. 

Wings. Pale grey, veins yellow on fore border and _ base, 
blackish elsewhere. Costa with a few short black bristles at base. 
Tegule yellowish grey, upper scale small, quadrangular. Halteres 
very small, yellow. 

Described from a good series of ? @ in the Indian Museum 
taken in Calcutta inearly December,1907. I have seen no specimen 
that I can assume to be the @ of this species. 

It may be remarked that Dr. Annandale has repeatedly ob- 
served this fly distended with blood, and actually sucking open 
sores on cattle. ‘Though the proboscis itself is not sufficiently 
stiff to pierce the skin the toothed tip would easily enlarge a small 
hole, otherwise made. 


—- ~ - ~ 


: nd B =i 
; wy ‘ 7, : Dini al 
es ad a, 7 7 = ~ ; 
J n a a* 
. au 
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ee ral . — ’ 
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} wr . * y —_ : ‘y ha 
aa _ + 7 
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ms * - = Se 
14% a . - 
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pa ae, s ue ‘ “4 : - > 
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. . 
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Mw some sO ad Pag “4 . e 
Pew gett. Secs : 
ae fag : ; re ae | ee, . 
Aan ane ee Aa 7 my * aa 
PP es CERES Ute ae 
ree ao Ns ay oor) ata 
a wi See At r ae 
Pine e Pen Se Livi he Aap cages 
; . “h - ie a at ee hand 
"> ae] ' min pat Ec: 
ry i" 4 ‘ee <5 Ale ‘= sf] , “ad, bs 
we b r Peay a he 
Dm J ines m4 Pah " 
_ rh ahve @ Vere i ' 
] F Yi eal J Ds" us 7 ' fi 
4 4 is a +» ? 
. uk aA >, 4 s 
® L a : a _ 


uit? 


os) 


EXPLANATION OF PLATE VII. 


1.—Stomoxys calcitrans, 1,., head o , front view. 


2.— vd., head @ , profile. 

3.— wd., head ¢ , front view. 

4.— Ves arista. 

5.— 1d., thoracic bristles, viewed from above. 
6.— id., rd., lateral view. 

— ta.., wing. 

8—I11.— 4a. abdomen (8, typical). 

12.—S. indica, Picard. 

13.—S. triangularis, Brun., sp. nov. r 


14.—S. oblongopunctata, Brun., sp. nov. 
15.—Bbdellolarynx sanguinolentus, Aust., head &. 


16,.— id., heads ¢ 
Ge td., arista. 
18.— id., wing. 


19.--- id., abdomen. 


Rec. Ind. Mus.,Vol.IV, 1910. Plate VI. 


15 x16. 18x8. 


“Out ) skeet 


a deli a Wee ppexeit eeagsch er ee ti. 
eer ihs hens Pua atte Gey! soeadaey ay 


Fe ae Oha2 LUN Ate on] > BPS) eee Bhi - 
Pais t ev, 


pated 9. Je > 
So )s eee io : 
> ‘ 
ata aS 
. ae rf Lt ean at '. 7 
- a 
: , AD a aha - 
reaviuen Fo) ae uh a ; 
2 i 7, - i : * wns brie - 5 aa! 3 242 
Or 2 ree > Ee ‘Sao. © rt a ER ; ; 
awed “tatebey ity hela Peek ei Pesatss es Oe LL ee oa : 
$e var 7 = i 
; na so priors : ‘to > ofite 4 
»* ong Pry) Sozst)if Pe, ba . 1d a 
e a tatohe “ .* 
i valy ene pos ek: *s a . ri 
‘ 1 7 2 ” 
a A taahicut) sant : ' tee 


bette ate ve 2 a 7 
: a, a ar 
ale >t ape Lae bs VAP STNG TVS A 4 rr 


sy 
4 : is 
; 
4 
: 
e 
» 
- 
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- 
7 =e P 
al 
¢ ? e 
a 7 4 
+ 
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’ 
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. 
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EXPLANATION OF PLATE VIII. 


1.—Lyperosia minuta, Bezzi, head ~ , front view. 


2.— Las, head o@, profile. 
3.— td., head 9 , front view. 
= 4d. arista. 
5.— tae thoracic bristles, dorsal view. 
6.— Mi biee id., side view. 
’ ae td., wing. 
8.—-L. exigua, Meij., head a. 
9 vd., head @. 
10.— id., abdomen. 
II.— td., hind leg o. 
12.—Philematomyia insignis, Austen, head, profile. 
13.— vd., proboscis. 
14.— id., head o, profile (small var- 
iety). 
15.— td., head @ , front view. 
16.— 1d., thoracic bristles. 
17.— 1a., wing. 


18.—Pyistirhynchomyia lineata, Brun., sp. nov., head ¢ , front view. 
19.— id., head, profile. 


4 


Rec. Ind. Mus., Vol.IV, 1910. | 


‘ 
7 


‘ 


Vo A NEW: ARRANGEMENT -O) DEE 
EN DIAN <A N OP HEL INA. 


by S. P. James, M.D., Major, Indian Medical Service ; 
Secretary to the Committee for the Study of Malaria in India. 


The following tabular statement shows (1) the names of the 
groups or genera in which the anopheline mosquitoes of India are 
at present arranged by Mr. Theobald, and (2) the characters by 
which Mr. Theobald identifies the genera and distinguishes them 
from one another. 


Abdominal Thoracic | Form of wing | Form of head 
Genus. ornamentation. | ornamentation. | scales. scales. 
| — 
Anopheles, Hair-like Hair like Large and Upright fork- 
Meigen. curved scales. curved scales. | lanceolate. ed, but no 
| flat scales. 
Myzomyia, se | re Mostly small. as 
Blanchard. | long and nar- 
| Tow or slightly 
| lanceolate. | 
Stethomyia, rr a3 | Lanceolate. | Median area 
Theobald. | | of head with 
some flat 
| scales. 
Pyretophorus, | Hairy. | Narrow curved Small and lan-| Not stated. 
Blanchard | scales. | ceolate. 
Myzorhynchus, Apical ventral) Hair-like | Dense, large, | Not stated. 
Blanchard, tuft of scales. curved scales tanceolate. | 
Nyssorhynchus, Wateraltuftsand Narrow curved) Not stated. | Not stated. 
Blanchard. small dorsal) or — spindle- 
patches of flat shaped scales. | 
scales. 
Cellia, Nearly complete-- With scales. Not stated. Not stated. 
Theobald, ly scaled with 
long irregular 
scales and with | 
lateral tufts. 
Neocellia, Similar to Cellia With scales. Not stated. Not stated. 
Theobald. but no lateral 
tufts. 
Aldrichia, Completely With scales. Not stated. | Not stated. 
Theobald. scaled with | 


large flat scales 
as in Culex, 


It has to be noted also that Mr. Theobald has stated that the 
common Indian species vosst cannot be placed in any of these 
genera and that a genus Pseudomyzomyia would be created for its 


96 Records of the Indian Museum. [Vor IV, 


reception. So far as I am aware the characters of this genus have 
not yet been published. 

A careful examination of the above table will convince the 
reader that it would be very difficult, if not impossible, correctly 
to arrange the Indian species generically by its aid. Those who 
have worked at the subject will be aware also that the defining 
descriptions can be criticised as being, (1) incorrect (e.g., the 
description of the abdominal ornamentation in the genus 
Nyssorhynchus; there are no lateral scale tufts in mosquitoes of 
this genus), or (2) inadequate (e.g., the descriptions of the genera 
Cellia and Neocellia), or (3) confusing and indefinite (¢.g., the des- 
criptions of the forms of wing scales). The difficulties caused 
by these defects have led to much criticism of the system of classi- 
fying the Anopheline on a basis of scale and hair covering, but it 
is probable that the fault lies not so much with the system as with 
the inadequate and confusing nature of the generic definitions. 
In this revision I shall try to avoid similar defects, but limits of 
space prevent me from giving more than a very brief summary of 
my work ; and for the same reason I am prevented from explain- 
ing in full why I differ from Mr. Theobald in regard to points of 
detail. 

A consideration of the best way in which to arrange the 
Indian anophelines according to scale and hair ornamentation is 
simplified if one starts by separating all the species in two great 
groups, the first containing those without scales on the abdomen, 
and the second containing those with scales on some part or the 
whole of that region of the body. According to the published 
descriptions of species the following come in the first of these 
groups. 

( atthent, James. 
| immaculatus, Theobald. 
| culictformis, James and Liston. 
lindesayt, Giles. 
gigas, Giles. 
culictfacies, Giles. 
Species without scales } listonz, Liston. 
of any kind on the } leptomeres, ‘Theobald. 
abdomen. jeyportensts, James. 
turkhudt, Liston. 
punctulata, James and Liston. 
| elegans, James. 
| nigrtfasciatus, Theobald. 
| nurset, Theobald. 


a= 


These 14 species are at present placed by Mr. Theobald in 
the following groups or genera :— 


In the genus Anopheles: aitkent, immaculatus, lindesayi 
and gigas. 
In the genus Stethomyia : culiciformis. 


IgI0.] Ss. P. James: The Indian Anopheline. 97 


In the genus Myzomyta: culicifacies, listont, leptomeres 
and turkhudt. 

In the genus Pyretophorus : jeyporrensts, punctulata, 
elegans, nigrifasciatus and nurset. 


I have examined many specimens of nearly all these species 
and the following remarks contain the chief conclusions at which 
I have arrived. (1) I have carefully examined specimens of 
maculipennts, Meigen, and bifurcatus, Linneeus, and taking them 
as the types of Mr. Theobald’s genus Amoheles, I am unable to 
place in this genus any Indian species at present known to me. 
(2) I find that the distinguishing character of the genus Stetho- 
myta, namely, the presence of a few “‘ flat scales”’ on the head, is 
not confined to the species placed by Mr. Theobald in that genus. 
Mr. Theobald found that the original character upon which he 
founded this genus (namely, the character ‘‘ mammilated prothora- 
cic lobes ’’) was not distinctive, and now that the second character 
has been found to be not distinctive the genus should be sunk. 
Probably all the species now assigned to it would come in my new 
genus Neostethopheles. (3) The use of the shape of wing scales 
as a means of distinguishing between the genera of this first large 
group of anophelines appears to me to be open to the objection 
that it must always be doubtful whether, for example, the wing 
scales are ‘“‘ large and lanceolate’’ or whether they are ‘‘ mostly 
small, long and narrow, or slightly lanceolate.’’ I find, however, 
that this character is the only one by which the genera Anopheles 
and Myzomyta can be separated, and that unless it is agreed to 
sink the latter genus under the former we must accept it. The 
wing scales of Anopheles maculipennis are distinctly longer and 
broader than those of Myzomyia culicifacies and Myzomyia listont, 
but they are only very slightly longer and broader than those of 
Myzomyia turkhudi. ‘The wing scales of Anopheles bifurcatus are 
not so long nor so broad as those of Myzomyia turkhudi. If the 
character is regarded as of general application to all the genera 
endless confusion ensues. As I do not place any Indian species 
in the genus Anopheles and asI neglect the character of the wing 
scales for the grouping of the Indian species the subject need not 
be further considered here, but it is very desirable to find a better 
character for separating the genera Anopheles and Myzomyia. (4) 
I believe that the Indian anophelines without scales on the abdo- 
men can be arranged in groups based on characters much more 
distinctive and easily recognized than those which Mr. Theobald 
has chosen, 

For the arrangement of the species without scales on the 
abdomen I retain two of Mr. Theobald’s groups, namely, Myzomyia 
and Pyretophorus, and create two new groups called respectively 
Neostethopheles and Patagiamyva. 

The following definitions include the characters by which the 
four groups in which I arrange the 14 species without scales on the 
abdomen may be identified and distinguished from one another. 


98 Records of the Indian Museum. [Vou. IV, 


Genus 1. NEOSTETHOPHELES, nov. gen. Abdomen with 
hairs but without scales of any kind. Thorax with hairs and as a 
rule without scales of any kind, but in one or two species a few 
long, exceedingly narrow, false scales! may be present on the anterior 
promontory only.  Prothoracic lobes with hairs but without scales. 
Upright forked scales of the head very narrow in their whole length 
(vod shaped). 


Type of the genus: aitkent, James. The distinguishing char- 
acters of the genus are shown in Plate ix, fig. I. 
I place the following species in this genus :— 


attkent, James. 
emmaculatus, Theobald. 
culictformis, James and Liston. 


Genus 2. MYZOMYTIA, Blanchard. Abdomen with hairs but 
without scales of any kind. Thorax with the dorsum clothed with 
long, very narrow, sharp-pointed, curved scales more numerous ante- 
riorly and forming on the anterior promontory a bunch projecting 
over the neck. Nearly all the scales are false scales. Prothoracic 
lobes with hatrs but without a tuft of scales. Upright forked scales 
of the head of the usual anopheline type, that ts broadly expanding 
from the base to the apex. 


Type of the genus: cwlicifacies, Giles. The distinguishing 
characters of the genus are shown in Plate ix, fig. 2. 
I place the following species in the genus :— 


culicifacies, Giles. 

listont, Liston (= christophersi, Theobald). 
culicifacies, variety punjabensis. 

turkhudt, Liston. 

leptomeres, Theobald.?* 


Genus 3. PATAGIAMYIA, nov. gen. Abdomen with hairs 
but without scales of any kind. Thorax with the dorsum clothed 
with long, narrow, curved, sharp-pointed scales which form on the 
anterior promontory a thick bunch projecting over the neck. Some of 
the scales are false scales and some are true scales. Prothoracic lobes 
with a conspicuous tuft of rather broad true scales projecting anteri- 
orly. Upright forked cephalic scales of the usual broadly expanding 
type. 

Type of the genus: gigas, Giles. At present the only other 


1 In this revision false scales are defined as those without clearly marked striae. 
There may be an appearance of linear marking, but separate striae cannot be 
counted. True scales always have definite bars or striae passing from the base to 
the apex of the scale. These striae often project beyond the apex, and when 
the scale is examined under the microscope they can be easily counted. 

2 When a species 1s marked with an asterisk it means that I have not examined 
its seal ornamentation in detail and that it is placed in the genus only provi- 
sionally, 


detente ene 


ees Wd 


oe ss eT 


_—— 


1910. ] S. P. JAMES: The Indian Anopheline. 99 


Indian species in the genus is lindesayi, Giles. The distinguish- 
ing generic characters are shown in Plate ix, fig. 3. 


Genus 4. PYRETOPHORUS, Blanchard. Abdomen wtth hairs 
but without scales of any kind. Thorax and scutellum with moder- 
ately broad, rather short, true scales some of which are blunt ended 
(ob-lanceolate). Prothoracic lobes with hairs and sometimes with two 
or three scales scattered irregularly, but always without a bunch or 
tuft of scales. Upright forked cephalic scales of the usual broadly 
expanding type. 

Type example of the genus: palestinensis, Theobald.' The 
generic characters are shown in Plate ix, fig. 4. 

I place the following species in the genus :— 


elegans, James. 

jey portensts, James. 
nigrifasciaius, Theobald.* 
nurset, Theobald.* 

punctulata, James and Liston.* 


I now take up the arrangement of the species which have 
scales on some part or the whole of the abdomen. Omitting 
certain varieties and doubtful forms they number 1g in all. At 
present 17 of these species are usually classified in Mr. Theobald’s 
genera as follows :— 


In the genus Nyssorhynchus: maculatus, Theobald. 
fuliginosus, Giles. 
jamest, Theobald. 
theobaldi, Giles. 
macultpalpis, James and Liston. 
karwart, James. 
stephensi, Liston. 
wtllmort, James. 

In the genus Myzorhynchus : barbirostris, Van der Wulp. 
sinensis, Wiedmann. 
nigerrimus, James and Liston, 
nigerrimus, Giles. 

In the genus Cellia: pulcherrima, Theobald. 


1 I name palestinensis as my type species because Mr. Theobald on page 71 
of vol. iii of his monograph has figured the thoracic scales of this species The 
figure shows quite broad true scales some of which are blunt ended, and on page 
74 Mr. Theobald says that these scales ‘‘ of quite a different structure to the 
narrow hair-like scales on turkhudi, clearly relegate it to the genus Pyretophorus.’’ 
According to my view a simple definition of the generic characters of Pyvetophorus 
would be ‘‘ with Nyssorhynchus-like thoracic scales but with no scales on the 
abdomen,’’ Costalis, whichI have not examined, is sometimes named as the type 
species of the genus, but if its thoracic scales do not correspond to this simple 
definition I should remove it. 


100 Records of the Indian Museum. [VoL. IV 


In the genus Neocellia: indica, Theobald. 
intermedia, Rothwell. 
dudgeontt, Theobald. 


In the genus Aldrichia: error, Theobald. 

Rossi, Giles, for which Mr. Theobald proposed a genus called 
Pseudomyzomyta, and halli, James, which is a new species not yet 
assigned to a genus, are not included in the list; they will be 
considered separately in this account. 

For the arrangement of the species in this second large group— 
the group in which scales are present on the abdomen—I retain 
the names of the groups Nyssorhvnchus, Myzorhynchus, Cellia, and 
Neocellia, but I alter Mr. Theobald’s definitions of these groups so 
as to make them represent clearly the scale characters of the 
groups. In the second place I change the position of one or two 
species that have been placed in wrong groups, and in the third 
place I create two new groups, one for vosst, the other for hall. 
I am unable to say anything about the genus Aldvichia, as I 
have not seen the species which represents it. 

Dealing first with the group Nyssorhynchus I agree with 
Mr. Theobald in regarding the Indian species maculatus as a suitable 
type of a group of anophelines characterized by the presence of 
scales on only the last one, two, or three segments of the abdomen, 
and I find that the scale structure of this species agrees in all 
important respects with that of fuliginosus, jamesi, theobaldt, 
macultpalpts and karwart. My definition of the group characters is 
as follows : 


Genus 5. NYSSORHYNCHUS, Blanchard. Abdomen with the 
first five or stx segments ornamented with hairs only. The last three 
or two segments and the genital processes carry in addition a number 
of rather long, blunt-ended true scales on both the dorsal and ventral 
surfaces. On the 8th segment and the genital processes the scales may 
be arranged in patches or may cover the surface more or less evenly ; 
but they are never aggregated together to form tufts of any kind. 
Thorax with the dorsum covered with quite broad true scales usually 
avranged in more or less parallel lines. On the anterior promontory 
the scales ave wong and sharp-potinted and form a small bunch projec- 
ting over the neck on each side of the middle line; on the mid region 
and posteriorly they are broader and some are blunt-ended. The 
scutellum carries a number of similar scales. Pyrothoracic lobes witth- 
out a tuft of scales. Head with the usual kind of upright forked 
scales. 

Type of the genus, maculatus, Theobald. ‘The generic charac- 
ters are shown in Plate x, jigs. 1 to 6. 

I place the following species in this group :— 


maculatus, ‘Theobald. 
fuliginosus, Giles. 
jamest, Theobald. 
theobaldi, Giles 


1910. |] S. P. James: The Indian Anopheline. IOI 


karwan, James. 
maculipalpis, James and Liston. 


I have next to consider the species vossz, which in its scale 
ornamentation exhibits some of the characters of the group Myzo- 
myta and some of the group Nyssorhynchus. An examination of 
a large number of specimens of this mosquito has shown that 
although minor differences in the degree and character of the scale 
ornamentation are common, the chief features, as figured in plate x, 
igs. 7 to II, can always be made out on unrubbed specimens. I 
describe them thus :— 


Genus 6. NYSSOMYZOMYIA, nov. gen. Abdomen with the 
first seven or six segments ornamented with hatrs only. The eighth 
segment (sometimes also the seventh) and the genital processes carry in 
addition a number of scales similar in character and arrangement to 
those of the group Nyssorhynchus. Thorax with the dorsum clothed 
with hairs and narrow, curved, sharp-pointed scales of various 
lengths and quite similar to those of the group Myzomyia. In addi- 
tion there are on each side of the anterior third of the dorsum a few 
broader blunt-ended Nyssorhynchus-like scales. Prothoracic lobes 
without a tuft of scales. Head with the usual type of upright forked 
scales. 

The species is, therefore, representative of a group intermediate 
between Myzomyia and Nyssorhynchus, and is nearer to the latter 
than to the former group. The term Nyssomyzomyia suitably 
indicates that it possesses the characters of both groups and I 
therefore apply this name to the genus instead of the name Pseudo- 
myzomyia which Mr. Theobald proposed, but the characters of 
which he has not yet described. ‘The characters of the genus 
Nyssomyzomyia are shown in Plate x, figs. 7 to II. 


I take up next the group NEOCELLIA, Theobald, several 
members of which have usually been wrongly placed in the genus 
Nyssorhynchus. The characters of this group are very different 
from those of the group Cellia and the name Neocellia is therefore 
quite misleading. Neonyssorhynchus would have been a more 
suitable name, but the resemblance even to that group is not close. 

The type species of the group is the Indian species indica, 
Theobald. I define the group characters thus : 


Genus 7. NEOCELLIA, Theobald. Abdomen with the dorsum 
of each segment clothed irregularly with hairs and long, rather broad, 
blunt-ended scales. The scales are not aggregated together to form 
tufts of any kind, but they are more numerous and thickly set on the 
last two segments than on the others. On the ventral surface the first 
five segments are devotd of scales, but on this surface of the 6th, 7th, 
and 8th segments they are present in considerable numbers, being 
disposed irregularly but attached chiefly on each side of the mid line 
and not forming tufts of any kind. Thorax clothed with broad true 


102 Records of the Indian Museum. [Vou. IV, 


scales. Prothoracic lobes with or without a few scales irregularly 
disposed, but always without a definite bunch or tuft of scales. Head 
with the usual type of upright forked scales. 

The characters of this genus are shown in Plate xi. I place 
the following species in it— 


indica, Theobald. 

stephensi, Liston. 

willmort, James. 

intermedia, Theobald.* (This perhaps = stephenst.) 
dudgeonu, Theobald.* (This perhaps = willmort.) 


Lastly, I have to consider the arrangement of the species in 
which some of the abdominal scales are disposed so as to form 
distinct bunches or tufts projecting from the dorso-lateral or from 
the ventral surface of certain segments. These tufts form con- 
spicuous objects readily seen with a hand-lens, and their presence 
supplies an easy means of separating the following groups from 
any of those described above. ‘The Indian species provided with 
certain of these abdominal tufts of scales are (1) pulcherrima, 
Theobald ; (2) senensts, Wiedmann (= vanus, Walker); (3) nigerri- 
mus, James and Liston (which probably = sinensis, Wiedmann, and 
vanus, Walker); (4) migerrimus, Giles; (5) barbivostris, Van der 
Wulp; (6) hallx, James; and they must be arranged in three quite 
distinct groups, the names of which are (1) Cellia, Theobald ; (2) 
Myzorhynchus, Blanchard; and (3) Christophersia, nov. gen. 


The following are my definitions of these groups :— 


Genus 8. CELLIA, Theobald. Abdomen with the dorsum of 
each segment clothed with very large and broad orbiculay and square- 
ended scales which stand out somewhat from the surface and overlap 
one another. In addition at the postero-lateral corner of each segment 
from the ist to the 7th is inserted a bunch of large scales jorming a 
tuft which projects laterally. Six tufts on each side are plainly 


vistble with a weak lens. The ventral surface of each segment ts 


clothed more or less evenly with very broad scales like those of the 
dorsum, but the clothing ts not so thick as on that surface. The 
postero-lateral tufts oj the dorsal surface ave visible on each side, but 
there are no ventral tufts of scales projecting downwards. Thorax 
clothed with very broad scales. Prothoracic lobes with a few scales 
projecting forwards. 

Type species of the genus, pulcherrima, ‘Theobald. The 
generic characters are shown in Plate xi. 


Genus 9. MYZORHYNCHUS, Blanchard. Abdomen with the 
dorsal surface clothed with hairs only. On the ventral surface from 
the apex of the seventh segment in the middle line a prominent bunch 
oy tuft of rather long, black, true scales projects downwards. On the 
ventral surface of the 6th, 5th, 4th and 3rd segments a few small white 
scales may be present; if so, they are arranged somewhat irregularly 


1gI0. | S. P. JAMES: The Indian Anopheline. 103 


and do not form tufts. Thorax with the dorsum clothed with hairs 
and narrow, sharp-pointed scales like those in the genus MyzoMyta. 
Prothoracic lobes with a dense tuft of broad true scales projecting 
anteriorly. Head with short very broadly expanding upright forked 
scales. 

Type species of the genus, barbivostris, Van der Wulp. The 
generic characters are shown in Plate xii. The other species in 
the genus are sinensis, Wiedmann, nigerrimus, James and Liston, 
and nigerrimus, Giles. 


Genus 10. CHRISTOPHERSIA, nov. gen. Abdomen with the 
dorsum of each segment thickly clothed with hairs and lanceolate and 
hlunt-ended scales which are not aggregated together to form tufts of 
any kind. The ventral surface of each segment 1s devord of scales 
except that from the apices of six segments in the mid line prominent 
tufts of long, blunt-ended scales project directly downwards. These 
tufts resemble the single abdominal tuft present tn the genus Myzor- 
uyncHUS. Thorax clothed with rather narrow lanceolate and blunt- 
ended true scales. Prothoracic lobes with a prominent tuft of true 
scales. Head with the usual kind of upright forked scales. 


Type of the genus: /alli, James. The generic characters are 
shown in Plate xii. 


It only remains now to summarize in tabular form the essen- 
tial differences between the groups I have defined, to indicate 
briefly how to place an anopheline in its correct genus, and to show 
in the form of a table the changes made in the generic position of 
the different Indian species. 

How to ascertain the generic position of an anopheline. 
(1) Using a microscope with a two-thirds inch objective and a 
high power eyepiece (No. 8 or No. 12), examine the dorsal, lateral, 
and ventral surfaces of the abdomen of the mosquito. If scales 
are not immediately seen search carefully the last segment and 
the genital processes. Upon the decision whether the abdomen 
carries scales or not depends the accuracy of the subsequent 
determination. The decision is always very easily made, but 
especial care should be taken to examine the lateral and ventral 
aspects of the abdomen as well as the dorsal aspect. If no scales 
are present, the mosquito belongs to one of the first four genera. 
(2) In that case next examine the head. It will be seen at once 
whether the upright forked scales are of the usual broadly expand- 
ing type (all the spotted-winged anophelines have scales of this type) 
or whether the upright forked scales are very narrow in their whole 
length (linear or rod-shaped). If they are rod-shaped the mosquito 
belongs to the genus Neostethopheles. (3) If the upright forked 
scales are of the usual broadly expanding type, next examine the 
prothoracic lobes from the dorsal aspect. ‘To see one of them clear- 
ly slant the pin so that the mosquito is turned more or less on its 
side. A very little practice will overcome any initial difficulty 


[Vou. IV 


‘ 


Records of the Indian Museum. 


104 


| “sayeos 


| Jz0y4s ‘IOpeoigq sUIOS Y}IM pue "SHYIUAYAOSSANT SUAS 944 
‘ad4} Surpuedxa ‘saqeos | ‘viAmozd py snuad ay} Ul se sayeos | UI se SaTeos ani} SutAz1e9 sjuem “uas 


A]peoiq jensn ay} JO | Jo yn e IOUT | payarod-dieys ‘Morren ‘Su0] WHIM | -89S z ISP] OY} IO YseT oN} YUMA | “ow ‘VIAWOZAWOSSAN 


-9133e JOU 91e YOIYM sapeos ons} 


‘S}JN} WIOF 0} pozes | 


“SLANL ATVOS 
TVNINOGEV LAOH LIM 


‘ad4} Suipued *sa]BOsS | Azied sjusuises zc oO £ 4sel oy} *preyo 
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“xo ATpvorq [ensn aq} JO yo yn} e ynoYA\ ‘SaTeos ans} peoiq oymb yA ‘SoTBOS JNOYFAL Inq Suey UAL | “Wel ‘saXOHdOLANAd | = 
| ‘soles | “AT = 
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| G 
| * A[IOTIO}J Ue | > 
snoijounim atom ynq ‘a0ejins mS 
‘ad 4} Surpued ‘SoTBOS | [BSIOP BJOYM 9} WO SaTeIS paAino ee 
-xo ATpBOIq [eNsN 94} JO | Jo yn} & YONA peiutod-dreys ‘morse “SuOT ITM, ‘Sa[VoS JNOYFIA 3Nq sIrey GUA | “‘preyouelg ‘VIAWOZAIL | z 
(‘snua sty} wor; | & 
| _satoeds & apnoxa you saop 410} 5} 
*xode 94} -uomoid JOM9}Ue IY} WO saTeos = 
spieMo} Surpuedxs AT | Moen AIA =‘ poyutod-dieys 2 
~peoig you ad4q padeys | ‘sayeos ynoyya ‘Buoy, Moy e& yo aouasoid euL) Wai, 
“por oystiajoeIeYyO BIO | INQ sey UAL ‘soqeos Noy 4nq sIrey WIM ‘SoTBOS JNOYFIA jnq SIV YIM | ou ‘sq IHHAIOHLALSONN 
é : es ee RENEE Poe REET ele WAS E00 pa nba a me 
‘peo, oy} 


| ‘saqo] o1oe10yyoIg ‘xeIoy | ‘TaTMOpqy | ‘snes 


| | 
| 
Se Oe ew se (LS? BA 


‘SABJIVADYD IUAaUaH fo a7qv T 


JO safeos poyxs10y yYSs1IdyQ 


105 


ne. 


The Indian Anophel 


S. P. JAMES 


IQIO. | 


‘ad4} Surpuedxe 
jensn eq} JO 


Aypeo1iq 


‘ad A} Surpued 


-xo A]peoiq jensn 9} JO 


‘addy Surpuedxe 
Ajpeoiq yensn 24} JO 


‘addy Surpuedxe 


A[peoiq 


jensn 


2q7 JO 


‘Savas JO yn} 
quourmoid ve TM 


*sa]BOS anI} jo 


| 37Ny asuap B TIA 


‘sprea 
-10j Surjoafoid 
soyeos Moy B IAA 


‘so]eos 
JO }JN} & JNONWA 


| 
i} 


| 
| 


‘savas 
ani} peorq = Ajaze1opour YA 


“DiAUL 
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sajeos pojurod-dieys ‘suo, WWM 


‘soTBoS ona} peorq AISA YIM 


‘saTeos ona} peoiq YM 


aU] Pru oy} m0Iy 
SpivauMop Apoemp surjoefoid 
saTeos and} JO $}jn} JueuTMOId xts 


| UJIAA O0RJINS [eIJMOA sy, ‘s}yNy 
| UIIO} JOU Op YOIYA\ sayeos paps 


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pUotudses [v9 JO 9dRJINS [eSIOp 9, | 


| ‘OUI] PIU 94} UI UoUIsaS T}MWIAVS 


ay} jo xode oy} woIy spieM 
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‘uas 


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TYNINOG 
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106 Records of the Indian Museum. [Vor ty 5 


Table of Species. 


Nene err nn nnn nn nee enn 
Genus according to | CG aineee 
Mr. Theobald’s | Bete ECON See 
the arrangement 


Specific name, 
described here. 


| Monograph of the 


Culicidae. 
r= eae ee a= ———— = se 
aitkeni, James. | Anopheles. | Neostethopheles. 
immaculatus, Theobald. _ Anopheles. | Neostethopheles. 
culiciformis, James and Liston, | Stethomyza. Neostethopheles. 
gigas, Giles. | Anopheles. Patagiamyia. 
lindesayt, Giles. Anopheles. | Patagiamyia. 
lindesayi, variety maculata, Theo- | Anopheles. | Patagiamyia. 
bald. | | 

culicifacies, Giles. | Myzomyia. | Myzomyia. 
listoni, Liston. | Myzomyza. | Myzomyia. 
culicifacies, variety punjabensis. eae? | Myzomyia. 
jeptomeres, Theobald. | Myzomyia. | Myzomyia.* 
tuvkhudi, Tiston. Myzomyia. | Myzomyia. 
elegans, James. _Myzomyta, 1903. | 

| Pyvetophorus, 1907. | Pyretophorus. 
jeyporiensis, James. | Pyretophorus. | Pyretophorus. 
punctulata, James and Liston. | are Pyretophorus.* 
nigrifasciatus, Theobald. Pyvetophorus. | Pyretophorus.* 
nursei, Theobald. | Pyretophorus. | Pyretophorus.* 
maculatus, Theobald. Nyssorhynchus. | Nyssorhynchus. 
fuliginosus, Giles. Nyssorhynchus. | Nyssorhynchus. 
fuliginosus, variety nagpori, J. | eves | Neesornncnaie 

and IL. | 

fuliginosus, variety adie, James, | a | Nyssorvhynchus. 
qamest, Theobald. | Nyssorhynchus. Nvssorhynchas. 
theobaldt, Giles. Nyssorhynchus. Nyssorhynchus. 
maculipaipis, James and Liston, | ae a | Nyssorhynchus. 


maculipalpis, variety indiensis, | Nyssorhynchus. 
Theobald. 

kavwari, James. 

yosst, Giles. 


| Nyssorhynchus. 
Myzomytia, 1903. 

| Pseudomyzomyia, 1907. 
indica, Theobald. | Neocellia. 

intermedia, Rothwell. lSnteoceiina: 


Nyssorhynchus.* 


| Nyssorhynchus. 


Nyssomyzomyta. 
Neocellia. 
Neocellia.* 


stephensi, Liston. | Nyssovhynchus. | Neocellia. 
willmori, James. | Nyssorhynchus. | Neocellia. 
dudgeonti, Theobald. | Neocellia. | Neocellia.* 
pulchervima, Theobald. | Cellia. | Cellia. 
barbivostris, Van der Walp. Myzorhynchus. Myzorhynchus. 
sinensis, Wiedmann. | Myzorhynchus. | Myzorhynchus. 
nigervimus, James and Liston. | Myzorhynchus. Myzorhynchus. 
nigervimus, Giles. | Myzorhynchus. | Myzorhynchus. 
halli, James. | ee | Christophersia, 
pet 4 


evvory, Theobald. | Aldvichia. 
} 


er ee —— 


— 


1910. | S. P. JAmes : The Indian Anopheline. 107 


that may be experienced in obtaining a good view of one of these 
bodies. It will at once be seen whether or not the lobe carries a 
cocade of scales projecting anteriorly. If the lobe carries such a 
cocade the mosquito belongs to the genus Patagiamyia. (4) Ifa 
tuft of scales is not present on the lobe next examine the dorsal 
surface of the thorax. If the scales are long, sharp-pointed, and 
very narrow (that is, if it is rather difficult at a first glance to say 
whether they are scales or hairs) the mosquito belongs to the genus 
Myzomyia. Tf the scales are short and moderately broad (that is, 
if it can at once be seen that they are true scales) the mosquito 
belongs to the genus Pyretophorus. 

In the second case we shall have detected the presence of 
scales as well as hairs on one or more of the abdominal segments. 
In that case proceed as follows: (1) The first point to decide is 
whether or not some of the scales are aggregated to form tufts. 
The appearance of these tufts is shown in Plate XII; they are 
very definite objects and the mistake must not be made of regard- 
ing as tufts the irregularly disposed scales that in some species 
overhang the sides of the terminal abdominal segments. The 
single tuft projecting downwards from the mid line of the ventral 
surface of the 7th abdominal segment in anophelines of the genus 
Myzorhynchus is the least easy to detect. In specimens of that 
genus we usually see no scales during the examination of the 
dorsal surface, and even when the mosquito has been turned 
upside down for the examination of the ventral surface, the 
ventral tuft may not be recognized because its very dark scales 
are viewed against the equally dark background of the abdomen. 
it is not until the mosquito has been turned on its side that the 
ventral tuft is easily seen. That is why it is so important not to 
neglect the examination of the last abdominal segments from 
every direction of view. If it is found that the abdomen carries 
only this single ventral tuft of scales, we can say at once that the 
mosquito belongs to the genus Myzorhynchus. The determination 
should be confirmed by finding that the thorax carries long, sharp- 
pointed, myzomyia-like scales, and that the prothoracic lobes 
carry a cocade of scales. (2) If on examining the dorsal surface 
we see at once that it is clothed with very broad scales and that 
tufts of scales project laterally from the sides of each segment, 
the mosquito belongs to the genus Cel/ia. (3) If on examining the 
dorsal surface we see that every segment carries scales and that 
there are no lateral tufts, but that when the mosquito is turned on its 
side a number of very prominent ventral tufts come into view, the 
mosquito belongs to the genus Christophersia. 

If our examination has shown that none of the abdominal 
scales are aggregated to form tufts, it has to be decided whether 
all the segments are provided with scales on their dorsal surface 
or whether only the terminal segments carry scales. In the first 
case the mosquito belongs to the genus Neocellia. In the second 
case examine the thorax; if the scales are short and broad the 
mosquito belongs to the genus Nyssorhynchus, if they are long, 


108 Records of the Indian Museum. DVO felis 


sharp-pointed, and very narrow, it belongs to the genus Nyssomy- 
zomyia. 

Hints. (1) When beginning the study of classification by 
scale ornamentation always select good, unrubbed specimens. If 
mosquitoes bred from larvee are being used do not kill them until 12 
hours have elapsed from the time when they hatched out. The 
best specimens are those which, bred under favourable conditions 
from larve, have been allowed one meal of blood and then kept 
in suitable jars until the meal has been digested. (2) Do not use 
card-discs for mounting the mosquitoes to be used in studying 
this subject. Cut off the head of the fine silver pin and mount 
the mosquito on the pin so that an equal length of the pin pro- 
jects from the dorsum and from the venter. ‘Then stick either the 
point or the head end of the pin into a small block of pith or of 
cork. For examination, the mosquito can now be pinned with 
the dorsal, ventral, or lateral surface uppermost and a _ proper 
view obtained. When card-discs are used this cannot be done, 
because the disc is always in the way and hides the part one 
desires to examine. (3) A knowledge of whether the scales are 
false scales or true scales is not necessary for determining the 
genus, but if it is desired to study this matter thoroughly, the 
scales must be mounted flat on a slide and examined with a high 
power objective. It is essential to carry out the same procedure 
when it is desired to study thoroughly the shape of various scales. 
(I refer of course to a closer study than is necessary for ascertain- 
ing whether the upright forked scales of the head are rod-shaped 
or broadly expanding, whether the thoracic scales are sharp- 
pointed and very narrow or blunt-ended and broad, etc.). The 
following is the simplest method of obtaining the particular scales 
one wishes to study. Dissect out with a sharp knife the part of 
the mosquito that carries the scales and place it on a slide under 
a cover-glass. Tap and press the cover-glass slightly, then remove 
it and slide the part of the mosquito away with the point of a 
needle. Replace the cover-glass and fix it with strips of gummed 
paper. Many scales will have remained on the slide during these 
manipulations. Examine them with a 7: inch oil immersion lens 
and draw them with the aid of an eyepiece camera lucida. (4) In 
describing the shapes of scales it would be of great advantage if 
all observers were to adopt the exact nomenclature used by 
botanists in describing the shapes of leaves. This nomenclature is 
given in all the elementary books on botany and is very applicable 
to the shapes of scales. The botanical terms acicular, linear, 
oblong, elliptical, rotundate, orbicular, lanceolate, ovate, oblan- 
ceolate, obovate, spatulate, etc., have a definite meaning and 
significance. ‘‘Tanceolate’’ is a botanical term commonly used 
by entomologists, but it is used by them indifferently for scales of 
very various shapes; a lanceolate leaf is broadest at the base and 
scales with that character are seldom or never seen, though oblan- 
ceolate scales arevery common. ‘he use of such terms as “ slight- 
ly lanceolate,’ ‘‘long and narrow,’’ ‘‘ large and inflated,” etc., 


1910. | S. P. James: The Indian Anopheline. 109 


can be criticised not only because these terms are so indefinite, but 
because it would be easy to find botanical terms that would des- 
cribe the shape of the scale in a single word that conveys the 
same meaning to everyone. (5) It is particularly to be noted that 
no detailed arrangement by scale ornamentation can be made that 
will apply to both females and males. ‘The scale ornamentation 
(especially on the last segments of the abdomen and genital pro- 
cess) of male anophelines often differs considerably from that of 
female anophelines. The arrangement described in this paper 
applies throughout to the female insect only. 


26-5-10. 
ADDENDUM. 


The issue of vol. v of Mr. Theobald’s Monograph while this 
paper was passing through the Press has made it necessary to 
add in the form of an appendix the characters of a new genus, 
NEOMYZOMYIA, described by him. He has created the genus 
for the reception of elegans, James, a species that I have not seen 
for some years. The generic characters are shown in the following 
statement :— 


| 


Abdomen. Thorax. | Head. Prothoracic lobes. 
| 


: wate a : 
Only the last segment The anterior|With the usualtype With a tuft of 
carries scales. They| promontory, ofbroadlyexpand-| long out-stand- 
are arranged as in| carries agroupof| inguprightforked ing scales, 


the genus Nyssor-| sharp-pointed) scales which form | 


hynchus. | curved scales| dense tufts at | 
| which project | the postero-lateral) 
over the neck.| corners. | 


The remainder of | | 
| the dorsum is | 
| clothed with | 
| hairs, | | 

| 


| 


It will be seen that the genus is distinct from any of those 
described above. It comes near to the genus Patagiamyia, but 
the presence of scales on the last segment of the abdomen sepa- 
rates it very distinctly and makes us place it in the second of the 
two great groups. Mr. Theobald says that the scales of the wing 
are Myzorhynchus-like and that the palpi are densely scaled, so 
that the chief difference between the new genus and the genus 
Myzorhynchus must be that in Myzorhynchus the scales of the 
last abdominal segments are arranged as a ventral tuft while in 
Neomyzomyia they are not aggregated to form a tuft but are 
disposed irregularly as in the genus Nyssorhynchus. ‘The name 
Neomyzomyta is rather misleading. 

Deke ys 


5-9-10. 


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EXPLANATION: OF “PVATE 7 LS 


The figures show from left to right the upright forked scales 
of the head, the hairs or scales on the dorsum of the thorax, 
and the hairs or scales on the prothoracic lobes. The drawings of 
these structures relate respectively to 


The genus NEOSTETHOPHELES (Fig. 1.) 
ms ,,  MyYZOMYIA (Hise) 
Sy oy EAT AGTAMY TA (Fig. 3.) 
PYRETOPHORUS (Fig. 4.) 


In Fig. 2 the three thoracic scales shown are false scales ; in 
Fig. 3 one scale is a false scale and two are true scales; in Fig. 4 
all are true scales. 


Plate IX. 


Zl Ind. Mus.,Vol.IV, 1910. 


o 


D.Bagchi, lith. 


S.P. James, del. 


: t anh - a Sse Met ie , : ; 
si Nf 8a, 2 a i) a * mers) 1 Oe: re fe eres C = : 


y wr wt * 
: 5: eden! * : 
: es eA eee es ane vat ay Paal, : j x 
cote OF = ar 7 7 
SES? tse 
Fd iT 


ey Es a. 


te § Ss ouee meet a 
es 


5 7 a 
ele at > PS east ee = 


ae s 8A a os 
ee se eee De f Soh ae . Pee - 
| i tg ae ee : 
b oe, Ps at el sy" Te * +. . ae _ iy 
ee BE ey ys" 
eed} Pigezt Fhe if Me FE thi, a rt pire sts y a rer 


sts 272 eh eee kere lke hie add 53 sory Ree artic nie ik » 
ated ee ies lah Babe: Arad) mM ote Soe of. 
EY OHEP Satie are oer slag te Teg ks Sty omens * 
Bi eit rena tohiys MPP ee iis 3 enre!s 3? ee 
tee ay: Mg, ST) er ie Ms ae fick Bree AGS) hag aa 


a. > eehe i a aa 
+n 
@ 
ea 7 @ ‘ 
7 “is 
al 
— =. = 
* = = 
’ . 


EXPLANATION: OF PLATE 3 


The upper half of the plate (Figs. 1 to 6) 


relates to the genus .. NYSSORHYNCHUS. 
The lower half is the plate (Figs 7 £Ov EB) 
relates to the genus .. NYSSOMYZOMYVIA. 


Details of the upper half of the plate—-From left to right the 
drawings are, (1) abdomen from the dorsal aspect, (2) abdomen 
from the ventral aspect, (3) last segment of abdomen and genitalia, 
(4) thorax. Fig. 5 shows three of the abdominal scales as 
drawn with the aid of a camera lucida under a 75-inch oil immer- 
sion objective and low eyepiece. Fig. 6 shows some of the 
thoracic scales drawn by the same method. The three thoracic 
scales to the left are from fuliginosus and the three to the right 
are from maculatus. 

Details of the lower haif of the plate-—Genus Nyssomyzomyia. 
The thoracic and abdominal scale ornamentation of ross? is shown. 
Fig. 8 shows the ventral surface of the last abdominal segments. 
The long pointed false scales and the hair on the right of the plate 
are from the anterior promontory and mid region of the dorsum of 
the thorax; and the two true scales below the short black line. 
(Fig. 11) are from the ventral surface of the last abdominal seg- 
ment. 


Placee ee 


Rec. Ind. Mus., Vol. JV, 1910. 


SES 
= 


5.P. James, del 


EXPLANATION OF PLATE XI. 


The upper half of the plate (Figs. 1 to 5) 


relates to the genus ie 7 / . NBOCELLIA: 
The lower half of the plate (Figs. 6 to 10) 
relates to the genus ze CELLIA. 


In both cases the following structures are shown: (1 and 6) 
abdomen from the dorsal aspect, (2 and 7) abdomen from the 
ventral aspect, (3 and 8) abdomen from the side; (4 and 9) 
abdominal scales and (5 and 10) thoracic scales. As regards the 
genus Neocellia some of the scales (the larger ones) are from willmort 
and others (the smaller) from stephensi. As regards the genus Cellia 
all the scales are from pulcherrima. 


ale. 


Plate, XI. 


ec. Ind. Mus.,Vol IV, 1910. 


Sy 


py? 


PAs 


D. Bagchi, hth 


'S.P. James, del. 


ee “y 


¢ar™ 


oan 


‘ 


f 
“Se erp 


Nie Ge os 
tg 


far 


ai 


EXPLANATION OF PLATE XII. 


The upper half of the plate (1.) relates to the 


Genus =. ig ae MyZzORHYNCHUS. 
The lower half of the plate (2.) relates to the 
genus ave re CHRISTOPHERSIA. 


In both cases the scale ornamentation of the abdomen, thorax, 
and prothoracic lobes is shown. ‘The two scales below the lateral 
view of the abdomen in I. are from the ventral abdominal tuft. 


2g 


Plate 


fad Mas.Vel IV, 1910. 


3 
piay/ shat 


,lith 


) 
on 
‘a 
pr 
a 
a 

| Q 
IL pe o 
Ss = Ss - 2S SSS o 
2 SSS a 
——< S 
uP) 

aw : 


1 , : 
> 
’ 
7 ~~. 4 
s “- 
> “ 2 
= , 
7 
7 * 
‘ 
‘/;, = 
*~ 
. 
7 
yy 
. t 
2 
. 
ny 
‘ 
ply 
Ae . 
“a . 
x - 
r 
28 P 


c =Te 


Other Publications edited and sold by the Superintendent of the Indian 
Museum (also obtainable from Messrs. Friedlander & Sohn) issued 
by the Director of the Royal Indian Marine. 


‘Illustrations of the Zoology of the R.I.M.S. ‘‘ Investigator ’’ 1892, Fishes, Plates 1 to VII. 


_ €rustacea, Plates I to V, 1894. Fishes, Plates VII to XIII. Crustacea, Plates VI to VIII. Ech- 


inoderma, Plates I to III, 1895. Echinoderma, Plates IV and V. Fishes, Plates XIV to XVI. 
Crustacea, Plates IX to XV, 1896. Crustacea, Plates XVI to XXVII, 1897. Fishes, Plate XVII. 
Crustacea, Plates XXVIII to XXXII. Mollusca, Plates I to VI, 1898. Fishes, Plates XVITT te 
XXIV. Crustacea, Plates XXXIII to XXXV. Mollusea, Plates VII and VIII, 1899. Fishes. 
Plates XXV and XXVI. Crustacea, Plates XXXVI to XLV, 1900. Fishes, Plates XXVII to 
XXXV: Crustacea, Plates XLVI to XLVIII. Index, Part I, 1901. Crustacea, Plates XLIX 


. to LV. Mollusca, Plates-IX to XIII, 1902. Crustacea, Plates LVI to LXVII. Crustacea, Plates 


LXVIII to LX XVI. Fishes, Plates XXXVI to XX XVIII, 1905. Crustacea (Malacostraca), Plates 
LXXVII to L.XXTX. Crustacea (Entomostraca\, Plates IT and TI. Molinsca, Plates XIV to 
XVIII, 1907. Fishes, Plates XXXIX to XLIII. Crustacea (Entomostraca), Plates III to V. 
Mollusca, Plates XIX and XX, 1908.—Re. 1 per plate, Mollusca, Plates XXI to XXIII, 1909.— 


As, 8 per plate. 


RECORDS 
of the 


INDIAN MUSEUM 


Vol. I, 1907. 


Part I.—Coutributions to the Fauna of the Arabian Sea, Records of Hemiptera and 
Hymenoptera from the Himalayas. Further notes on Indian Freshwater Ento- 
mostraca. The Faunaof Brackish Ponds at Port Canning, lower Bengal, I—III 
A Sporozoon from the Heart of a Cow. Miscellanea, 


Part II.—Revision of the Oriental Stratiomyide. Description of an Oligochete 
Worm allied to Cha@togaster. The Fauna of Brackish Ponds at Port Canning, 
Lower Bengal, IV. Further Note on a Polyzoon from the Himalayas. Reports 
on a collection of Batrachia, Reptiles and Fish from Nepal and the Western 
Himalayas. The Fauna of Brackish Ponds at Port Canning, Lower Bengal, V. 
Notes on Oriental Diptera, I and II. Miscellanea, 


Part IIT.—Report on the Marine Polvzoa in the collection of the Indian Museum. 
The Fauna of Brackish Ponds at Port Canning, Lower Bengal, VI. A third note 
on Earwigs (Dermaptera) in the Indian Museum, with the description of a new 
species. Notes on Oriental Diptera, III. Description of a new snake from Nepal. 
Notes on a collection of marketable fish from Akyab, with a description of a new 
species of Lactarius. Description of two freshwater Oligochete Worms from the 
Punjab. Notes on Phosphorescence in Marine Animals. Notes on the rats of 
Dacca, Eastern Bengal. Notes on Freshwater Sponges, I—V. Miscellanea. 


Part IV.—Nudiclava monocantht, the type of a new genus of Hydroids parasitic 
on Fish. Preliminary descriptions of three new Nycteribiide from India| Anno- 
tated Catalogue of Oriental Culicide. Notes on Oriental Diptera. Notes on 
Freshwater Sponges, VI, VII. Description of a new Cyprinid Fish of the genus 
Danio from Upper Burma. Miscellanea, 


Vol. II, 1908-1909. 


Pari I.—The retirement of Lieut.-Col. Alcock, with a list of his papers, etc., on Indian 
Zoology. The Fauna of Brackish Ponds at Port Canning, Lower Bengal, VIT. 
Description of a New Dictyonine Sponge from the Indian Ocean. Notes on 
Freshwater Sponges, VIII. Remarkable cases of variation, I. Description of a 
new species of Lizard of the genus Salea, from Assam. The Fauna of Brackish 
Ponds at Port Canning, Lower Bengal, VIII. Description of a new Cavernico- 
lous Phasgonurid from Lower Siam. Descriptions of new species of Marie and 
Freshwater Shells in the collection of the Indian Museum. Notes on Oriental 
Syrphide, I. Description of a new variety of Spongilla loricata. Notes on 
Oriental Diptera, V. Miscellanea. 


Part IT.—Gordiens du Musée Indien. The Fauna of Brackish Ponds at Port Canning, 
Lower Bengal, IX. Description of a new species of Danio from Lower Burma. 
Rhynchota Malayana, I. Cimex rotundatus, Signuoret. Notes on Freshwater 
Sponges, IX. Fruit Bats of the genus Pleropus inhabiting the Andaman and 
Nicobar Archipelagos. A new species of Sun-Bird obtained near Darjiling. 
Three Indian Phylactoleimata. On two new species of Eagle-Rays (Myliobatide), 
Description of a new species of the genus Sesarma, Say., from the Andaman 
Islands. Descriptions of new species of Land, Marine, and Freshwater Shells from 
the Andaman Islands. 


Part IIT.—The Fauna of Brackish Ponds at Port Canning, Lower Bengal, X, XI. On 
some Oriental Solifuge with descriptions of new forms. The difference between 
the Takin (Budorcis) from the Mishmi Hills and that from Tibet, with notes 
on variation displayed by the former. On Caridina nilotica (Roux) and its 
varieties. Description of a new species of Charaves from the Bhutan Frontier. 
First Report on the Collection of Culicide and Corethride in the Indian Museum, 
with descriptions of new genera and species. Miscellanea 


Part IV.—Report on a collection of aquatic animals made in Tibet by Capt. F. H. 
Stewart in 1907, I. Notes on Aculeate Hymenoptera in the Indian Museum, I, 
Indian Psychodide, Description of a new species of mouse from the Madura 
District, Madras. Some Cleride of the Indian Museum. The Fauna of 
Brackish Ponds at Port Canning, Lower Bengal, XII. Description of a new 
species of Saw-Fish captured off the Burma Coast. A new Sting Ray of the 
genus Trygow from the Bay ot Bengal. New Micro-lepidoptera from India and 
Burma. Notes on some Chrysomelid Beetles in the collection of the Indian 
Museum. Six new Cicindeline from the Oriental Region, Description of a 
new slug from Tibet. 


ea Laan 


Mie TS FLOW SOF THESE CIES ‘OF 
Lea OS FROM THE OR LTEN TAT RE - 
GION, INCLUDING. NOTES.ON SPECIES 
FROM SURROUNDING COUNTRIES. 


By GERTRUDE RICARDO. 


The Oriental Region in this paper includes India, Burma, 
Federated Malay States, Siam, Annam, Sumatra, Borneo, Java, 
and other islands, the Philippines, Formosa, and the southern 
part of China, as defined by Wallace and others, but Celebes is 
not included, following the eastern dividing line adopted by 
Sclater, so that, as a rule, the species from Celebes are not 
included here, but relegated to the Austro-Malay Region ; judging 
from the Bigot and Walker types I have seen from Celebes, they 
certainly (with one or two exceptions included here) appear to 
have little affinity to the Indo-Malay, Borneo and Java species. 

The material on the examination of which this paper is based, 
includes the British Museum collection, a collection sent me by 
Dr. Annandale from the Indian Museum for identification, a 
smaller collection from Mr. F. M. Howlett, from the Pusa Agri- 
cultural Research Institute, Bengal; besides species sent by Dr. 
Kertesz from the Budapest Museum. ‘To Dr. A.’T. Stanton I am in- 
debted for an interesting series of species collected in the Federated 
Malay States. From Col, Alcock I have had a small collection to 
identify belonging to the London School of Tropical Medicine. 
Through the courtesy of the Director and of M. Jacques Surcoutf 
of the Paris Museum, I have been able to examine most of the 
Macquart and Bigot types in their possession. ‘Through the 
courtesy of the authorities of the Natural History Museum 
at Genoa, I was able to examine Rondani’s types in their 
collection. I am indebted to Mr. G. H. Verrall for his loan 
of the majority of the Bigot types, which, as a rule, would 
be very difficult to recognize from the descriptions, based in 
many cases on single specimens, and now at any rate in a 
very denuded condition. Bigot labelled many of the specimens 
in the Indian Museum collections, but most of them are merely 
MS. names, where he made a new species and published the descrip- 
tion, he evidently interchanged the specimens with the Museum and 
his own collection, so that some of Mr. Verrall’s Bigot types have 
co-types from the same series of specimens in the Indian 
Museum collection. His specification of localities is very vague, 


ale? Records of the Indian Museum. Vio. aye 


‘““India’’ alone frequently being given for Assam; Dr. Annandale 
informs me in a letter that M. Bigot destroyed most of the original 
labels and substituted others in which only ‘‘Inde’’ was written. 
Owing to denudation it is frequently difficult to identify species of 
Tabanus, and probably with the advent of fresh material much 
rectification may be necessary, especially as regards the identifica- 
tion of the older species. The synonyms given here are very 
numerous, but have been chiefly determined from the more 
important plastic characters, such as breadth of forehead, 
its. greater or less narrowness anteriorly above the eyes, 
and the shape of the frontal callus; beth these characters 
vary, but to a small extent only; the shape of the palpi is 
also a valuable character if not relied upon too minutely ; 
the absence or presence of an appendix on the wing is a very 
variable, uncertain, character; the closed or extremely narrow 
first posterior ‘cell at border of wing appears fairly reliable, 
but males and females probably vary in this character. Rondani 
formed the genus Bellaydia for species possessing this character 
(Archiv. Canestr., iii, p. 81, 1863), but it does not appear a good 
generic character and has not been adopted by other authors. 
The design and colouring of the abdomen must naturally be largely 
made use of, though owing to the easy denudation of the tomen- 
tose dust which usually forms the spots and stripes, it is often 
difficult to identify species; the colouring of the hairs on body and 
legs is useful, but it appears to vary in specimens of the same 
species to some extent, especially as regards the legs; the shining 
polished subcallus in some species appears a reliable distinct 
character, though here again denudation must be borne in mind. 
The subgenus Therioplectes, Zeller, formed for species with pubes- 
cent eyes and with an ocelligerous tubercle on vertex of forehead, 
appears applicable to the species of the Oriental Region, though 
their number is very small; but it is otherwise with the subgenus 
Atylotus, Osten Sacken, formed for species with pubescent eyes but 
no ocelligerous tubercle, some species such as the widely distributed 
Tabanus taenvatus, Macq., and other species from this Region have 
an Atylotus-like appearance, that is to say, they resemble the 
group containing fulvus, rusticus, and others, of a mealy coloured 
appearance, Brauer placed black and other species in this group, 
in his work on European Tabanidae ; but the eyes are practically 
bare in both sexes, so that this subgenus as in the South African 
fauna is not here applicable. The grouping of the species has 
been again attempted as in those of the South African Region, but 
where it is based upon the presence or absence of spots on the 
abdomen it, will probably need correction in the future, the group 
represented by Tabanus funufer, Walker, being one of the most 
difficult in this fauna. The table of species has been drawn up for 
the females, only males are included in a few instances where 
specified ; as is always the case, the males in the collections are very 
few in number. The measurement of the length of the forehead is 
based on the width anteriorly above the eyes. In the references 


IQITI.| G. Ricarpo: Revision of the species of Tabanus. II3 


here to Macquart’s Dipt. Exot. volumes the original pagination 
is used, unfortunately in some of the reprints of the work from 
Mem. Sci. Agri. et Arts de Lille, 1838—1854, in which publication 
the work first appeared, the original pagination has not been 
adhered to, and in this incorrect form has been in general use. 
As the descriptions by Walker and Bigot are, as a rule, too poor for 
identification of the species, it appears justifiable to delete from 
the list of species of Tubanus any types of either of these authors 
which are now lost or destroyed. All types, except those of Bigot 
and Macquart which are either in Mr. Verrall’s possession, or in the 
Paris Museum, or as in the case of Macquart in the Lille Museum, 
are in the British Museum collection unless otherwise specified. 

There were about 119 described species of Tabanus from the 
Oriental Region, now reduced by synonyms, deletion from list or 
not belonging to the Oriental Region to about 73 species, which 
with the addition of 40 new species described here (4 were pub- 
lished first in the Annals) brings the total to 117 species, including 
one species from Celebes, and 3 species from other Regions occur- 
tring here. The table for species does not include those species of 
which I have no knowledge, except in a few instances where the 
descriptions allow of their being included with some certainty. 

The drawings in the two plates have been executed by Miss 
Grace Edwards. 


TABLE FOR GROUPS. 


I. Eyes bare res ie 3 lee 
Eyes hairy (Thertoplectes) = GROUP esi.) (OL. 
II. Forehead with two separate calli, the first 

one usually transverse, the second irre- 

gular in shape, smaller. Forehead four 

to six times as long as it is broad. Small 

species 9—1I4 mm. in length ie TE 
III. Thorax and scutellum same colour. Ab- 

domen with no bands ey Pe pGROUP)I, I. 


Scutellum white or yellow-haired. Abdomen 
with white or yellow-haired bands .. Group II. 4. 

IV. Forehead with two small round spot-like 
calli, one above the other, never united .. Group III. 7. 
V. Forehead with no callus Ag .. GROUP a 

VI. Forehead with one square, narrow, or oblong 


callus, which is always prolonged towards 
the vertex by a more or less narrow line. . VII. 


VII. Wings with spots or bands .. .. Group V. Io. 


Wings distinctly marked with brown or yellow 
colouring, but not in the form of bands. 
Large species 18—23 mm. in length .. Group VI. 13. 


I14 Records of the Indian Museum. [Vox. IV, 


Species with one or more stripes usually 
continuous on abdomen. Forehead five 
to nine times as long as it is broad > GROUP VILLE a7s 


Species with median or lateral spots or both 
on abdomen, not usually forming a conti- 
nuous stripe. Forehead four to ten times 
as long as it is broad a 2. GROUP VITIE S36: 


Species with paler bands and sometimes spots 
onabdomen. Forehead four to ten times 
as long as it is broad ay +32 GROUP JEXe 93538 


Species with the abdomen unicolorous or 
almost so, sometimes darker at apex. 
Forehead five to ten times as long as 
it is broad 32 : nf GROUP X. 765. 


VIII. Species with eyes pabeseeat and an 
ocelligerous tubercle on forehead at 
vertex aN ye 1 GROUP DU: 


TABLE FOR SPECIES. 


Group I. 
1. Subcallus shining. ie with no 
cross-bands 2. 
Subcallus not shining. Eyes with 
cross-bands 2 


. Frontal callus reaching eyes. Ab- 
domen grey, covered with long 
white pubescence. Tibiae whit- 
ish with long fringes of white 
hairs ane . varus, 1. Sp. 
Frontal callus not reaching eyes. 
Abdomen black, shining. Tibiae 
whitish, but with no long fringes 
of white hairs on .. hirvitpalpis, n. sp. 
3. Frontal callus reaching eyes. Black 


species with three grey stripes on 
abdomen ie .. bicallosus, Ricardo. 


Group II. 

4. Dark brown or yellowish brown 
species with yellow-haired scutel- 
lum and yellow-haired bands on 
abdomen Be 3 5. 

Black species with white-haired 
scutellum and white-haired bands 
on abdomen 

5. Abdomen reddish brown or yellow 
with six narrow yellow bands. 


Igtt.] G. Ricarpo: Revision of the species of Tabanus. II5 


Palpi pale yellow with black and 

yellow pubescence. Fore tibiae 

pale on basal two-thirds only .. flavicinctus, n. sp. 
Abdomen dark reddish brown or 

blackish with two broad yellow 

bands. Palpi black with grey 

tomentum and white and black 

hairs. Fore tibiae pale to apices... leucocnematus, Bigot. 

6. Abdomen black with two white 

bands. Frontal callus almost 

reaching eyes. Fore tibiae pale on 

basal two-thirds only. Wings 

tinged brown on fore border .. bicinctus, n. sp. 
Abdomen with six white bands. 

Frontal callus not reaching eyes. 

Fore tibiae pale to apices. Wings 

clear a Ee .. sexcinctus, n. Sp. 


Group III. 


7. Forehead slightly narrower anterior- 

ly, three and a half to four times 

as long as it is broad bis 8. 
8. Abdomen blackish covered with 

golden yellow tomentum and ap- 

pressed yellow pubescence .. fulvus, Meig. 

Abdomen yellowish with median 
and lateral blackish or brownish 
Stripes: is. a _. ditaeniatus, Macq. 


Group IV. 


g. Abdomen brown, reddish at base, 

covered with grey tomentum and 

with two small round greyish spots 

on the segments. Wings with no 

appendix. Length7}to1I mm. virgo, Wied. 
Abdomen blackish brown with me- 

dian and lateral greyish tomentose 

spots. Wings with an appendix. 

Length 13 mm. 7 _. nemocallosus, Ricardo. 
Abdomen covered with greyish yel- 

low tomentum and with thick 

short yellow pubescence, 2: abdo- 

men the same but a black mediar 

stripe often appears, ~. Wings 

with noappendix. Length 113-12} 

mm. ae Be .. negativus, n. Sp. 


Group V. 


10. Wings with four dark spots. Abdomen 
reddish brown, with white spots javanus, Fabr. 


116 Records of the Indian Museum. {Vor,.. EV 


Wings with one cross-band, the first 
posterior cell open a 
Wings with two cross-bands, the 
first posterior cell closed or very 
narrow He 
11. Forehead narrowest at ‘vertex. Ab- 
domen reddish, darker at apex, 
with small white median spots 
Forehead narrowest anteriorly. Ab- 
domen yellowish, darker at apex, 
with larger white spots : 
12, Abdomen brownish yellow, with 
broad yellow posterior bands on 
each segment, under side yellow- 
ish. Antennae reddish yellow. & 
with equal facets to eyes 
Abdomen brownish red with narrower 
yellow bands, under side darker. 
Antennae reddish yellow, black at 
apex. o with unequal facets to 
eyes 


Group VI. 


13. Wings dark rich brown, the discal 
cell clear. Large black species 
with orange-red pubescence on 


face and breast. Antennae red .. 


Smaller species similar in Sain 
Antennae black 
Wings yellowish or brownish, very 
distinctly tinged with brown at 
apex and on _ posterior border. 
Scutellum yellow or greyish : 

14. First posterior cell of mass closed or 
very narrow 

First posterior cell open, not. very 
narrow . 

15. Abdomen reddish brown, narrow, 
with grey tomentose bands and 
median spots. Palpi slender. 
First posterior cell closed 

Abdomen brown with narrow grey 
tomentose bands but no median 
spots. Palpi large, broad. The 
first posterior cell very narrow at 
DOLGEL a As " : 

Abdomen black, livid at base. 
Smaller species. First posterior 
cell closed 


. optatus, Wik. 


. non-optatus, n. sp. 


. flexilis,. Wik. 


vanderwulpi, O. S. 


pratti, n. sp. 


. fuscicornis, n. sp. 


nephodes, Bigot. 


. albofasctatus, n. sp. 


basalts, Macq. 


J} s 


Ab 
I5. 
16. 


rgl1.j G. Rricarpo: Revision of the spectes of Tabanus. 117 


16. Large yellowish species with blackish 
bands on the abdomen .. auriflamma, Wlk. 


Group VII. 


17. First posterior cell closed.  Yellow-_ 
ish species with indistinct median 
black stripe on abdomen .. annamitus, &., Bigot. 
First posterior cell open 18. 


. Frontal callus oblong or narrow its 

lineal extension. Forehead at 

least five times as long as it is 

broad ats Pee IQ. 
Frontal callus large, somewhat pro- 

tuberant, almost square, with 

short lineal extension. Forehead 

three or four times as long as it 


is. broad... Ys ! 36. 
1g. Abdomen with a median stripe aed 
lateral stripes or spots 20. 


Abdomen with one median ne 
nuous stripe only, no lateral spots, 
or only isolated ones on the second 


SEQIMENE ae 27. 
Abdomen with lateral stripes only . 35. 

20. Frontal callus with a spindle- Aneel 
lineal extension ur als 


Frontal callus with.the lineal exten- 
sion not spindle-shaped 
21. Abdomen blackish or reddish brown 
with grey median and _ lateral 
stripes, almost. all the same 
tenet, 3.7. . styiatus, Fabr. 
Abdomen with the grey median stripe 
beginning only on the third seg- 
ment, and with lateral stripes end- 


ing on the third or fourth seg- . 
ment fe. a .. Atlaris, Wik. 


N 
N 


22. Abdomen with a median stripe, the 

side spots small.and indistinct 

Abdomen with a median stripe, the 

side spots large and distinct .. 24. 
23. Smaller species. Abdomen reddish 

yellow, darker at apex. Thorax 

blackish. Forehead eight times : 

as long as it is broad .. abbreviatus , Bigot. 
24. The median abdominal stripe com- 

posed of almost equal sized spots, 

not very large 


iS) 
Oo 


25. 


26. 


27. 


28. 


20. 


30. 


ae 


Bae 


33: 


34. 


Records of the Indian Museum. 


The median abdominal! stripe com- 
posed of spots of unequal size, 
those on the third and fourth seg- 
ments very large and conspicu- 
ous ae 

Abdomen aad thorax lilac or black- 
ish brown, femora blackish. 
Forehead about five times as long 
as it is broad 

Abdomen and thorax pboeecely oe 
dish brown, femora reddish brown. 
Forehead five to six times as long 
as it is broad : 

Abdomen and thorax reddish brown, 
femora blackish. Forehead about 
seven times as long as it is broad 

Median stripe of abdomen grey 
tomentose or white or yellow 
haired : os 

Median stripe of abdomen black... 

Median stripe of abdomen golden 
haired ae 

Forehead barely elie as wide an- 
teriorly as it is at vertex 

Forehead parallel or nearly so 

Small brown species. Femora brown- 
ish. Wings clear, brown on fore 
border 


Large black or reddish Grown species, 
medium abdominal stripe narrow. 
Femora biack. Wings tinged 
brown Se 

Brown species, the median abdo- 
minal stripe broad, and two round 
spots on second segment. Femora 
reddish. Wings brown on fore 
border 

Abdomen reddish, Bree at apex, a 
narrow grey stripe often apparent 
on the median black stripe 

Forehead parallel 

Forehead narrower anteriorly 


Abdomen yellowish brown, darker 
at apex, the median stripe dull 
golden haired indistinct 

Abdomen vellowish brown or dark 
brown, the median stripe broad. 
Fore femora yellowish. Wings 
brown, especially on fore border 


. rubidus, Wied. 


. albimedius, W1k. 


speciosus, n. sp. 


. hortistriatus, n. sp. 


. abscondens, Wik. 


. auristriatus, n. sp. 


. hybridus, Wied. 


[Vor IV, 


26. 


29. 
30. 


. monotaentatus, Bigot. 


. brunnipennis, n. sp. 


33. 
34. 


19g1II.} G. RICARDO: Revision of the species of Tabanus. 119 


Abdomen covered with yellowish 
tomentum, median stripe very 
broad. Fore femora black. Wings 
clear are # : 

35. Black species. Abdomen with ferru- 
ginous lateral stripes on the first 
four segments. Legs blackish 
brown 

36. Forehead narrowest anteriorly 

Forehead narrowest at vertex 

37. Small black species. Abdomen with 
a median grey stripe beginning on 
the third segment only, the lateral 
stripes terminate on the second 
Segment. =: Ae 

38. Small brown species. Abdomen with 
a median grey stripe 


Group VIII. 


39. Wings with first posterior cell closed 
or at least very much narrower at 
border. Forehead six to seven 
times as long as it is broad 

Wings with first posterior cell open, 
never very narrow at border 

40. Abdomen with three series grey spots 
on abdomen, reddish brown in 
male, blackish in female 

Abdomen with three grey spots on 
the first two or three segments 
only, a larger, broader bodied, 
blackish species 

41. Forehead narrow, six to ten times ; as 
long as it is wide anteriorly, with 
a long narrow frontal callus 

Forehead broader, four to five times 
as long as it is wide anteriorly, 
with a larger nearly square, or 
pear-shaped frontal callus ‘ 

42. Brown, reddish brown, or reddish 
yellow, medium sized or large 
species, the triangular greyish or 
yellowish spots not usually form- 
ing a continuous stripe 

Large unicolorous reddish species 
with small median indistinct white 
spots a 

Reddish brown or brown species with 
large prominent median whitish 


. aurotestaceus, W1k. 


. tristis, v. d. Wulp. 


37: 
38. 
. qucundus, Wik. 
. puteus, n. sp. 
40. 
4l, 


. amaenus, Wik. 


. mandayinus, Schiner. 


42. 


IZ0 


Records of the Indian Museum. 


spots on the third and fourth seg- 
ments only an 


Reddish species with black spots 
Small species with a narrow yellow 


abdomen irregularly marked with 
black and with three series irre- 
sular shaped grey median and 
lateral spots 


43. Medium sized species, 17-18 mm. in 


length. Abdomen reddish yellow 
with small median white spots. 
Forehead almost parallel or bare- 
ly a third narrower anteriorly, six 
times as long asitisbroad. Fron- 
tal callus oblong not reaching the 
eyes. Wings clear, faintly tinged 
yellowish brown 


species usually 18-25 mm. in length, 
with abdominal median _ spots. 
Forehead from one-third to halt 
narrower anteriorly, frontal callus 
long and narrow 


44. Forehead eight to nine times as long 


as it is broad, about one-third 
narrower anteriorly } 


Forehead very narrow, eight to oa 


times as long as it is peoaa only 
half as wide ‘anteriorly as at ver- 
tex 


45. Abdomen reddish, or reddish yellow, 


large, broad, with median short 
triangular yellow spots often indis- 
tinct. Fore tibiae reddish yellow 
on basal half, with black pubes- 
cence. Wings large, tinged 
yellowish brown or almost clear. 
Forehead almost parallel 


row, with median long triangular, 
distinct whitish spots, usually 
joining each other. Fore tibiae 
white on basal half with white 
pubescence. Wings long, faintly 
tinged yellowish brown.  Fore- 
head about a third narrower an- 
teriorly .. Ae 


most hemispherical white median 


.. varicolor; Ricardo. 


.. vubicundus, Macq. 
Large brown or reddish brown 


. stantont, n. sp. 
Abdomen reddish brown, long, nar- 


.. indianus, 0. sp. 
Abdomen blackish brown with al- 


[Vou.1ve 


50. 
Se 


44. 


45. 


46. 


1git.] G. Ricardo: Revision of the species of ‘Tabanus. I21 


spots. Fore tibiae blackish. Legs 
and palpi blackish. Wings clear, 
tinged brown. Forehead about a 
third narrower anteriorly .. brunneus, Macq. 
46. Forehead eight times longer than it 
is broad, the frontal callus not 
reaching eyes 47. 
Forehead ten times as long as it. is 
broad, the frontal callus 1 reaching 
eyes sé 48. 
47. Abdomen dark reddish proven or eas 
dish yellow, with indistinct small 
greyish or yellow-haired median 
spots, often absent. Thorax black- 
ish with some yellowish grey 
tomentum. Fore tibiae pale on 
basal half. Wings slightly tinged 
with brown. Male with unequal 
facets on eyes ne ... fumtfer, Wik. 
Smaller species. Abdomen reddish 
brown with distinct bright yellow- 
haired median spots and similar 
lateral spots on the 2-5th seg- 
ments. Thorax blackish with 
greyish yellow tomentum . malayensis, n. sp. 
Larger species. Abdomen broad, 
reddish brown with indistinct 
median spots and some lighter 
segmentations. Thorax reddish 
with some grey tomentum. Wings 
nearly clear. Fore tibiae pale on 
basal two-thirds: Male with 
equal facets to eyes .. .. factiosus, W1k. 
48. Abdomen reddish brown, median 
spots indistinct, small, whitish. 
Thorax reddish, thickly covered 
with ashy grey tomentum. Wings 
clear 7 . dissimilis, i. sp. 


49. Abdomen and thorax fedaee Phiniie! 
neatly devoid of pubescence. 
Wings clear. Palpi ending in an 
acute point 33 .. tenobilis, Rondani., 
Abdomen and _ thorax reddish. 
Wings tinged yellowish brown. 
Palpi ending in an obtuse point .. nexus, WIk. 
50. Red-brown species with the large 
median white spots on third and 
fourth segments of abdomen, and 
white spots on the lateral borders 


122 Records of the Indian Museum. [Vor..1V; 


of the first four et a Tibiae 
reddish .. .. Signijer, 7, WIk. 
Brown species with the same median 
spots, but with lateral white spots 
on the first two segments only. 
Tibiae white a .. significans, n. sp. 
51. Abdomen reddish, the apex dull 
black with large black median 
spots on the second and third seg- 
CETUS 8 yo fuscomaculatus, n. sp. 
52. Frontal callus square. ‘Small black 
species with white-haired median 
abdominal spots. Legs blackish, 
tibiae yellowish 3 .. leucopogon, Bigot. 
Frontal callus pear- shaped. Large 
reddish brown species, abdomen 
with median triangular spots. 


Legs reddish brown .. .. discrepans, n. sp. 
Group IX. 
53. First posterior cell closed ats 54. 
First posterior cell not closed Si 55- 
54. Large blackish species with narrow 
fulvous bands on abdomen .. servillet, Macq. 


55. Abdomen blackish brown, the first 
two segments covered with grey 
tomentum ae .. crassus, & , WIk. 
56. Forehead narrow, seven or eight 
times as long as it is broad. Fron- 
tal callus narrow with a lineal ex- 
tension. Black or brown species 57. 
Forehead broader, four or five times 
as long as it is broad. Frontal 
callus large with a stout, broad, 
or spindle-shaped extension ie 62. 
57. Large black or brown species 2I mm. 
in length, with very narrow te 
bands on abdomen .. 58. 
Smaller blackish species, 15-17 mm. 
in length, with whitish bands on 
abdomen = oe 59. 
58. Brown species. Scutellum and sides 
of thorax with pale tomentum. 
Legs blackish, tibiae obscurely 
reddish .. as .. alboscutatus, Bigot. 
Black species. Scutellum and sides 
with nosuch paletomentum. Legs 
black, tibiae yellowish white .. explicatus, Wik. 
59. Forehead narrower anteriorly ie 60. 


IgtI.] G. Ricarpo: Revision of the species of Tabanus. 123 


Forehead hardly narrower anterior- 
ly, but parallel z3 a 61. 
60. Frontal callus large, reaching the 
eyes. Antennae, legs, and palpi 
wholly black. Wings clear or 
tinged smoky brown .. micobarensis, Schiner. 
Frontal callus narrow. Antennae 
and legs blackish. Palpi yellow. 
Wings brownish ase . justorius, Rondani. 
61. Frontal callus narrow, not reachine 
eyes. Antennae and legs black: 
Palpi black, paler on _ insides. 


Wings faintly tinged brown .. khastensts, n. sp. 
62. Brown or black species with whitish 
bands and spots on abdomen... 63. 
Reddish yellow species with whitish 
bands and spots on abdomen... 64. 


63. Abdomen reddish brown or blackish 
brown with broad white bands 
and large triangular median spots. 
Palpi short and stout . sanguineus, W1k. 
Smaller species. Abdomen black 
with narrow white-haired seg- 
mentations and small triangular 
spots. Palpi slender .. oxyceratus, Bigot. 
64. Abdomen reddish yellow, darker at 
apex, with narrow yellowish white 
segmentations and median spots. 


Legs black, tibiae yellowish .. orventis, Wik. 
Abdomen similar, but the last four 
segments are black .. .. fulvimedius, Wik. 


Smaller species. Abdomen similar 
to that of orentis. Legs wholly 
yellow .. ate .. consanguineus, Macq. 


GRouP X. 


65. Black or brown species. Forehead 
narrower anteriorly, seven to ten 
times as long as it is broad a8 66. 
Large reddish yellow species from 
20-24 mm. in length. Forehead 
six to ten times as long as it is 
proada. ree 
Dark reddish brown species from 
18-22 mm. in length. Forehead 
seven to ten times as long as it is 
broad a 75. 
Reddish or reddish yellow species 


Records of the Indian Museum. [Vor. IV, 


from 16-19 mm. in length. Fore- 

head six to ten times as long as it 

is broad .. By, zi 706. 
Small species reddish, yellowish, 

brownish, or blackish, from 12-16 

mm.in length. Forehead five to 

ten times as long as it is broad .. 79. 
. Thorax paler than abdomen with 

yellowish tomentum and_ pubes- 

cence a iy 67. 
Thorax not paler than abdomen eS. 68. 
. Dark brown or blackish species with 

yellowish brown thorax. Legs 

black, tibiae white. Wings tinged 

yellowish brown ui . birmanicus, Bigot. 
Reddish brown species with light 

yellow or greyish thorax. Legs 

black, tibiae white, middle and 

posterior femora yellowish. Wings 

brownish, clear at apex .. flavothorax, n. sp. 


. Wings yellowish brown, yellow on- 

fore border. Black species. Tho- 

rax with short red pubescence .. ténctothorax, n. sp 
Wings brownish or black. Brown 

or black species te 69. 
. First posterior cell of wings ieee 

Abdomen deep black .. nigrotectus, Bigot. 
First posterior cell not closed fe. 70. 
. Blue-black species. Thorax red at 

base. Antennae reddish brown. 

Posterior tibiae red at base .. caerulescens, Macq. 
Forehead almost parallel. Abdomen 

dull reddish brown with greyish 

blue tomentum. Antennae red- 

dish yellow, black at apex. Tibiae 

dull red or blackish. Beard and 

palpi black bis tenebrosus, Wk. 
Forehead narrower, not parallel .. pee 
. Abdomen blackish. Antennae red- 

dish yellow. ‘Tibiae dull reddish. 

Beard white and black. Palpi 

yellowish 5s .. peraktensis, n. sp. 
Abdomen dull blackish, conical. 

Antennae black. Tibiae black. 

Beard and palpi black .. inobservatus, 1. sp. 


2. Forehead parallel, six times as long 


“NJ 


as it is broad : 73- 
Forehead narrower anteriorly, cight 
to ten times aslong asitis broad . 74. 


tgtt.] G. Rrcarpo: Revision of the spectes of Tabanus. 125 


73. Broad-bodied reddish-yellow species. 

Antennae yellow. Legs reddish 

brown ... fulvissimus, Rondani. 
74. Thorax brownish. Antennae reddish, 

the third joint black. Femora 


reddish brown .. wuventris, Wik. 
Thorax yellowish. Antennae, palpi 
and legs yellow aS. . flavissimus, 1. sp. 


75. Abdomen reddish brown, cogered 
posteriorly with yellowish brown 
tomentum, and yellowish pubes- 
cence. Forehead seven times as 
long as itis broad. Beard white.. pauper, Rondani., 
Abdomen dull reddish brown with 
black pubescence. Forehead seven 
times aslongasitis broad. Beard 
brownish, scanty. Fore coxae with 
white hairs .. leucohirtus, n. sp. 
Abdomen dull reddish Grown with 
black pubescence. Forehead ten 
times as long’ +as’4t-is. broad: 
Beard brown, thick. Fore coxae 
with black hairs ap .. atrohirtus, 1. sp. 


76. Forehead ten times as long as it is 
broad. Frontal callus long and 
narrow. Abdomen reddish yellow, 
darker atapex. Legs black, tibiae 


whitish at base Ke. .. pallidepectoratus , Bigot. 
Forehead six to seven times as long 
as it is broad a a IGE 


77. Forehead narrowest at the vertex, 
abdomenreddish. Legs wholly red nigropictus, Macq. 
Forehead narrowest anteriorly a 78. 


78. Abdomen narrow, reddish brown, 
under side with bands. ‘Thorax 
reddish. Legs black, tibiae white 
at base .. Joidus, Bigot. 
Abdomen conical, reddish vellow, the 
apex black, under side the same, 
Thorax blackish. Legs black, fore 
tibiae dull reddish yellow on basal 
half, others wholly so .. stamensts, 1. Sp. 


79. Forehead five times as long as it is 
broad, narrower anteriorly. Fron- 
tal callus club-shaped. Small red- 
dish brown species. Palpi slender, 

hairy Pi . palpalis, Ricardo. 
Forehead five to six anes as long ¢ as 
it is broad, parallel or nearly so. 


126 


80. 


8I. 


82. 


83. 


84. 


85 


86. 


Records of the Indian Museum. 


Frontal callus oblong or club- 
shaped 

Forehead seven to ten times as long 
as it is broad, narrower anteriorly. 
Frontal callus long and narrow or 
oblong . sits a 

Yellow species. Thorax and scutel- 
lum covered with grey tomentum 
and yellow pubescence. Legs and 
antennae yellow. Palpi stout, end- 
ing in along fine point. Frontal 
callus yellowish, often indistinct. . 

Reddish brown species. Frontal 
callus reddish brown, always dis- 
tinct : che ee 

Thorax and _ scutellum blackish. 
Palpi slender, long. Antennae 
black at tip. Legs “black, tibiae 
reddish yellow 

Thorax and scutellum us asually red- 
dish. Palpi stouter. Antennae 
not black at tip 

Abdomen reddish brown, unicolor- 
ous. Legs reddish yellow, femora 
often darker 

Abdomen marked with grey tomen- 
tum, giving it the appearance of 
having zigzag indistinct bands. 
Legs blackish, tibiae yellowish 

Subcallus not bare, shining or tuber- 
Culous) ye: ; aa 

Subcallus bare, shining, tuber- 
culous 

Palpi stout. Femora black or reddish 
brown : 

Palpi slender. Femora yellowish . . 


' Forehead two-thirds narrower anteri- 


orly than at vertex. Abdomen 
yellow, anteriorly rather trans- 
parent. Legs yellowish, fore legs 
darker ; 

Forehead one-third narrower an- 
teriorly than at vertex. Reddish 
brown or reddish yellow species .. 

Abdomen reddish brown. ‘Thorax 
blackish. Legs black, tibiae yel- 
lowish. Wings clear or ene 
tinged ; 

Thorax reddish yellow, ‘almost same 
colour as abdomen. Legs yellowish, 


flaviventris, Bigot. 


. diversifrons, n. sp. 


obconicus, WIk. 


. borniensis, Ricardo. 


. fuscicauda, Bigot. 


[Vor. IV, 


80. 


83. 


Si: 


82. 


. subcinerascens, n. sp. 


84. 
88. 


85. 
87. 


86. 


Tg1I.] G. Ricardo: Revision of the species of Tabanus. 127 


fore femora darker. Wings tinged 
vellow on fore border 
87. Abdomen dusky reddish brown or 
reddish yellow. Forehead about 
a third narrower anteriorly than 
at vertex, frontal callus pear- 
shaped) <. 
Abdomen olive-colouréd. Forehead 
about half as narrow anteriorly as 
it is broad at vertex, frontal callus 
very narrow ae 
88. Medium sized species. Abdomen 
yellowish with darker apex. Palpi 
vellow. Legs reddish yellow, the 
fore tarsi darker. Forehead seven 
times as long as it is broad 
89. Small species. Legs black, tibiae 
white. Eyes with cross-bands 
go. Reddish brown or yellowish species. 
Beard white or yellow. Palpi 
usually pale. Forehead ten times 
as long as it is broad 
Black, sometimes reddish brown 
species. Beard black. Palpi black- 
ish. Forehead pu times as rae 
as it is broad 4 


Group XI. THERIOPLECTES. 


gt. Forehead with two transverse calli. 

Subcallus bare and shining. Ab- 

domen black with white-haired 

bands. ; 

Forehead with one callus. Sub- 
callus not bare or shining 

g2. Frontal callus black, club- ape 

with spindle-shaped extension. 

Abdomen black, reddish yellow at 

sides of first two or three seg- 

ments. Palpi stout with short 

apex. Appendix usually present. 

Legs black and reddish yellow 

Frontal callus small, indistinct. Ab- 

domen blackish, only reddish on 

second segment. Palpi slender 

with long acute apex. No appen- 

dix to wing ie 2 

93. Smaller species. Legs black and red- 

dish yellow 


. uniformis, n, sp. 


. subhirtus, Ricardo. 


. formostensis, a. sp. 


. tuberculatus, n. sp, 


go. 


. simplissimus, W1k. 


. ceylonicus, Schiner. 


. subcallosus, n. sp. 


. hirtus, Wik. 


93. 


. wyvillet, n. sp. 


128 Records of the Indian Museum. [VoL 11V;,; 


Frontal callus broad, yellow. Abdo- 
men black, reddish yellow at sides. 
Palpi slender. No appendix to 
wing. Legs yellow .. .. albilateralis, Macq. 


Group I. 


Forehead with two callosities, the first one usually transverse, 
the second one smaller, irregular shaped, separated from each 
other, forehead broad, four to six times as long as it is broad. 
Small species g—14mm. Scutellumsame colour as thorax. Abdo- 
men not banded. 


Tabanus rarus, ? , 0. sp. 
(Pl: xiit, fig.-1.) 


Type @ from Kuala Lumpur, Federated Malay States, 
(H. C. Pratt), from London School of Tropical Medicine. 


A small species of unusual appearance, the abdomen being 
covered with grey tomentum and with long white pubescence. 
Subcallus shining. Legs black, all tibiae whitish with fringes of 
long white hairs, which are also present on the femora. Wings 
clear. Length 12 mm. ; 

Face covered with grey tomentum, and with long white pubes- 
cence. Beard white. Palpi yellow with white pubescence which 
is long below and short above. Antennae reddish yellow, the first 
two joints pale yellow with black pubescence, the third narrow 
at base with only an obtuse angle representing the tooth. Sub- 
callus shining, reddish yellow, furrowed in the middle. Forehead 
five times as long as it is broad, very slightly narrower anteriorly, 
covered with greyish yellow tomentum ; the frontal callus reddish 
yellow, reaching eyes, transverse, the median one black, irregu- 
larly heart-shaped, on the vertex appears another reddish brown 
small callus; some black hairs on forehead. Hind part of head 
with long white hairs. Thorax and abdomen black but appearing 
grey, being covered with dense grey tomentum, the thorax with 
whitish pubescence, shoulders reddish with grey tomentum and 
white hairs; breast and sides the same. Scutelium identical with 
thorax, fringed on its outer border with long white hairs. 
Abdomen has the white pubescence thickest on the last five 
segments; under side the same, but the pubescence slighter. Legs 
black, the coxae and femora with some grey tomentum, the fore 
tibiae white, black at their extreme apex, the other tibiae straight, 
slender, yellowish, the long white pubescence is thickest on the 
fore legs, on the tarsi the pubescence is black and short. Wzings 
clear, stigma yellow, veins pale brown. 


Igtt.] G. Ricarpo: Reviston of the species of Tabanus. 129 


Tabanus hirtipalpis, 2 , n. sp. 
(Pixiit, fies: 2225 


Type from Bichiakoh, Nepal; sent in Indian Museum col- 
lection, 


A small black species allied to Tabanus ceylonicus, Schiner, but 
at once distinguished from it by the wide forehead with two callosi- 
ties and the hairy palpi, and by the metatarsi being black. Length 
g mm. 

Face black covered with ashy grey tomentum and with long 
white hairs. Palpfi stout ending in a fine but short point, pale 
yellowish covered with grey tomentum, and with long white hairs 
at base and below, some few biack hairs visible on dorsum. 
Antennae reddish yellow, black at apex, the first two joints with 
black hairs, the third joint rather short with small tooth. Fore- 
head about four times as long as it is broad, slightly narrower 
anteriorly. Subcallus shining yellow, convex, divided by a narrow 
interval from the frontal callus which is black, transverse, not 
reaching the eyes, furrowed laterally ; beyond it appears, separated 
from it however, a small indistinct black callus; the forehead dark 
with brownish tomentum. Thorax and abdomen black, shining, 
the latter with very narrow yellowish red segmentations, some 
white hairs at sides and on under side. Legs black, the fore tibiae 
to apex, and the others wholly, yellowish white, fore coxae and all 
femora with some white hairs. Wangs hyaline, veins and stigma 
brownish yellow. 


Tabanus bicallosus, 7 2, Ricardo. 
(Pil aati fics 4) 


Ann. Mag. Nat. Hist. (8), it1, p. 489 (1909). 


Type (o) from Pusa, Bengal. 
Type (@) and three other females from the same place. 


These specimens were sent to me for identification by 
Mr. F, M. Howlett from India. 


This small species might at first sight be taken for a small 
specimen of Tabanus striatus, Fabr., but is at once distinguished 
from it by the two separate calli of the forehead. It is very nearly 
allied to Tabanus gratus, Loew, which is distributed over South, 
Central, and West Africa, reaching up to Egypt, as I had a speci- 
men sent me from the Suez Canal. The wholly yellow legs, 
narrower median stripe of abdomen the same width throughout (in 
the African species it is wider on the third and fourth segments), 
and the prolongation of the thoracic stripes on to the scutellum, 
seem the only differences between the Indian and the African 
specimens. 


130 Records of the Indian Museum. [Vor IV, 


Black, with five grey stripes on the thorax and three on the 
abdomen. Legs and antennae yellowish. Wings clear. Length 
Io mm. 

9. face covered with grey tomentum and with white pubes- 
cence. Beard white. Palpi greyish, withsome white hairs at base, 
and black hairs elsewhere, but these last are not very numerous. 

Antennae: the first joint yellow, the second and third red, 
tooth at base small. Forehead broad, quite a third narrower an- 
teriorly, and four times as long as it is wide, covered with yellow- 
ish brown tomentum and with some black pubescence; the frontal 
callus yellowish brown, nearly square, and almost reaching the 
eyes ; beyond it and sometimes connected by a very fine line is an 
irregular-shaped black or brownish callus isolated in the middle of 
the forehead. Eyes with three cross-bands. Thorax: the median 
stripe is linear, the other ones broad, sides of thorax greyish, with 
black pubescence. Scutellwm: the lateral stripes are continued 
here, leaving the centre blackish. Abdomen narrow, all three 
stripes reaching from the first to the sixth segment; sides grey, 
with white pubescence; under side covered with grey tomentum. 
Legs uniformly reddish yellow, only the tarsi a little darker ; the 
femora with grey tomentum and white pubescence. Wungs clear, 
the stigma and veins yellow. 

@. The male is identical. Eyes with large facets above ; the 
small ones on the lower third of eyes are continued round the head 
as a very narrow border. 


Group II. 


Forehead with two callosities, the first one usually transverse, 
the second one smaller, oblong, or irregular shaped, separated from 
each other ; forehead broad, four to six times as long as it is broad. 
Small species,9—14mm. Scutellum white or yellow haired. Abdo- 
men with white or yellow-haired bands. ‘Tibiae white, fore tibiae 
curved, and sometimes slightly incrassate. 

The species as yet described in this group are conferred to 
India, Assam, Burma and Federated Malay States. 


Tabanus flavicinctus, 2, n. sp. 


Type 2 from Shillong, Assam, in Howlett collection ; 2 other 
females from N. Khasi Hills, Lower Ranges (Chenneli collection), 
in the British Museum collection; and another female from the 
Nilgiris in Howlett collection. 

A species nearly allied to Tabanus leucocnematus, Bigot, but 
distinguished from it by the presence of narrow yellow-haired 
bands on every segment of abdomen except the first one. The sub- 
callus and palpi are yellowish, the latter with the long-haired pubes- 
cence yellow, the tibiae are yellowish, with the apices of the 
fore pair more widely black, and the white pubescence on them 


ro1r.] G. Ricarpo: Revision of the species of Tabanus. 131 


and the coxae replaced by yellow. ‘The antennae are dark at the 
apices. The abdomen is a lighter reddish brown or reddish yellow. 
The wings are clear, with no appendix. The forehead is almost the 
same width throughout, and a little shorter, hardly more than 
four times as long as it is narrow. Length 9-10 mm. ; 

There is a female in the British Museum collection from N. 
Kanara, India (Bell), which appears to be the same species, but 
the middle callus of forehead is much narrower, and the forehead 
itself slightly narrower and anteriorly narrower. ‘The bands on the 
third and fourth segments are a little produced in the middle, but 
this is the case in some of the other specimens. 


Tabanus leucocnematus, ¢ , Bigot. 
Mem. Soc. Zool. de France, vii, p. 656 (1892). 


Type (now headless) @ , from India, is a faded denuded | speci- 
men which was evidently described by Bigot in this condition ; it 
would be impossible for anyone to verify the species from his des- 
cription in which he makes no mention of the pubescence on scutel- 
lum and on thorax so characteristic of this species and others in the 
group. Fortunately fresh specimens from Lushai Hills, Burma, 
in Dr. Kertesz’s collection and from Khasi Hills, Assam, in Mr. 
Howlett’s collection serve to clearly identify the species, and the 
following description is based on them. 

A small species dark brown in colour with two yellow pubes- 
cent bands on the abdomen, scutellum entirely covered with yellow 
pubescence, all tibiae white, wings tinged brown on the fore border. 
Length I0-IIt mm. 

Face covered with grey tomentum, a dark brown band is 
present on each side of the antennae, pubescence white, consisting of 
long hairs, beard white. Palpi stout, ending in a point, pubes- 
cent, appearing black but covered with dense grey tomentum, on 
the inside they are yellowish, the pubescence is white and long on 
the first joint, on the second short and black, with a few white 
hairs below and on apex. Antennae reddish yellow with a small 
tooth on third joint, the first two joints with black pubescence. 
Subeallus shining, brown or reddish brown, convex, furrowed in 
the middle, produced between the antennae, divided from the frontal 
callus by a narrow interval. Forehead about five times as long 
as it is narrow, and about a third narrower anteriorly, covered 
with grey tomentum and with two separate calli, the frontal callus 
almost square, furrowed in the middle, almost touching the eyes, 
the middle callus distinct from it in fresh specimens, 1s narrower 
and oblong, sometimes with a short linear extension, both blackish 
brown in colour. Thorax dark brown, on the anterior half covered 
with grey tomentum and with yellow appressed pubescence, on 
posterior half with black pubescence bordered with a fringe of 
yellow hairs; sides of thorax with long white hairs ; breast covered 
with grey tomentum, and with long white pubescence. Scutellum 


132 Records of the Indian Museum. [Vou..EVs 


chestnut-brown when denuded, butin fresh specimens covered with 
grey tomentum and with dense golden yellow pubescence of the 
same shade as that on the posterior border of thorax. Abdomen 
reddish brown, appearing darker with dense short black pubes- 
cence, on the third and fourth segments are wide bands of bright 
golden yellow pubescence, which also appears on the sides of the 
first two segments, and often as a narrow fringe on the posterior 
border of the sixth; under side with the yellow bands narrower but 
present on most of the segments, the second segment with white 
pubescence. Legs black, the knees, fore tibiae except at their apices, 
and the middle and posterior tibiae wholly, whitish, with white 
hairs, the fore coxae covered with grey tomentum and with white 
hairs, the others less distinctly so, the pubescence on femora thick, 
black; the fore tibiae are curved but hardly incrassate. Wings 
tinged with brown on the fore border as far as the third or fourth 
longitudinal vein, clearer at the apex, veins, stigma and costal 
border yellowish, short appendix present. 


Tabanus bicinctus, 9, n. sp. 
(Pl. xiii, fig: 52) 


Type @ and another from N. Kanara, S. W. India (Bell) ; 
others from Pahang, Perak and Kuala Lumpur, Federated Malay 
States (Pratt, Durham). 

A handsome black species with white-haired scutellum, white- 
haired bands on the third and fourth segments, whitish tibiae, and 
wings tinged brown on the fore border, distinguished from Tabanus 
leucocnematus, Bigot, by its colouring and by its more heavily 
fringed black femora. Length: type 18 mm., others ro—18 mm. 

Face covered with whitish tomentum and with long white hairs. 
A dark brown band extends between the eyes and the base: of the 
antennae, with dark pubescence. Beard white; thick. Palpz stout, 
ending in a point, yellowish but appearing grey on the outside, 
being thickly covered with grey tomentum and with short black 
pubescence, a few white hairs are visible chiefly at the base, and 
longer ones below. Antennae reddish yellow, the first two joints 
with black pubescence, the third joint with very small tooth crowned 
with a few black hairs, apex of joint darker. Subcallus shining, 
dark brown. Forehead about six times as long as it is broad, about 
one-third narrower anteriorly, covered with grey tomentum, the 
two calli the same colour as the subcallus, the frontal callus 
irregularly square, separated from the subcallus by a narrow inter- 
val, convex, furrowed in the middle, almost reaching the eyes ; the 
median callus is oblong and narrower, on each side of it and at the 
vertex the forehead is brownish. 

Thorax black, covered with an anterior band of grey tomentum 
and short whitish pubescence, and with a heavy white fringe of 
hairs on its posterior border; on the posterior dark part of dorsum 
the pubescence is black and dense. Scutellum covered with whitish 


IglI.] G. RicarDOo: Revtsion of the spectes of Tabanus. 133 


grey tomentum and dense white pubescence. Breast with long 
white pubescence. 

Abdomen black, type with only the two white-haired bands, 
but on some of the other specimens narrow fringes of white hairs 
appear on the fifth and sixth segments, the rest of the dorsum with 
dense black pubescence, sides of first two segments with white 
hairs, black on the others ; under side similar, but the second seg- 
ment is covered with white pubescence. 

Legs black, tibiae white, but black at their apices, fore coxae 
with some long white hairs, and tibiae with white pubescence, else- 
where black, the femora with thick black fringes on each side. 
Wings tinged brown on the fore border as far as the third vein, 
less so on some specimens, veins and stigma brown, costal border 
vellow. 


Tabanus sexcinctus, ¢ , n. sp. 


Type,.a. temale from. Lushai Hills; Burma. (Capt: Ee Cz 
Macleod), sent by Dr. Kertesz for identification. 


A species nearly allied to Tabanus bicinctus, n. sp., but distin- 
guished by white bands on every abdominal segment except the 
first, and the under side is almost entirely covered with white pubes- 
cence. The wings are quite clear. The femora with grey tomen- 
tum, and some white hairs which are long and numerous on the fore 
pair, the thick black fringes not present. The palpi are more white 
haired. The forehead is almost the same width throughout, and a 
little broader, the frontal callus not reaching the eyes. The thorax 
appears less black, with no anterior white-haired band, but with 
some grey tomentum on the dorsum and black pubescence. TLength 
II mm. 


Group ILL 


Species with the callosities on forehead represented by two 
small calli separated from each other, both about the same size, 
usually round, small spots; distinguished by their Azylotus-like 
appearance (fu/vus group only) though the pubescence on eyes is 
often practically non-existent. So far the only species in this group 
from the Oriental Region are also common to the Palaearctic 
Region, and are represented by the typical Tabanus ditaeniatus, 
Macq., a South African species whose distribution is very wide, and 
Tabanus fulvus, Mg. (Atylotus). 

The identity of Tabanus agricola, Wied., recorded by Schiner as 
occurring at Hongkong, seems very doubtful; the species is not 
included in the table. 


Tabanus fulvus, 7 ¢? (Afylotus), Meigen. 


Syst. Beschreib., ii, 61, 40 (1820) (see Kertesz Cat. Dipt. iii, 
1908, for authorities). 


134 Records of the Indian Museum. [VOns IVE 


Atylotus bituberculatus, Bigot, Mem. Soc. Zool. de France, v, 
659 (1892). 

Bigot gives China with a query as the locality for his type, 
which is identical with Meigen’s species. 


In British Museum collection are two females from Chinkiang 
Yangtze River, China (B: G. Tours); and another from China is 
in the Kertesz collection, so that the range of the species to the 
east coast of China and within the line of the Oriental Region 
is well established. 

The type of Tabanus hoang, 2? , Macq., Dipt. exot., Suppl. v, 
p. 46 (1855), is apparently lost as, though the label is existent 
among the Bigot types lent me by Mr. Verrall, there are no speci- 
mens which answer to the description given by Macquart, in which 
he says it is related to Tabanus fulvus, Mg.; it was possibly identical 
with this latter, judging from the description. In any event it 
may now be deleted from the list of Tabanus. 


Tabanus ditaeniatus, 7 9? Macquart. 


Dipt,-exot., 1,.(2),.Ps5304 (1838). -Walker,) Lists. Dipt.; vs 
pp: 236,. 355, (1854); Bezzix Ann; Mus. Civ.. Genovas (2), xii 
(xxxii), p. 184 (1892) ; Austen, Report Gordon Memorial College 
(1906), p. 57, fig. 22; td., African Blood-Sucking Flies, p. £18, 
pl. xi, fig. 82 (1909). 


Tabanus pyrrhus, Walker, Dipt. Saund., i, p. 47, pl. i, 
figs. 4, 3 (1850). | 

Tabanus bipunctatus, v. d. Wulp, Notes Leyden Museum, 
vii, p. 75, pl. v, fig. 5 (1885). 

Atvlotus nigromaculatus, Ricardo, Ann. Mag. Nat. Hist. (7), 
vi, p- 165 (1900); Newstead, Ann. Tropical Medicine, 1907, 
Pp. 44. 


The type was recorded ;by Macquart from Mauritius; it is 
a common species in South Africa; Coquiilet records. it from 
Japan as Tabanus...pyrrhus,—see Proc. U. States Mus., xxi, 
p. 311 (1898); as may be seen from the localities given below, 
it has a-wide distribution ranging from Africa to India, China 
and Japan. 

In Indian Museum coll. and Howlett coll. are specimens 
from Perso-Baluchistan Frontier; Pusa; Belgatchia, Calcutta, in 
Bengal; Madras Coast; Umballa, N. W. India; Balighai in 
Orissa, S. E. Bengal. In Brit. Mus. coll. specimens from Wei 
Hai Wei, China ; “ biting cows”’ (Muat). 

A greyish yellow species, the abdomen yellowish with a black- 
ish brown broad median, and narrow lateral stripes, on the median 
one appear greyish yellow tomentose triangular spots, the lateral 
stripes do not reach the sides of abdomen and ‘at apex coalesce 
with the median stripe; the abdomen often varies in appearance 


Igii.] G. Ricardo: Revision of the species of Tabanus. 135 


the yellowish ground colour not always being very apparent. 
Antennae and legs yellowish. Length 11—14 mm. 


‘Tabanus agricola, 9, Wied. 


Auss. Zweifl. Ins., i, p. 556 (1828); Schiner, Reise Novara 
Dipt., p. 82 (1868). 

Ashy grey: sides and segmentations of abdomen, antennae 
and legs luteous: Jength 6 lines, @.. From Red Sea. 


It is somewhat similar to Tabanus rusticus, Fabr.. Antennae 
yellow (the last joint wanting). Palpi very pale yellow. Beard 
yellowish white. Face and forehead mouldy grey. Thorax 
appears as if covered with ashy grey tomentum because it has 
whitish pubescence on the black-grey ground colour. Abdomen 
clay-yeilow on the sides, and more narrowly so on the hind borders 
of the segments, so that not much remains of the black-grey 
ground colour on each segment. Under side almost yellow. 
Dorsum and under side with white pubescence. Wings colourless 
with yellow veins. Legs clay-yellow; the anterior tibiae at the 
base inside, all tarsi, but the posterior tarsi only at the apex, 
brownish black. (Rtippel in the Frankfort Museum.) Wied., Auss. 
Zweifl. Ins., i, p. 556. 

Wiedemann’s description fits a male and two females from 
Hongkong so well that I am induced to refer them to this species, 
and add the following particulars to complete the description :— 
The antennae are pale red-yellow, the first and second joint short, 
not produced above and with few hairs, the third somewhat deeper 
in colour, slightly incised above, so that the angle projects but 
little. The femora have tolerably thick but extremely short 
whitish pubescence ; the fore tibiae brownish black at the apex 
(Wiedemann says, no doubt by mistake, brownish black at the 
base, which is contrary to the usual colouring of species of 
Tabanus). Wings quite clear, the first posterior cell wide open, an 
appendix present. Eyes of male very large and convex: they 
join completely on the forehead. Everything else as described by 
Wiedemann ; the correctness of the identification is chiefly corro- 
borated by the allusion to the similarity of the species with 
Tabanus rusticus, Fabr. (Schiner, Reise Novara Dipt., p. 82). 

From the comparison with Tabanus rusticus, 1,., nec Fabr. 
(Atylotus), itis to be presumed this species has two small calli on 
forehead. Tabanus rusticus has not yet been recorded from Asia. 
It seems probable that the specimens Schiner identified from 
Hongkong as Tabanus agricola, Wied., belonged to Tabanus ditae- 
niatus, Macq. 


Group IV. 


A small group of species, at once distinguished by the absence 
of callosities on the forehead, which'is broad, from four to five times 


136 Records of the Indian Museum. [Vou. IV, 


as long as it is wide. In old or denuded specimens it occasionally 
appears as if a frontal callus were present. Species from 74 to 13 
mm. in length. 


Tabanus erythrocephalus, Wulp, from Halmaheira Island, and 
Tabanus leucopterus, Wulp, from Aru Island, in the Australian 
Region, appear to belong to this group. 


Tabanus virgo, 2 o, Wiedemann. 


Anal. Entomol., 22 (1824); 7d., Auss. Zweifl. Ins., i, p. 184 
(1828). 


Tabanus albulus, 9 @, Walker, Dipt. Saund., i, p. 46 (1850). 
Tabanus puella, 7, Walker, l.c., p. 53. 


Types of Tabanus albulus, male and female, from India 
(Saunders coll.). 


Type of Tabanus puella, male, from India (Saunders coll.). A 
male from Mysore (H. K. Slater). In Indian Museum coll. Speci- 
mens from Calcutta and Pusa, Bengal. The Wiedemann type 
came from East India. Slaty grey with red-yellow antennae and a 
twofold white spotted abdomen. Length 4—4} lines, 7 9°. 


Antennae bright rusty yellow; face rather grey haired. Palpi 
very light yellow. Beard white. Forehead light yellowish green 
without a callus or line. In the male it is ochre-yellow behind the 
antennae. Thorax slaty grey, very faintly white striped, the 
median line extremely narrow. The shoulders and posterior sides 
of thorax almost chamois-leather colour. Breast sides whitish. 
Abdomen slaty grey with narrow yellowish white-haired segmenta- 
tions: on each segment two white-haired roundish spots. Wings 
very hyaline with ochre-yellow veins and halteres. Legs ochre- 
yellow with almost chamois-leather coloured femora. In the 
Royal Museum. Wiedemann, Auss. Zweifl. Ins., i, p. 184. 

This small species is easily recognized by the unmarked fore- 
head and very pale wings, yellow antennae and legs and spotted ab- 
domen. The female specimens above mentioned vary from 9—II 
mm. One male only measures 74 mm. 

The description given by Wiedemann of the colour of the 
abdomen only applies to very fresh specimens, the grey colour 
being caused by the grey tomentum covering the abdomen, viewed 
best from behind ; the ground colour of the abdomen is greyish 
brown, rusty coloured at the sides of the first three or four seg- 
ments, the pubescence black, but white onthe spots and seg- 
mentations ; under side lighter with white pubescence, darker at 
the apex. The thorax is blackish brown covered with grey tomen- 
tum and with some appressed fulvous hairs, the stripes hardly 
visible. The forehead is about five times as long as it is wide, and 
a third narrower anteriorly, yellowish, showing no callus when 
fresh , if denuded one is sometimes visible. Antennae reddish yel- 
low, the first two joints pale yellow. Face ashy grey with white 


rgiI.] G. RIcARDO: Reviston of the spectes of Tabanus. 137 


pubescence. Beard white. The palpi very pale yellow, stout at 
base, ending in a point. 

The male has the abdomen more reddish in colour, darkening 
at the apex. Large facets of the eyes very distinctly divided from 
the small ones, which take up barely a third of the eye and reach 
the vertex as a very narrow border. 


Tabanus nemocallosus, 2? , Ricardo. 
(RIS xin figs6,) 
Ann. Mag. Nat. Hist. (8), ili, p. 4g0 (1909). 


Type @ and another female from Pusa, Bengal, in Howlett 
coll. 


A medium-sized species, distinguished by the rather broad 
forehead, not narrower anteriorly, and with no callus or spots. 
Abdomen blackish brown marked with greyish yellow median 
and lateral spots. Length 13 mm. 

Face and forehead covered with greyish tomentum, the former 
with white pubescence. Beard scanty, white. Palpi yellowish 
white with white pubescence, stout ending in a short point. 

Antennae reddish yellow, the first two joints pale yellow, the 
third with hardly any tooth. J orehead barely three times as long 
as itis broad. Thorax, scutellum and abdomen blackish brown, the 
former with five distinct greyish yellow stripes, the dorsum with 
some grey tomentum and with appressed pale fulvous pubescence ; 
scutellum with the same and with grey tomentum. Abdomen with 
an almost continuous median stripe tormed of grey tomentose 
narrow, triangular spots, and with irregular-shaped, roundish, grey 
tomentose spots on the sides, the pubescence on these, short, pale 
fulvous, some black pubescence on the dark parts, the sides of the 
first two or three segments pale reddish, under side lighter, covered 
with grey tomentum. Legs pale yellowish red, the tarsi brown, 
the pubescence white on femora and tibiae, black on the tarsi. 
Wings hyaline with brown veins and yellow stigma. a long 
appendix on branch of third vein. 


Tabanus negativus, ” ?,n. sp. 
Type ~, type 2, from Formosa, in Dr. Kertesz’s coll. 


In Brit. Mus. coll. one female from Hongkong very much 
mutilated. 


A species allied to Tabanus nemocallosus, Ricardo, but distin- 
guished from it by the absence of spots on the abdomen and of 
stripes on the thorax. Wings with no appendix, veins pale 
yellow. Length of female type 13 mm.; of male specimens 114-12} 
mm. 

9. Face, beard, and antennae the same as in the above- 
mentioned species. Palpi pale yellow, the same shape, but with 


138 Records of the Indian Museum. [VoL. IV, 


black pubescence. forehead with no callus; almost parallel, about 
four times as long as itis broad. Thorax and abdomen uniformly 
covered with greyish yellow tomentum, through which the dark 
ground colour hardly appears, and with thick short pale yellow 
pubescence on the dorsum of abdomen, on the thorax it is less 
thick and replaced chiefly by black pubescence anteriorly, shoulders 
the same colour. Breast rather greyer with white pubescence. 
Under side of abdomen similar but almost devoid of pubescence. 
Legs pale reddish yellow, the apices of fore tibiae and the tarsi 
blackish, the other tarsi more or less brown, pubescence pale 
yellow on the coxae and femora, chiefly black elsewhere. Wings 
clear, the veins very pale yellow and stigma hardly noticeable, 
no appendix. 

@. Similar ‘Ge in some of the specimens, owing probably to 
denudation, black median markings or a black median stripe appear. 
Abdomen with very little yellow pubescence, almost bare. Eyes 
large, convex, the large facets occupying fully two-thirds of the 
surface of eyes, the small ones reaching to the vertex as a narrow 
border. Palipi pale yellow with yellow pubescence and black 
pubescence on apex. Frontal triangle and subcallus coloured 


alike. 


GROUP V. 


Species with wings spotted or banded. Forehead with one 
square, narrow, or oblong callus, always prolonged towards the 
vertex as a more or less narrow line; five to ten times as long as 
itis broad. Species from 9—2I mm. in length. 


Tabanus javanus, 2, Fabr. 


Syst. Antl., 103, 46 (1805); Wied., Dipt. exot., p. 93 (1821); 
id., Auss. Zweifl. Ins., i, p. 183 (1828). 


Thorax striped; abdomen with white triangular dorsal spots, 
wings spotted. Smaller than the preceding species (Tabanus 
vittatus and longicorms). Head white haired. Thorax brown, 
with white stripes. Abdomen brown, with a large triangular spot 
on the dorsum of each segment. Body pale, darker at the apex. 
Wings hyaline, with five black spots. Legs testaceous. From 
Java, Pabr., Syst. Antl.; p. 103. 

The species is unknown to me. Wiedemann redescribed it in 
Auss. Zweifl. Ins., i, p. 183, as follows: Thorax black, white 
striped, the sides rusty yellow, abdomen brown, ferruginous at 
the sides, with triangular white median spots; wings spotted with 
brown, 44 lines ¢ , from Java. 

The whole head reddish white. Beard white. Forehead with 
a chestnut-brown smooth stripe, widening below into a broad callus. 
Dorsum’ of thorax with three whitish lines, the sides almost 


rgit.]| \G. Ricarpo: Revision of the species of Tabanus. 139 


chamois-leather coloured behind the usual. seam, with a small 
black stripe; breast sides almost  chamois-leather coloured ; 
scutellum black. Abdomen on the first segment brown, 
rusty. brown on the sides, the second one at the base with 
an interrupted whitish band and another more wavy one at the 
apex; from the middle of it the triangular spot proceeds, the third 
to the sixth each with the same shaped whitish band at the apex 
and larger triangular spots; on the seventh the border is hardly 
whitish ; the three last’ segments are not at all rusty yellowish ; 
under side almost chamois-leather coloured, brownish towards the 
apex, with whitish segmentations. Wings almost hyaline, stigma 
and four spots brown. Legs reddish; tarsi gradually brown. ‘The 
fore legs are wantingin the single specimen in Fabricius’s 
collection. A very denuded specimen in the Leyden: Museum 
is of a deep pure brown colour, with no trace of the stripe and 
spots of abdomen, only the segmentations are yellowish white. 


Tabanus optatus, 2 , Walker. 


Proc. Linn. Soc., i, p. III, 33 (1856). 


Tabanus alboscutatus, 2, Rondani, Ann. Mus. Civ. Genova 
vii, p. 456 (1875). 


Tabanus equestris, 7 @, v. d. Wulp, Notes Leyden Museum, 
Wil P77, 34, Diav, 112. 0/1885). 


Walker’s type, 2, came from Sarawak, Borneo (Saunders 
coll.); another female in Brit. Mus. coll. from Kajoe-Tanan, 
Sumatra (Dr. Hagen), and a male from India (Saunders coll.). 
In Indian Museum coll. a female from Belgatchia, Bengal. 

On comparison of one of the Brit. Mus. coll. specimens with 
the type of Tabanus alboscutatus, the identity of the two species 
was at once established. Rondani’s type also came from Borneo. 

From van der Wulp’s description and figure of wing there 
is no doubt he was describing the same species, and his description, 
as being the best one, is given below. 

Reddish, face and palpi pale yellow; eyes naked ; antennae 
with the third joint darker; abdomen with triangular dorsal white 
spots; the anterior legs and all femora brownish; wings at the 
base and a median band brownish, 7, @. Length 15°5—17 mm. 

Face yellowish white; beard of the same colour ; palpi pale 
ochraceous ; front rufous, in the 9 moderately broad, with a 
brown cordiform spot extending upwards ina line. Eyes bare, 
in the @ reddish bronze above, with large facets ; below for a 
third part and on the outward borders blackish, with much smaller 
facets, even after having been moistened no trace of cross-bands is 
to be seen. Antennae ochraceous ; third joint infuscated, rather 
slender and on its upper side with a triangular tooth. ‘Thorax, 
scutellum and abdomen rufous, the thoracic dorsum a little infus- 
cated, with indistinct stripes; hind margin of the scutellum 


140 Records of the Indian Museum. [VOLEEV; 


greyish ; last segments of the abdomen darker ; their hind margins 
yellowish, with small triangular, white, dorsal spots; on the under 
surface the pale margins of the segments are broader. Legs and 
halteres fulvous; the femora and the whole front legs fuscous. 
Wings at the base brownish yellow, in the middle with a rather 
broad brownish cross-band, which issues on the stigma and em- 
braces the discal cell, but does not quite reach the hind margin ; 
the costal cell is yellow. This species seems to be not uncommon 
in the Sunda Islands ; it is represented in the Leyden Museum by a 
¢ from Sumatra (v. Lansberge), 2 2 from Java (Blume and Ludeking) 
and 4 ¢ from Borneo (Muller) ; it was along time ago designated in the 
collection by the late Mr. Snellen van Vollenhoven under the 
name equestvis,n.sp., without having been described. The speciesis 
related to my species Tabanus pictipennts (Tijd. v. Ent., xi, p. 100, 
1868), the name of which having been used before, is changed by 
Osten Sacken into Tabanus vanderwulpi (Berlin Ent. Zeit., xxvi, 
p. 97, 1882); it differs, however, by its fulvous posterior legs, its 
more slender antennae, the third joint of which is darker, and by 
the pattern of the wings, showing instead of two cross-bands, 
but a single one which passes over the discal cell. Notes Levden 
Museum, vii, p. 77, 31, pl: v, fig. 6, 1885. 

Tabanus pictipennis is the same as Tabanus flexilis, Walker, a 
species from Celebes, but not the same as Tabanus vandcrwulpr. 
The forehead against the general rule is slightly wider anteriorly, 
about a third, and is five and a half times as long as it is wide, the 
frontal callus small, pear-shaped, with a linear extension. The 
legs have the fore tibiae obscurely reddish yellow on their basal half, 
the extreme apices of other tibiae and all tarsi blackish. The cross- 
band of wing in the description, said not to reach the hind border 
of discal cell, does so in the figure and in these specimens. 


Tabanus non-optatus, @ , n. sp. 


Type o from Purneah District, in Indian Museum coll. 


Type @ from unknown locality and two males in Brit. Mus. 
coll. from unknown locality. 


This species is very nearly allied to Tabanus optatus, Walker. 
The female type is so similar to the male type, and the two males 
identical with latter, that I think it is safe to assume that the speci- 
mens with no locality specified came from India or an adjacent 
region, especially as the species with banded wings are not very 
numerous in the African or Oriental Region, and probably not in 
any fauna. 

This species is distinguished from the Walker species by the 
almost wholly yellowish abdomen, only becoming darker on the 
last two or three segments, and the pale spots are larger, more 
triangular in shape, the abdomen is longer and more conical in 
shape, and the forehead in the female is not narrower at the 


1giI.] G. Ricarpo: Revision of the species of Tabanus. I4I 


vertex. Length of male type 14 mm., of female type 19 mm., 
the other males 17—19 mm. 


9. Face covered with greyish tomentum, pubescence and 
beard whitish. Palpi yellow with longer slender point, and a few 
black hairs. Antennae reddish yellow, the first two joints with some 
black hairs, the third broad at base, deeply incised, with a distinct 
tooth. Subcallus and forehead same colour as face, the latter with 
some black pubescence, almost parallel, about five times as long 
as it is broad; the frontal callus reddish brown, almost square, not 
reaching the eyes, posteriorly narrower with a short lineal extension. 
Thorax dull chamois-leather coloured, with some greyish tomentum, 
no stripes visible, a few yellowish and black hairs visible on dorsum, 
shoulders pale reddish with black pubescence; breast covered with 
grey tomentum and with greyish white pubescence, tufts of white 
hairs round base of wing; scutellum same colour as thorax, but 
with whitish grey tomentum on its posterior border. Abdomen 
reddish yellow, black at apex, the spots most distinct in the second, 
third and fourth segments, pubescence black but very scanty and 
short, the segmentations somewhat lighter ; under side similar with 
more distinct white segmentations. Legs blackish, the tibiae reddish 
with black pubescence, the pubescence on femora white and on 
coxae, which latter are covered with grey tomentum, tarsi reddish 
brown with black pubescence. Wings with a dark band across the 
middle of wing, similar to that of Tabanus optatus. 


@. Similar, the thorax rather darker in colour. The eyes 
with large facets on upper part, the small ones attaining the vertex 
as a narrow border. 


Tabanus flexilis, 7 @ , Walker. 


Proc. Lirfh. Soc., iv, p. 104 (1860) ; v, p. 104 (1861). 


Tabanus pictipenms, 2, v.d. Wulp, Tijd. v. Entom., xi 
p. 100, pl. iti, figs. 1, 2 (1868); nec vanderwulpi, Osten Sacken. 


Tvpe o@ from Celebes. 


b] 


Type @ from Macassar, Celebes (A. R. Wallace), and others 
from Manado, Celebes, all in Brit. Mus. coll. 


This species, though as yet only recorded from Celebes, is 
included here owing to the great similarity to it of Tabanus 
vanderwulpi, Osten Sacken, from the Philippines. 

From the figure and the description of van der Wulp’s there is 
no doubt he was redescribing a specimen identical with Walker’s 
species, both from the same locality. His description is given, as it 
is sufficient for identification, of the female only however, as he did 
not know the male. 


9. Ochraceous. Forehead very narrow witha blackish median 
line; eyes naked; antennae testaceous; the anterior margins of 
the abdominal segments brown; legs blackish; wings with two 


142 Records of the Indian Museum. [VOL LV 


brown bands; costal border testaceous. 9 8? lines. PI. iii, figs. 1 
and 2. 


Head ochre-yellow, below with some coloured pubescence; the 
forehead very narrow, stripe shaped, with a black median line 
hardly broader below. Eyes naked, bronze coloured, without 
markings. Antennae cinnamon-brown, the first joint ending in a 
point, above the second very short, prolonged above in a thorn-like 
point, the third moderately incised above. Palpi’ ochre-yellow. 
Thorax and scutellum blackish with ochre-yellow tomentum and 
on the breast sides with conspicuous ochre-vellow pubescence. 
Abdomen black-brown, with thick pale ochre-yellow almost golden 
yellow tomentum, which leaves the fore borders of the segments 
tolerably wide, so that cross-bands more or less darker are formed, 
somewhat broader at the sides. Legs black, the coxae the same 
colour as the abdometi, and the ground colour of the femora 
becomes more or less modified owing to the rather thick ochre- 
yellow pubescence. Halteres yellow. Wings with greyish ground 
colour and black-brown veins; the base and the marginal cell 
brown-yellow; a brown shadow lies across the transverse veins, 
which forms a narrow half zigzag band in the middle of the wing, 
and on the jast third of the wing appears a broad brown cross- 
band, which stretches from the fore border at the ending of the 
first vein to the upper branch of the third longitudinal vein; this 
band, which becomes paler on the posterior border, is rounded on 
the inside and outside towards the apex of wing notched triangu- 
larly, the angle of the third vein has no appendix; the first 
posterior cell is closed at the border with a short stalk. Celebes 
(Rosenberg). v.d. Wulp, Tijd. v. Entom., xi, p. 100. 


In Walker’s type the abdomen is brownish black, but the 
yellow bands are covered with yellow pubescence, besides tomen- 
tum ; the under side is more yellow with whitish segmentations, 
blackish at the apex. Legs black, the fore tibiae at the base 
obscurely yellowish, the hind tibiae obscurely reddish yellow, 
the pubescence on coxae and femora yellow with grey tomentum, 
black on the tibiae and tarsi, the hind tibiae with black fringes. 
Forehead is barely more than half as wide anteriorly as it is at 
vertex and about ten times as long as it is broad, the frontal! 
callus very narrow, pear-shaped, with lineal extension. Length of 
female type 19 mm., the other 16 mm. 


@”. Eyes with equal facets. Face and subcallus with yellow- 
ish grey tomentum and yellowish white hairs. Beard the same. 
Palpi yellow with yellow hairs and a few black ones at the apex. 
Antennae reddish yellow, the first two joints pale yellow, with black 
hairs, the third long and slender with a small tooth, slightly dusky 
at tip. Abdomen more yellow than in female, the darker bands only 
beginning on the fifth segment, the segmentations narrowly whit- 
ish, the under side identical. Wangs with the first posterior 
cell narrowed at opening. Length of type 16mm., of others 
Ig mm. 


Igtr.]| G. Ricarpo: Revision of the species of ‘Tabanus. 143 


Tabanus vanderwulpi, 2 o, Osten Sacken. 


Berlin. Ent. Zeit., xxvi, p. 97 (1882); nec pictipennis, V. d. 
Wulp. 


One female in Brit. Mus. coll. from the Philippines. 


This species is very nearly related to Tabanus flexilis, W1k. 
Osten Sacken thought it identical with Tabanus pictipennis, v. d. 
Wulp (which is undoubtedly the same as the Walker species), and 
renamed it as above, the first name being preoccupied. His re- 
production of van der Wulp’s description is not given here as he 
made additions to it to suit his specimens from the Philippines, 
I v and4 2. Judging from the specimen in the Brit. Mus. coll. 
the female differs from Tabanus flexilis, Wlk., in the following 
particulars :— 

Antennae black at apex. Forehead about half as wide an- 
teriorly as it is at the vertex and fully ten times longer than it is 
broad, the frontal callus longer and narrower, very little wider 
than its lineal extension. Abdomen brownish red with the yellow 
posterior bands narrower, produced inthe middle as triangular 
spots, but the under side is darker with black and yellow bands. 
Legs black or dark brown with black pile as Osten Sacken des- 
cribes, but in this specimen the femora have some yellow hairs, 
the fore coxae same colour as breast. Length 21 mm. 

@. According to Osten Sacken’s description the eyes have 
large and small facets very sharply divided. 


Group VI. 


Forehead with one square or narrow or oblong callus, always 
prolonged towards the vertex as a more or less narrow line, usually 
nine to ten times as long as it is broad (in Tabanus auriflamma 
Wlk., however, it is much broader). Large species, 18-23 mm. in 
length, with the wings coloured brown or yellow or both, usually 
clear at extreme apex and sometimes in the discal cell, never 
forming regular bands. 

Tabanus speculum, Wik., trom Celebes will belong to this 
group. 


Tabanus pratti, 2, n. sp. 


d 


Type @ in Brit. Mus. coll. from Kelanton, Malacca (pur- 
chased from H. Rolle, 1904); and another female from Sarawak 
(A. R. Willington, 1908). Presented London School ‘Tropical 
Medicine coll. In the latter collection is one female from Batu 
Tiga, Selangor (Pratt). This very handsome black species is 
distinguished by the bright orange-red pubescence on face and 
sides of breast ; the beard is also the same colour. Abdomen black 
with white-haired bands at sides. Wings dark brown. ‘Tibiae white 
at base. Length of type 24 mm., the other female 23 mm. 


144 Records of the Indian Museum. [VOE..LV. 


Face thickly covered with bright orange-red pubescence, the 
hairs in middle of face and a tuft at sides rather more golden 
yellow, a few brown hairs are visible on cheeks bordering the eyes, 
and a tuft-like fringe between the antennae and the eyes. Beard 
orange-red, thick. Palfi dull reddish, appearing black by reason 
of the thick black pubescence, stout, almost the same width 
throughout. Antennae red, the first two joints covered with black 
hairs, the third large, broad, the last four divisions forming hardly 
a fourth of the whole iength; the tooth is represented only by a 
very slight angie indeed. Subcailus blackish, covered with reddish 
brown tomentum. fovehead darker, narrow, about nine times as 
long as it is broad, and a third narrower anteriorly; frontal callus 
black, shining, long and narrow, almost reaching the eyes with a long 
stout linear extension. Thorax, scutellum and abdomen dull black- 
ish brown, covered with close black pubescence, on the shoulders 
and sides of thorax it is longer ; breast with dense orange-red long 
pubescence ; in the centre it is nearly bare, covered with reddish 
brown tomentum. Abdomen with white-haired segmentations on all! 
the segments but the first, thickest at the sides, hardly ever reach- 
ing the centre but usually represented on the median line by a few 
scattered white hairs ; the type itself is denuded and only shows the 
white bands on the second and third, the segmentations under the 
white pubescence at sides only, appear golden yellow, sides of ab- 
domen with thick tuft-like black pubescence ; under side black with 
white-haired segmentations. Legs black with black pubescence, the 
tibiae white on the basal half with white pubescence, but black at 
the extreme base. Wungs a dark rich brown, a clear streak is visible 
in the costal cell below stigma, another in the marginal cell, the discal 
cell is whollv clear, the extreme apex, the basal cells and posterior 
border much paler if not quite clear, veins and stigma blackish 
brown, first posterior cell a little narrowed at opening. Squamae 
brown. Halteres brown, the club at apex yellow. It is named 
after the gentleman who has enriched the London School of 
Tropical Medicine and Brit. Mus. coll. with a considerable number 
of specimens from the Malay Archipelago. 


Tabanus fuscicornis, 2 , n. sp. 


Type ¢? in Brit. Mus. coll. from Punkio, Formosa (A. E. 
Wileman, 19090). 

This species is nearly allied to Tabanus pratti, n. sp., in 
colouring, but is at once distinguished by the black antennae and 
black palpi and by the more hyaline wings, quite clear at apex. 
Length 17 mm. 

Face brownish yellow, paler at base, with a few black hairs, 
the bright orange-red pubescence is here confined to the beard. 
Palpi blackish with black pubescence more pointed than in 
I. pratt. Antennae wholly black, the first two joints with 
black pubescence, the third broad at base with a perceptible 


1gtI.] G. Ricarpo: Revision of the species of Tabanus. 145 


tooth. Subcallus brown. Forehead ashy grey, narrow, the 
same width throughout and about seven times as long as it is 
broad, the frontal callus black, almost the same width through- 
out, to within a short distance of the vertex, very similar to 
that of T. pratti. Thorax and scutellum in this species bluish 
black, with ashy grey tomentum, devoid of pubescence on dor- 
sum, sides with long black hairs; breast with similar orange-red 
pubescence to that of T. pratti. Abdomen black with median 
white-haired spots, the segmentations white-haired at the sides 
only, pubescence on dorsum black; under side black with white 
segmentations. Legs black with black pubescence, some yellow 
hairs on the fore coxae and yellow tomentum on the others. 
Wings marked with dark rich brown colouring, most intense on 
fore border and across middle of wing, leaving the discal cell how- 
ever clear, and not extending beyond the base of fork of third 
vein, so that the apex is wholly clear, posterior border almost 
wholly clear, veins and stigma blackish, no appendix present. 
Halteres brown. 


Tabanus nephodes, 2? , Bigot. 
Mem. Soc. Zool. France, v, p. 656 (1892). [Atvlotus.] 


Type (female) recorded from India; a label is affixed to it with 
“‘Naga Hills,’ which are on the Burma-Assam boundary. There 
is another female in the Indian Mus. coll. from Sibsagar, Assam, so 
that the habitat of this species is probably in these regions. 

A dark reddish brown species with indistinct grey tomentose 
bands and median spots on the abdomen. Legs reddish brown. 
Wings deeply tinged yellowish brown, with the first posterior cell 
closed. 

Antennae yellow. Forehead very narrow. Length of type 21 
mm., the other female is Ig mm. 

Face red, covered with grey tomentum, above near the 
antennae and the subcallus with more yellowish brown tomentum, 
the pubescence brown, on the lower part of the face the pubescence 
is white. Beard of long white hairs. Palpi yellow with black 
pubescence, long, slender, not much stouter at base, ending in an 
obtuse point. Avntennae reddish yellow, the first two joints with 
black pubescence, the third blackish at its apex with a few hairs, at 
the base broad, with a small tooth crowned with some black hairs. 
Forehead reddish, covered with grey tomentum, about ten times as 
long as it is broad, almost reduced to half its width anteriorly, the 
frontal callus reddish, very narrow, cylindrical, with long, raised 
lineal extension ; the callus is hardly wider than this last. Thorax 
and abdomen reddish brown, the former with grey tomentum, sides 
reddish yellow, shoulders with black hairs; breast blackish, covered 
with greyish white tomentum, pubescence probably white. Scwtel- 
fum similar to the thorax. Abdomen with grey tomentose bands on 
the first four segments about a third of the width of segment ; on the 


146 Records of the Indian Museum. [VOL Lvs 


last three the segmentations are very narrowly lighter, hardly per- 
ceptible; there are traces of median white-haired spots with a 
ground-work of grey tomentum on the second to fifth segments ; 
pubescence otherwise seems black, but on the sides of the first 
four segments there are white hairs; under side similar with white 
hairs on the segmentations. All the specimens are more or less 
denuded. Legs reddish brown, the fore tibiae at extreme base and 
the middle pair appear reddish yellow, fore coxae with grey 
tomentum and some white hairs, femora also with a few, other- 
wise the pubescence is black. Wings with almost a dark band, 
the dark brown colouring extending from the upper end of the 
stigma to the apex on fore border, but not reaching much 
beyond the branch of third vein, leaving the apex clear, extending 
across the wing towards posterior border over the upper border of 
discal cell, shading the veins proceeding from the cell and graduaily 
fading away ; costal border and stigma yellow, base of wing with a 
faint yellow tinge, veins yellow, brown on upper half, the first pos- 
terior cell closed, appendix present. 


Tabanus albofasciatus, ?, n. sp. 


Type ° from Shillong, Assam, sent in Indian Mus. coll. 


A large brown species, distinguished by the pale greyish basal 
band on scutellum, in this respect allied to Tabanus albocostatus, 
Bigot, but with the posterior cell very much narrowed at its open- 
ing on border. The abdomen has narrow white tomentose bands. 
Wings yellowish on basal half, brown on apical half. Legs black. 
Length 21 mm. 

Face covered with yellowish tomentum and with some pale 
yellowish hairs. Beard yellow. Palpi yellow with black pubes- 
cence, large, ending in a fairly acute point, antennae bright red, 
dusky at apex, the first two joints pale yellow with black pubes- 
cence, the third wide at the base with a prominent tooth. Fore- 
head with tomentum a little darker than that of the face, about 
nine times as long as it is broad and barely half as wide anteriorly 
as it is at vertex. Frontal callus long and narrow with lineal ex- 
tension. Thorax brown, reddish yellow at the sides with golden 
yellow hairs at base of wings, black hairs on the shoulders, dorsum 
of thorax with traces of short golden yellow pubescence anteriorly ; 
breast covered with greyish yellow tomentum and with yellow 
pubescence, some black hairs intermixed. Scutellum brown but 
the basal two-thirds covered with grey tomentum. Abdomen 
blackish brown with greyish white narrow tomentose bands on the 
first five segments, the first one does not extend beyond the sides, 
traces of white pubescence on them, otherwise the pubescence on 
dorsum where present is black ; under side identical. Legs blackish 
brown, the femora with some yellowish brown tomentum below, 
pubescence black. Wings yellowish as far as the apex of discal 
cell, more intensely so on fore border, apex brown extending to 


Igit.] G. Ricarpo: Revision of the species of Tabanus. 147 


the fourth posterior cell but the second submarginal cell is almost 
wholly clear. The type is not in very good preservation but is too 
distinct a species to be confused with others, though it is allied to 
Tabanus jotdus, Bigot, in colour of wings and shape of antennae 
and forehead; the palpi are however larger and broader, and the 
wings have no appendix, the posterior cell open at border though 
very narrow. 


Tabanus basalis, 2? , Macquart. 


Dipt=-exot..5 £. (1); ‘p.230°.(1838) evan, der Waulp,: Notes 
Leyden Museum, vii, p. 72, 26 (1885). 


Black. Scutellum and basal segments of the abdomen ferru- 
ginous. Wings yellow at the base, brown at the apex. Length 8 
lines. From East India. M. Marc. Museum. 

Face and forehead brown, this latter narrow, especially an- 
teriorly, with a raised line and a small anterior callus, blackish. 
Antennae brownish testaceous, the third joint elongated, with a 
very short tooth, the last divisions brown. ‘Thorax brown; sides, 
outer border and scutellum pale ferruginous. Abdomen with the 
first two segments pale ferruginous, transparent, with light whitish 
reflections ; the others shining black. Iegs blackish; tibiae yellow- 
ish black at the apex. Wings on anterior half yellowish and pos- 
teriorly of a light brown. Macq., Dipt. exot., i (I), p. 130. 

Two females from Sumatra (V. Lansberge). In the most 
essential points they agree with the description. There are how- 
ever some disagreements to be mentioned. 

Ist. According to the description the third and following seg- 
ments of abdomen ought to be shining black ; in my specimens the 
colour of these segments is fuscous and rather dull; moreover the 
livid margins of segments show remains of a white hair-fringe ; the 
under surface is wholly fuscous and there the white fringe is com- 
piete, even on the second segment. 

and. According to Macquart the legs are blackish, the tibiae 
yellow with black tips; this may be the truth in regard to the 
front legs ; in the posterior ones, however, I find the tibiae brown 
and not darker at the end. 

This species may be easily recognised by the light brownish 
yellow colour of the first two abdominal segments, on account 
of which it resembles more or less the N. American Tabanus 
cinctus, Fabr. The eyes are bare, reddish bronze, without cross- 
bands, even after having been moistened, and with very small 
facets all over. V. der Wulp, Notes Leyden Museum, vii, 
Darya: 

Type in Paris Museum seen by me, has the first posterior cell 
of the wing closed, a fact not mentioned by Macquart or V. der 
Wulp. 

The abdomen is elongated, the first two segments livid, the 
remaining ones brownish black. Scutellum livid. Thorax brown, 
livid at base. Wings tinged brown, clear at the extreme apex, 


148 Records of the Indian Museum. [Vor21V5 


yellowish at the base. Only the fore pair of legs are remaining; 
these are brown, the tibiae pale. Forehead about ten times as long 
as it is broad, narrowed anteriorly. Frontal callus oblong, no 
linear extension apparent. Palpi large, broad, ending in a point, 
reddish, flat. Antennae reddish with hardly any tooth. Length 
18mm. A specimen in Brit. Mus. coll. from Chantabun, Siam, 
which I believe to belong to this species—though owing to its 
bad preservation it is not possible to be certain—has the scutellum 
covered with grey tomentum, the Jegs as Schiner describes, and 
measures 15 mm. ‘The beard is brown. 

The female of Tabanus crassus, Walker, has not vet been dis- 
covered; it is just possible this may prove to be it, though it is a 
very differently shaped insect. The closing of the first posterior 
cell is not always present in both sexes. 


Tabanus auriflamma, ? , Walker. 


List Dipt. Brit. Mus., i, p. 155 (1848). 


Type (female) and two others from Sylhet, Assam, in Brit. 
Mus. coll. 


In Indian Mus. coll. one female from Sibsagar, Assam, 
another from Naga Hills and another from Cachar, both 
localities on the border of Burma and Assam. 

In Kertesz coll. one female from Sylhet, Assam. A handsome 
yellow-and-black species with yellowish wings, brown at apex 
and on posterior border, abdomen yellowish with black bands. 
Legs yellowish. Antennae yellow. Length of type 22 mm., 
others from 20—24 mm. 

Face, subcallus and forehead covered with bright golden yellow 
tomentum; hairs on face and beard yellow or yellowish brown. 
Palpi reddish yellow with brown pubescence, stout, ending in an 
obtuse point. Antennae reddish, the first two joints covered with 
greyish tomentum and with brown hairs, the third joint slender, 
with no tocth and hardly a perceptible angle. Forehead parallel, 
about four times as long as it is broad, the frontal callus red-brown, 
shining, almost square, not reaching eyes, with hardly any lineal 
extension in fresh specimens ; in denuded ones it appears. Thorax 
and abdomen yeliowish. the former with some greyish brown tomen- 
tum and browner on the dorsum; pubescence scanty, consisting of 
short yellow hairs, shoulders with longer soft yellowish brown 
hairs. Breast the same colour. Scutellum yellow. Abdomen with 
broad blackish brown bands on the second, third and fourth seg- 
ments, and a very narrow one on the fifth, all situated on the 
anterior half of segments, taking up half the width of segment on 
the second, rather narrower on the third and fourth, and on the 
fitth hardly noticeable ; in some specimens the bands are reddish 
brown, the pubescence on the yellow parts golden yellow, thick, on 
the bands black ; under side similar. Legs wholly reddish yellow, 
in the type the fore tarsi and tibiae are darker, owing to discolora- 
tion, pubescence reddish yellow. Wangs large, deeply tinged with 


rgtt.] G. Ricarpo: Revision of the species of Tabanus. 149 


yellow, brown on the apex, beginning from the base of the fork of 
third vein and extending along the posterior border, gradually 
becoming narrower as it approaches the base of wing, the colora- 
tion very distinct and the two colours sharply divided, veins yellow, 
brown on the brown part, stigma yellow, no appendix, all posterior 
cells widely open. 


Group VII. 


Forehead with one square or narrow or oblong callus always 
prolonged towards the vertex as a more or less narrow line, from 
five to nine times as long as it is broad. 

Species of medium size or small, with one or more stripes on 
abdomen, usually continuous, the median stripe always so. 

Tabanus sirtatus, Fabr., is the typical species of this group. 
Tabanus auristriatus, n sp., hybridus, Wied., and aurotestaceus, W1k., 
form almost a subgroup. Tabanus immanis, Wied., is not known 
to me and is not included in the table; neither is Tabanus bubalh, 
Doleschall, which will prove hard to identify from its very meagre 
description. 


Tabanus annamitus, @ , Bigot. 


Mem. Soc. Zool. France, v, p. 630 (1892) (Bellardia). 
Type, male, from Saigon, Cochin China. 


A species with a fulvous abdomen on which appears an in- 
distinct blackish median stripe, the apex blackish. Wings clear. 
Legs yellowish. Length 15 mm. 

Eyes large, with the large facets very distinct, taking up fully 
two-thirds of the eye, the small facets on lower part continued to 
vertex as a narrow border,no appearance of hairs on eyes. Frontal 
triangle chestnut-brown. Subcallus and face covered greyish 
tomentum and a few white hairs. Antennae now incomplete, 
described as yellow, black at apex; the first two joints with black 
pubescence. Palpi yellow, with paler pubescence. Thorax blackish 
with grey tomentum. Abdomen reddish yellow, with some grey 
tomentum, especially visible on the black narrow median stripe 
which becomes broader towards apex; under side reddish yellow, 
apex blackish. Legs yellowish, the fore femora darker, covered with 
grey tomentum and with black pubescence, the posterior pair with 
chiefly white pubescence, elsewhere it is mostly black. Wings quite 
clear, veins and stigma yellow, the first posterior cell closed at 
border. 

I have seen no specimens identical with this species. 


Tabanus striatus, 7 2, Fabr. 
(Pl. xiii, fig. 7.) 


Ent. Syst., iv, p. 371 (1794); zd., Syst. Antl., p. 103 (1805) ; 
Wied., Dipt. exot., p. 79 (1821); zd., Auss. Zweifl. Ins., 1, p. 155 


150 Records of the Indian Museum. [VoEssly- 


(1828); V. d. Wulp, Sumatra Exped. Dipt., p. 16 (1881) ; id., Notes 
Leyden Museum, vii, p. 71 (1885). 

Tabanus dorsilinea, ¢ , Wied., Anal. Entom., p. 22 dee id. 
Auss. Zweifl. Ins., i, p. 184 (1828). 

Tabanus sinicus, 7 , Wlk., List Dipt., i, p. 163 (1848). 

Tabanus tenens, 2 , Wik., Ins. Saund. Dipt., i, p. 49 (1850). 

Tabanus mégalops, 7, Wlk., List Dipt., v, Suppl. i, pp. 247, 
407 (1854). 

Tabanus partitus, 2 , Wlk., Proc. Linn. Soc. London, i, p. 9 
(1856). 

? Tabanus manilensis, @ , Schiner, Reise Novara Dipt., p. 84 
(1868), 

Atylotus macer, @ , Bigot, Mem. Soc. Zool. France, v, p. 649 
(1892). 

Tabanus rufocallosus, 2 , Bigot, /. c., p. 679. 


d 


Reddish brown; thorax with white stripes, abdomen with 
three wide white stripes. Length 5-6 lines, 7 9. From China 
and Java. Fabr., Ent. Syst., iv, 371, 39 (1794). 

Ashy grey, abdomen brown with three abbreviated white lines. 
Fabr., Syst. Antl., 103, 47 (1805). 

Antennae rusty yellow. Face grey-haired. Beard whitish. 
Forehead yellowish with a smooth band widened above and below 
into a square callus. Thorax in certain lights appearing white, 
with four broad whitish stripes and a line in the middle always 
apparent. Sides of breast white-haired. Abdomen in unrubbed 
specimens with three broad white unabbreviated stripes; the 
side borders also white so that really five stripes are present. 
Wings hyaline. Halteres brownish with white knob. Femora 
reddish rusty brown, tibiae paler. Tarsi blackish brown. Wied., 
Auss. Zweifl. Ins., i, 155, 69 (1828). 

Van der Wulp records a male from Soerian Sumatra, in 


Sumatra Exped. Dipt., 16, 3 (1881), and gives the following 
details :— 


““o@. Eyes are naked, flatter above than in other species, with 
a coarse network and a purple transverse band, below which the 
network becomies very fine. The white stripes on the thorax are 
indistinct, as are the two side stripes on the dorsum of 
abdomen, which latter ends in a point. The wings are quite clear, 
but have a pale grey tinge, the costal cell is yellow, the stripe- 
like stigma yellow-brown.” 

In Notes Leyden Museum, vii, 71, 24 (1885), he records 7 
and ¢ from Java and Sumatra and adds— 

““Fyes bare, in # reddish bronze above with large facets, the 
lower third and the borders blackish with small facets ; the two parts 


distinctly separated; in the 9 the eyes are uniform bronze with 
small facets. ’’ 


To1t.}| G. Ricardo: Revision of the species of Tabanus. I5I 


This species has a wide distribution, being found from India 
to Sumatra. 

In the Brit. Mus. coll. are ~ 2 specimens from N. India, 
Pusa Bengal, Bassein Bombay, Calcutta; Trincomalee, Tamble- 
gam, Kandy and other places in Ceylon; Upper Burma; Kuala 
Lumpur, Federated Malay States; Cavité, Philippines; and from 
Kajoe-Tanan, Sumatra (Hagen). 

In Indian Mus. coll. from Bunkutwa, Gonda District, U. P.; 
Calcutta; Murshidabad, Purneah District; Port Canning; Bengal ; 
Madras Coast ; and Balighai, near Port Orissa E. India. 

In Howlett coll. from Belgatchia and Pusa, Bengal; others 
identified from Sylhet, Assam; and Manipur, Burma. 

The only species likely to be confused with this species are 
Tabanus albimedius , Walker, and Tabanus rubidus, Wied., from both 
of which it may be distinguished by the spindle-shaped continuation 
of the frontal callus, by the almost straight median stripe, narrow 
almost continuous lateral stripes and by the slender more or less 
pointed abdomen. Length from 12—20 mm. A female specimen in 
the Paris Museum placed under this species and supposed to be the 
original type is a specimen of Tabanus albimedius, Walker, from 
java, but as the original description very clearly states there are 
three stripes, the type has probably been lost or confused; there is 
a specimen side by side with this supposed type, determined by 
Macquart as Tabanus striatus, with the usual striped narrow 
abdomen. 

Some small specimens, females, from Mohmand Pass, N. W. 
Frontier, in the Howlett coll., which I took at first to be a new 
species, only measure 11 mm. but I can find no other specific 
character to divide them from Tabanus striatus. 

9. Face greyish, with white pubescence. Beard white. Palpi 
very pale yellow with white pubescence and some black hairs, 
curved on upper side, with a rather long apex. Antennae reddish 
yellow, darker at apex. Forehead rather narrow, slightly narrower 
anteriorly, about six times as long as it is wide, covered with 
yellowish grey tomentum, some short white pubescence is visible, 
on the vertex are some black hairs. Frontal callus shining red- 
dish brown, oblong, hardly reaching the eyes anteriorly, posteriorly 
receding from them and continued in a fine line for a very short 
distance, then becoming broad, at least half as wide as the callus 
itself, often starting direct from the callus as a broad stripe. 
Pubescence on hind part of head short and white. Thorax reddish 
brown, or brown with four fairly distinct greyish white tomentose 
stripes with white pubescence, elsewhere on the dorsum the pubes- 
cence is black, sides of thorax with biack hairs, breast grey with 
white pubescence. Scutellum same colour as thorax, with black 
pubescence and white hairs round the posterior border. 

Abdomen usually long and slender, ranging from reddish brown 
to blackish brown, the median stripe continuous, reaching the sixth 
segment, composed of oblong grey tomentose spots slightly narrower 
at their apices on the second and third segment, the side stripes 


152 Records of the Indian Museum. PViOL-ShVE 


usually prolonged very distinctly to the fourth segment, appearing 
on the fifth as an indistinct spot, the sides of abdomen white with 
white hairs, the pubescence on dorsum black, white on the stripes. 
Under side reddish yellow or darker, covered with grey tomentum. 

Legs reddish yellow, the femora more rusty reddish, often ap- 
pearing darker owing to the black hairs situated chiefly on the upper 
sides, in some specimens the middle and posterior pair are blackish 
covered with grey tomentum, the pubescence is always predomi- 
nantly whitish ; on the coxae it is white and on the tibiae with black 
hairs on the upper sides, the tarsi reddish brown with black pubes- 
cence. Wings hyaline, stigma yellow, veins reddish. Halteres 
yellow or brown with lighter knob. 

In a specimen from Malacca the fore coxae and femora are shin- 
ing black with grey tomentum and wholly white pubescence. 

a”. These are all reddish brown in colour of abdomen; in 
Tabanus sinicus ¢ type and another specimen from Calcutta, the 
femora and the greater part of the fore tibiae are blackish. The eyes 
are as Van der Wulp describes them, but the large facets are 
yellowish green in colour with a large brown band across the centre, 
only in the type of Tabanus sinicus, is this not apparent. The 
males vary from I2—-20 mm. in size. 

Tabanus dorsilinea, », said to be nearly related to Tabanus 
styviatus in the description but much smailer and different, has 
been fong placed as a synonym and apparently correctly, though 
I have never seen a specimen of Tabanus siriatus with ‘‘ segmenta- 
tions of abdomen widely ochre-yellow,’’ as Wiedemann describes. 
His type came from E. India. 

Tabanus sinicus, ¢ , from Hongkong, has rather darker legs 
than usual, measuring 20 mm. length. 

Tabanus tenens, 2, from India, has legs rather paler than 
usual. 

Tabanus megalops, 7, from Java, is more reddish in colour 
than usual. 

Tabanus partitus, @ , from Singapore, has the femora rather 
darker than usual. 

Atylotus macer, # , from India is labelled as Atylotus mucro- 
loma, but from the description it is evidently the type of A. macer ; 
it is a small specimen in very bad condition, as remarked by Bigot 
in his description ; it appeats to be a small specimen of Tabanus 
striatus, the three stripes clearly visible though Bigot speaks of two 
only. The type of Tabanus rufocallosus is from Java, nearly all 
the legs were wanting when described, the abdomen is partly des- 
troyed which accounts for Bigot describing the median stripe as 
ending on the third segment ; in reality it appears to end on the 
fourth segment, as the fifth is incomplete; the fore femora are 
rather darker than is usual. 

Tabanus manilensis, 2 , is apparently from the description a 
specimen of Tabanus striatus, with the side stripes indistinct and 
the legs rather darker than usual; it is described as from Manila. 
Length 13 mm. one specimen only. Tabanus chinensis, Thunberg 


Igii.] G. RicarDo: Revision of the species of Tabanus. 153 


(N. Acta. Reg. Soc. Upsaliensis, ix, 53-62, 1827), recordedas from 
China and Cape of Good Hope, was probably a specimen of Tabanus 
stytatus, or of Tabanus taeniola, P. B., from the Cape, or Thunberg 
may have had specimens of both species before him, the African and 
Indian species being very nearly related, but distinct ; the frontal 
callus alone would divide them, but his description is not full enough 
to make it possible to decide; the species may well be deleted from 
the list of Tabanus species. See Bezzi, ‘‘ Nomenkiatorisches wber 
Dipteren’’ in Wien. Ent. Zeit., xxvii, vol. ii and iii (Feb. 
1908). 

Tabanus costalis, Lichtenstein, Catalogus, p. 213, Hamburg 
(1796); Austen, Ann. Mag. Nat. Hist. (8), i, p. 346 (1908); from 
Coromandel. This species, described in under a dozen words and 
absolutely indeterminable but apparently thought by the author 
to be near Tabanus striatus, should be deleted from the list of 
species of Tabanus. 


Tabanus hilaris, 7 @ , Walker. 


Ins. Saund. Dipt., p. 40, pl. ii, fig. 3 (1850). 


In Brit. Mus. coll.; the type ~ from India, others from the 
Punjab and N. W. India. 


In Indian Museum coil. ~ and 2 specimens from Bhogaon, 
Purneah district, N. Bengal, ‘‘resting in numbers on tree trunks 
during the day, common in the evening on the stomachs of cows ”’ 
(C. A. Paiva). In Howlett coll. 7 and 9 specimens from Belgatchia 
and Pusa, Bengal, others sent for determination came from Sylhet, 
Assam. 

A species distinguished from T. stviatus, F., by the short median 
stripe of abdomen which does not begin till the thivd segment and 
by the shorter lateral stripes which usually terminate on the third 
or fourth segment. The frontal callus is very similar in shape, 
shining red-brown, rather protuberant with a spindle-shaped exten- 
sion, the forehead almost the same width throughout. The palpi 
shorter and stouter. Legs darker, the femora being all blackish 
or reddish brown. In colouring the abdomen is _ blackish 
brown with the stripes grey. The thorax with two stripes which 
fade away on the anterior border. ‘The males are similar, in the type 
the abdomen is more reddish brown than black. Eyes in males 
with a broad band of very large facets, leaving the lower half of 
eyes and a narrow border continued to the vertex composed of 
small facets; across the yellowish large facets is a well-marked 
broad brown band. 

Length of male specimens 11} mm., of females 14 mm. 


Tabanus abbreviatus, ? , Bigot. 


Mem. Soc. Zool. France, v, p. 670 (1892). [Atylotus.| ? Aty- 
lotus conicus, @ , Bigot, /. c., p. 650. 


154 Records of the Indian Museum. VoL. IV; 


Type @ and another from Java, in bad preservation. In Brit. 
Mus. coll. a female from Singapore (Falshaw), ‘‘ Biting horses and 
cattle, more frequently seen in the last six months than in former 
years;”’ a female from Selangor (Butler), ‘‘Common, bites men 
and horses.’’ In Howlett coll. females from Calcutta and from 
Belgatchia, Bengal. 

in Kertesz coll. four females from Formosa. The type had 
the antennae incomplete when described and is now in a dirty 
denuded condition ; the description given below is based largely on 
the modern specimens. 

A reddish yellow species with the thorax blackish, abdomen 
with a narrow grey median, and fainter lateral stripes. Antennae 
and legs reddish brown. Length type 16 mm., other specimens 
{from I4—17 mm. 

This species is distinguished from Tabanus albimedius, W1k., by 
its smaller size, narrower forehead especially above antennae, more 
slender palpi, covered with black pubescence, darker legs and 
thorax, abdomen reddish yellow, blackish at apex. face covered 
with grey tomentum and with silvery white pubescence. Beard 
the same colour. Palpi pale yellow thickly beset on their outer 
side with black hairs, long and slender, the apical part as long as 
the stouter basal part. Awtennae red-brown, blackish on the last 
half of the third joint which is broad at its base with a prominent 
tooth, the first two joints with black hairs. Forehead narrower 
anteriorly, covered with yellowish brown tomentum and with black 
hairs about eight times as long as it is wide anteriorly, the frontal 
callus oblong, not reaching the eyes, diminishing to a long narrow 
line, reddish brown or brown in colour. Thorax blackish covered 
with yellow brown and grey tomentum and with white and black 
hairs on dorsum, the grey stripes indistinct, sides with black hairs, 
but tufts of whitish or yellowish hairs round base of wings; breast 
grey with white pubescence. Scutellum as thorax. Abdomen red- 
dish yellow, on each side of the median stripe it is usually brown, 
the last three segments largely black, the median stripe is narrow, 
usually continuous, covered with grey tomentum and with black 
hairs, some yellow hairs are often present, the side stripes are faint, 
indicated by grey tomentum and by yellow hairs, the pubescence 
on the dorsum is otherwise black, thicker at the apex: under side 
lighter. Legs reddish, femora darker, especially the anterior pair, 
with grey tomentum and largely yellowish pubescence, fore tibiae 
at apex and fore tarsi blackish, other tarsi brown. Wdngs clear, 
costal border yellow, veins brown, stigma yellow. 

In one of the specimens from Formosa the median stripe is 
broken up into narrow triangular spots which however join; in 
fresh specimens the side stripes appear more distinct as oblong 
narrow spots forming more or less continuous stripes. 

Atylotus conicus @, Bigot, from India is probably the ~ of 
this species. Type is in fair condition; eyes with large facets 
above. 


19g1I.] G. Ricarpo: Revision of the species of Tabanus. 155 


Tabanus rubidus, ? , Wied. 


Dipt. exot., 69, 14 (1821). 


9. Lilac brownish. Thorax with whitish stripes. Abdomen 
with three whitish stripes composed of spots. 8 lines. From 
Bengal, Wied., Dipt. exot., 69, 14 (1821). 


Related to Tabanus autumnalis. Antennae rather reddish, the 
end joint almost wholly blackish. Face and beard white. Palpi 
very pale yellowish. Forehead grey. Frontal stripe narrow end- 
ing below in a narrow egg-shaped reddish callus. Thorax lilac red- 
dish brown, with grey tomentum and five whitish, in certain lights, 
grey-haired stripes; breast sides rather hoary white with white 
tomentum. Abdomen lilac reddish brown, median stripe com- 
posed of narrow triangular side stripes of nearly square spots: side 
borders yellowish white, in certain lights the abdomen appears a 
little grey haired. Wings in the middle rather yellowish ; halteres 
rather reddish. Femora very light lilac reddish, above with a. 
blackish stripe, tibiae more yellowish, both with white pubescence, 
tarsi brown. Wied., Auss. Zweifl. Ins., i, 127, 25 (1828). 

Van der Wulp in Notes Leyden Museum, vii, 71, 23 (1885), 
records females from Java and Sumatra. Eyes with nocross-bands. 

In Brit. Mus. coll. 2 specimens from Moulmein, Burma ; 
Bombay ; Kuala Lumpur and Singapore, Federated Malay States ; 
Siam (Woolley). On the female from Singapore is the following 
note: ‘‘ Biting horses and cattle, more frequently seen during the 
past six months than in former years.’’ (Falshaw.) 

In Howlett coll. from Pusa and Belgatchia. In Indian Mus. 
coll. from Sibsagar, Assam; Bhogaon and Purneah District, N. 
Bengal: ‘“‘ Resting in numbers on tree-trunks during the day, 
common in the evening on the stomach of cows;’’ from Khasi 
Hills, Assam; from Partabgarh, U. P. 

This species is distinguished from Tabanus striatus, Fabr., in 
general by its broader, larger abdomen; the frontal callus is not 
prolonged in a spindle-shaped thick line but as a narrow short line; 
the median stripe of abdomen is not so straight, the sides are 
serrated, the spots composing it being more triangularlin shape, 
the side stripes do not appear so regular and continuous, being 
composed of very similar spots, but their apices as a rule are 
turned inwards so that the stripe appears irregular. The /egs as a 
rule are darker, especially the femora, and the apices of the fore 
tibiae, which are blackish, their basal part yellowish white. The 
palpi are pale yellow or whitish with mostly black pubescence, 
stout and large. The face is covered with grey tomentum and 
with white hairs. Forehead slightly narrower anteriorly, about 
five times as long as it is broad. Inthe specimen from Assam the 
wings are tinged with brown. _ Length of specimens 17—20 mm. 
The males of this species if they do not retain the dark colour of 
the females will be difficult to distinguish from those of Tabanus 
albimedius, W1k., only the darker femora serving to divide them. 


156 Records of the Indian Museum. [Vor gi 


I have not yet seen any males which appear to belong to this 
species. 


Tabanus albimedius, ° o , Walker. 
(Piven ste 26.) 
Dipt. Saund., p. 48 (1850). 
Tabanus vagus, 2 , Walker, Dipt. Saund., p. 50. 
Tabanus umbrosus, ° , Walker, Dipt. Saund., p. 52. 
Tabanus priscus, 2 , Walker, List Dipt., i, p. 176 (1848). 
? Tabanus calidus, @ , Walker, Dipt. Saund., p. 57. 


Aiylotus lachrymans, 2? , Bigot, Mem. Soc. Zool. France, v, 
p. 669 (1892). 

In Brit. Mus. coll. the type (2) of albimedius from E. India 
(Saunders coll.), other specimens from Nepai; India; and a long 
series of males and females from Ceylon (Yerbury coll.) ; these lat- 
ter are said to be common on the road from Trincomalee to Kan- 
thalla. 


In Howlett coll. females from Umballa, goo ft., ‘‘ lying close 
on tree trunk;’’ males from Calcutta; and males and females from 
Pusa, Bengal. 

This species is very nearly allied to Tabanus rubidus, Wied., 
being chiefly distinguished from it by the lighter colour of abdomen 
which is not uniformly blackish brown or lilac-brown, as is the case 
in typical specimens of the Wiedemann species, but is reddish 
brown, usually darker brown each side of the median stripe, which 
latter varies from narrow to wider triangular spots forming a more 
or less continuous stripe. Under side reddish yellow. Thorax red- 
dish brown with grey tomentum and indistinct grey stripes. Legs 
are lighter in colour, the femora reddish brown, the tibiae yellowish 
but the apices of the fore tibiae blackish, the tarsi reddish brown. 
9. Forehead five to six times as long as it is broad, slightly 
narrower anteriorly, frontal callus reddish brown, not quite reach- 
ing the eyes, with a narrow linear extension, which at once dis- 
tinguishes it from Tabanus striatus. Palpi, very pale yellow with 
few or no black hairs. Antennae red, darker at the apex. Length 
1343—181 mm. ‘The male is difficult to distinguish from the males 
of Tabanus striatus, but the shape of median stripe is different, 
being not so straight or continuous. The males of the three 
species striatus, albimedius and yvubtdus are all somewhat dificult 
to distinguish from each other. 

The species is very nearly allied to Tabanus rubsdus, Wied., 
and may perhaps eventually be regarded as only a form of the 
above, as many of the specimens of both species are variable in 
the colouring of the abdomen and of the legs and therefore 
difficult to divide from each other. 

The series from Ceylon have the forehead rather narrower 
anteriorly and the abdomen is rather more reddish yellow. 


I9g1t.] G. RicaRDO: Revision of the spectes of Tabanus. 157 


Tabanus vagus, type, @, is from KE. India (Saunders coll.), 
another female from same locality and one from Java were 
identified by Walker as this species, but I find no difference in the 
specimens from those of Tabanus albimedius. 


Tabanus umbrosus, type, 2 ,from E. India (Saunders coll.), has 
the legs somewhat darker than usual, and the median stripe is 
straighter and narrow. 


Tabanus priscus, type, 2 , from an unknown locality, appears 
to be identical with T. albtmedius, but priority is not given to it 
owing to the doubt as to locality. 

? Tabanus calidus, type, 2 , from Asia, but in the description a 
query accompanies the locality ; this is presumably a specimen of 
Tabanus albimedius, very similar to the series of specimens from 
Ceylon. 

Atylotus lachrymans, 2 , Bigot, from Java, appears identical 
with the Walker species. 


Tabanus speciosus, 2 , n. sp. 


Type o from Travancore. 

Type @ from india (Wroughton), and another female from 
Tamblegum, Ceylon (Yerbury), in Brit. Mus. coll. In Indian 
Museum coll. female from Travancore, S. India. 

A distinctly marked conspicuous species with a superficial 
resemblance to Tabanus reducens, Wik., from Celebes, but dis- 
tinguished from it by the presence of side spots. It is allied to 
Tabanus rubidus, Wied., but the conspicuous spots on the third 
and fourth segments of abdomen with wide bases will distinguish 
it; on the second segment the spot is very narrow so that the 
stripe has a much more unequal appearance; and it is larger in size. 

A black species, with three series of white spots on abdomen, 
legs blackish with yellow tibiae, antennae red, wings almost clear. 
Length 19 mm. 

@. Face covered with greyish white tomentum, with silvery 
white pubescence and beard. Palpi pale yellow with some black 
pubescence, stout, ending in a short acute point. Antennae red, 
duskier at the apices, the first two joints pale yellow with black 
pubescence, tooth of third joint small but distinct. Subcallus same 
colour as face. Forehead with rather darker tomentum and with 
some white pubescence, about seven times as long as it is wide, 
only slightly narrower anteriorly, the frontal callus reddish brown, 
large, oblong, not reaching the eyes, the short linear extension not 
always visible. Thorax reddish brown in type, probably blackish 
brown when not denuded, with five distinct grey tomentose stripes, 
with yellowish white pubescence on them, sides with black hairs, 
round the base of the wings with white ones; breast covered 
with grey tomentum and with white pubescence. Scutellum 
same colour as thorax, with white pubescence at sides and on 
posterior border. Abdomen blackish brown, the median stripe 


158 Records of the Indian Museum. [VorL. IV, 


composed of four whitish tomentose spots, the one on the second - 


segment narrow triangular, on the two following segments broad 
triangular, on the fifth the same but a little narrower, all with 
their apices reaching the border of the next segment, the spots on 
the sides distinct and large but less defined in shape and only 
present on the second, third, and fourth segments, oblique, inclining 
inwards from the outer border, the one on the fourth segment 
smaller; all these spots are white haired, pubescence otherwise on 
dorsum short, thick, black, sides yellowish with white pubescence ; 
under side reddish brown with an indistinct median black stripe, 
pubescence white. Legs blackish, the coxae and femora covered with 
dense grey tomentum and with white pubescence, tibiae reddish 
yellow, the fore pair only so on their basal half, with white pubes 
cence, which is also present but less thick on the other tibiae; 
pubescence on apices of fore tibiae, the extreme apices of the others 
and on the tarsi black. Wungs hyaline, a hardly noticeable brown- 
ish tinge on the fore border, veins brown, stigma reddish yellow. 
Male identical with female, more reddish incolour. Eyes very large, 
the large facets occupy fully two-thirds of the eyes, the lower third 
composed of small facets which reach to the vertex as a narrow 
border, a dark brown band crosses the coppery coloured large facets. 


Tabanus hirtistriatus, 2 , n. sp. 


Type ( @ ) and five other females, the type from Perak, the others 
from Durien Tipus, Negui Sembilan, Federated Malay States, in col- 
lection sent me by Dr. Stanton. A small brownish species, with a 
very narrow forehead, anteriorly barely half as wide as it is at the 
base, the abdomen with a grey tomentose median stripe clothed 
with yellowish or whitish hairs, this stripe often seems almost 
obsolete in rubbed specimens. Wings tinged with brown on the 
fore border. Length of type 114 mm., others from I2—r4 mm. 

Face covered with grey tomentum and some white hairs, sides 
of cheeks and subcallus with yellowish brown tomentum. Beard 
white. Palpi small, a little stout at base, ending in a point, pale 
yellowish with black pubescence. Antennae slender with an acute 
but very short tooth at base of third joint, bright red, black at apex, 
the first two joints with black pubescence. Forehead covered with 
same coloured tomentum as subcallus, the frontal callus very 
narrow, oblong or cylindrical with a fine linear extension, not 
reaching the eyes, brown in colour, about nine times as long as it 
is wide. Thovax blackish brown with fine greyish yellow short 
pubescence, at sides with black pubescence. Scutellum is similar. 

Abdomen blackish brown, more yellow-brown on the first two 
segments, sometimes reddish brown and black at apex, the 
pubescent stripe continuous with straight sides; under side 
yellowish brown covered with grey tomentum, and with white- 
haired segmentations. Legs brownish, the tibiae yellow, black at 
apices. Wings hyaline, tinged with brown on the fore border 
reaching the apex, a short appendix usually present. 


a 


tgtr.| G. RicaArpo: Revision of the species of Tabanus. 159 


There are a series of males in the Indian Museum coll. from 
the base of the Dawna Hills, Lower Burma (Annandale), which 
may possibly prove to be the male of this species ; the abdomen and 
legs are yellower; all the facets of the eyes almost the same in size. 
A note is attached to them, wiz., ‘‘ Hovering over cart track in the 
bright sun.” 


Tabanus monotaeniatus, 9°, Bigot. 
Mem. Soc. Zool. France, v, p. 655 (1892). [Atylotus.] 


Type (?) and four others from India (only three specimens 
are mentioned by Bigot in his description) and one other ( 2 ) from 
N. Khasi which was placed under his species Tabanus montlifer. 


In Indian Museum coll. three para-types from India, one of 
them from Sibsagar, and another from N. Khasi, Assam. In 
Howlett coll. seven @ from Nangpoh, Assam, ‘‘ caught on animal.’’ 
In Brit. Mus. coll. @ specimens from Dibia and Sadia, Khasi 
Hills District: and N. Khasi Hills, lower ranges (Chennell); and 
Nangpoh, Assam. A blackish brown or reddish brown species, 
the abdomen with a narrow distinct grey tomentose median stripe 
reaching to the sixth segment, the sides straight: the thorax blackish 
with indistinct stripes. Legs blackish, with red tibiae, antennae 
and palpi. 

Length of the type 18 mm., other specimens range from 
I4—194 mm. 

Face covered with greyish tomentum and with white pubes- 
cence, a few black hairs on cheeks. Palpi large, reddish or yellow, 
covered with black hairs, the inside bare, ending in rather an obtuse 
point. Beard white. Avztennae reddish, the third joint dusky, and 
blackish at apex, ouly red at base, the tooth obtuse, the first joint 
with grey tomentum and with black pubescence, the second with 
black pubescence on its outer border. Forehead covered with 
yellowish tomentum, with some short black pubescence, very nearly 
the same width throughout, very slightly narrower anteriorly, quite 
six times as long as it is wide, the frontal callus club-shaped, the line 
proceeding from it being thick; the callus does not reach the eyes 
and is dark brown in colour. Hind part of head with short white 
pubescence, some black hairs at vertex. Thorax black with yellow- 
ish brown tomentum and with traces of four indistinct grey stripes, 
the shoulders and sides of thorax in some of the specimens reddish 
brown, the pubescence of dorsum short, black, hairs at sides black. 
Scutellum black with grey tomentum and black pubescence ; breast 
and sides covered with greyish tomentum and with white hairs. 
Abdomen blackish brown, or reddish brown, with some yellowish 
brown or greyish tomentum which in some specimens appears as 
indistinct roundish spots, the dorsum rather thickly covered with 
short black pubescence, with the exception of the median stripe on 
which a few white hairs are visible, the hairs on sides of first 
three segments are black, afterwards white, the segmentations very 
narrowly lighter; under side blackish brown or reddish brown 


160 Records of the Indian Museum. [Vor ive 


covered with grey tomentum, the segmentations yellowish, the 

pubescence chiefly black. Legs: the coxae and femora black with 
grey tomentum and with white pubescence, the tibiae reddish, darker 
at extreme apex, with chiefly black pubescence, the tarsi the same 
colour but appearing darker on account of the thicker black pubes- 
cence. Wangs grey, tinged with brown on fore border, and round 
veins. Halteres reddish brown, knobs pale yellowish red. 

The two females from N. Khasi are rather paler in colouring, 
the abdomen more reddish, the thorax with sides and shoulders 
reddish, the legs slightly redder, the wings clear with stigma and 
veins yellow. 

The type and the majority of the specimens have a blackish 
brown abdomen, but some are reddish brown. 


Tabanus brunnipennis, @ , n. sp. 
(Pipsais figs) 


Type (¢) and two other females from Basi, N. Kanara, S. W. 
India (T. R. Bell), 1908; one 92 from Bangkok, Siam (5S. 8. 
Flower), 1908. 


In Howlett coll. one @ from Gorakhpur, U. P., India.' 


A well-marked brown species with a rather broad very dis- 
tinct grey median stripe on abdomen, and two grey spots on the 
second segment. Legs and antennae reddish. Wings grey, tinged 
brown on fore border as far as the second longitudinal vein. 
Length of type 16 mm., other specimens from 13—15 mm. It is dis- 
tinguished from T. monotaeniatus, Bigot, by its smaller size, and 
the wings have not the cross-veins shaded, but the tinged fore 
border sharply defined. 

Face covered with whitish grey tomentum and a few scattered 
white hairs. Beard white. Palfz yellow with rather numerous 
black hairs, stout at base ending in a long obtuse apex. Antennae 
reddish, the first joint appearing paler with whitish tomentum and 
some very fine black pubescence, the third slender with a very 
slight tooth, brown at apex. Forehead covered with yellowish 
brown tomentum and with a few black hairs, the same width 
throughout, about six times as long as it is wide, the frontal callus 
shining reddish brown, not reaching the eyes, club-shaped, the 
linear extension thick ending in a point beyond the middle of the 
forehead, in some of the specimens the callus appears larger, 
apparently owing to denudation, a slight furrow appears each side 
of the apex of callus converging towards the vertex. Thorax 
blackish (when denuded reddish), covered with greyish tomentum 
and black pubescence, two indistinct grey stripes apparent, with 
traces of yellowish pubescence anteriorly, sides grey with white 
hairs, shoulders reddish with black hairs, breast covered with 
grey tomentum and white pubescence. Scutellum reddish with 
grey tomentum and biack hairs, white hairs on _ borders. 
Abdomen dark reddish brown, usually redder on the anterior 


1g1l.] G. Ricarpo: Revision of the species of Tabanus. 161 


segments, covered with dense but short black pubescence, on the 
stripe and spots with short white hairs; the spots are situated one 
on each side of the second segment, small and round, indica- 
tions of one on the third segment are present, the stripe measuring 
nearly I mm. at its broadest from the first to sixth segment, sides 
straight ; under side paler, reddish, with white and yellowish pubes- 
cence. Legs reddish, apices of fore tibiae and tarsi darker, femora 
with grey tomentum and chiefly white pubescence, tibiae and tarsi 
with chiefly black pubescence. Wings with the dark colouring very 
noticeable reaching to the apex, veins and stigma brown. 


Tabanus abscondens, 2? , Walker. 


Trans. Entom. Soc. London, v, p. 275 (1860). 


Type 2 from Burma; one @ from N. Chin Hills, Burma 
(Watson); one @ from Dawna Range, Tenasserim, Lower Burma, 
500—1,500 feet (Bingham); one @ from China in Dr. Kertesz’s 
collection. 


Type and specimens are all in a denuded faded condition, 
so that the redescription given below is imperfect. The spe- 
cies appears to be nearly related to Tabanus glaber, Bigot, but is 
distinguished from it by the frontal callus. Abdomen reddish, 
with a broad black median stripe. Length 16 mm., one specimen 
¥7, Mim. 

Face greyish with thick short white pubescence. Beard white. 
Palpt reddish yellow with black pubescence, long with a not very 
pointed apex. 

Antennae gone in type. Walker describes them as small, 
with the tooth very small; in the specimens from Tenasserim and 
China they are long and slender, the tooth prominent, reddish, the 
third joint darker. Forehead considerably narrower anteriorly, 
about six times as long as it is wide, the frontal callus oblong, 
narrow, nearly reaching the eyes with a short linear extension. 
Thorax blackish, red at sides, scutellum blackish, red on its outer 
border. Abdomen appears to be reddish with a broad black stripe 
on which appear traces of a narrow grey stripe, the sides of seg- 
ments black and the three lastsegments wholly blackish. Legs red- 
dish, the fore femora, apices of fore tibiae, and tarsi blackish. 
Wings clear, stigma yellowish, veins brown. 


Tabanus auristriatus, 9, 0. sp. 


Type @ and three others from Gersoppa, N. Kanara, S. W. 
India (Bell), 1908. 

A dark brown species with golden hairs on thorax and ab- 
domen, forming a stripe on the latter. Antennae yellowish. Legs 
brownish. Wings pale brown, deeper brown on the fore border. 
Length of type 14 mm., other specimens from I11—134 mm. 


162 Records of the Indian Museum. [Vor 2rv- 


Face covered with pale greyish white tomentum on lower 
part; a narrow band stretching across just below the antennae, 
subcallus and the forehead covered with rich vellowish brown 
tomentum, pubescence on face scanty, black. Beard sparse, white. 
Palpi long, narrow. yellowish brown with black pubescence. 
Antennae long, slender, the first two joints pale yellow with black 
hairs, the third red, black at apex, with a small tooth. Forehead 
narrow, the same width throughout, about six times as long as it 
is wide anteriorly, the frontal callus long, narrow, not reaching 
the eyes, prolonged in a thick line just beyond the middle of fore- 
head. Thorax and scutellum thickly covered with very short yellow 
pubescence, sides with black hairs, breast grey with white hairs. 
Abdomen yellowish brown anteriorly, darker at apex, with black 
pubescence, the median stripe is indistinct, chiefly marked by 
the golden yellow hairs which also border the segmentations; 
under side brown with white-haired segmentations. 

Legs brown or yellowish brown, fore tibiae a little paler at 
base. Wangs tinged brown, paler on the posterior border, stigma 
dark brown, veins brown. 

This species is distinguished from Tabanus hybridus , Wied., by 
the forehead not becoming narrower anteriorly, by the darker legs 
and indistinct narrower median stripe. 


Tabanus hybridus, 2 ~, Wied. 


Auss. Zweifl. Ins., i, p. 557, 31 (1828); Walker, Proc. Linn. 
Soc., i, p. 110 (1857) (recording the species from Macao). 


Oak-brownish with a brown-yellow striped abdomen. Wings 
brownish on the fore border, 54 lines, 2 from Macao. 

Of a slender form. Antennae rusty yellowish, the last joint 
with a hardly perceptible tooth at base; palpi brownish; face 
yellow; beard whitish; forehead yellow with a linear like callus 
gradually somewhat thicker below. Thorax oak-brown with two 
hardly perceptible lighter stripes ; breast sides greyish. Abdomen 
brown with a broad yellow stripe and quite narrow yellow side 
borders, the last segment black-brown; on the under side the three 
last segments are blackish with yellow side borders. Wings on 
the fore border brownish. Legs brownish yellow with black-brown 
tarsi. In my collection. Wied., Auss. Zweifl. Ins., i, p. 557- 

Van der Wulp in his Cat. Dipt. S. Asia also gives Borneo, the 
type being recorded from Macao, S. China. 

In Brit. Mus. coll. are 2 specimens from Kuala Lumpur and 
Sunghai, 5. Perak, Federated Malay States; also specimens sent 
to me by Dr. Kertesz for identification from Perak, and from 
Sylhet, Assam, others from Batu Tisa, Federated Malay States, 
Stanton coll. A very pale faded specimen determined by Walker 
as this species from Sarawak, Borneo, is probably correctly 
identified. 

@. An easily recognised species with the median yellow 
stripe on the yellowish brown sometimes darker brown abdomen, 


1g1r.] G. Ricarpo: Revision of the species of Tabanus. 163 


the stripe in well-preserved specimens is covered with golden 
yellow hairs and is continuous and straight at sides; the thorax is 
covered with short golden yellow pubescence and some black hairs 
intermixed. The forehead is narrow, quite seven times as long as 
it is wide anteriorly, where it is almost a third narrower than at 
vertex. The frontal callus is very narrow with a fine linear exten- 
sion. ‘The palpi are yellowish covered with black pubescence, rather 
stout, ending in an obtuse point. The /egs are brownish yellow, but 
the anterior and middle pair paler yellow, the fore tibiae often appear- 
ing white owing to white pubescence ; in some specimens however the 
pubescence is black. The wings have the costal border shaded 
brown extending to the apex, in some specimens it reaches to the 
hind border, stigma yellow. 

o@. Similar, the large facets of eyes occupy the greater portion 
of eye, the small facets on the lower part occupying about a third 
of the surface of eye, continued as a narrow border to the vertex. 

Length of specimens from 114 to 15 mm. 


Tabanus aurotestaceus, 7, Walker. 


List Dipt., v, Suppl. i, pp. 214, 253 (1854). 
Type from Shanghai, and a @ from China (Walker coll.). 


A species near Tabanus hybridus, Wied., but distinguished by 
the wholly clear wing and black fore femora. 

The o@ has a very large head, Atvylotus-like, flat above, the 
small facets occupying the lower third of the eyes and continued 
as a narrow border to vertex. Thorax and abdomen covered with 
yellowish tomentum and with black and some yellow hairs. Legs 
pale yellow, fore femora black, tarsi blackish. The yellow stripe 
on abdomen is very broad, about a third of the width of abdomen. 
Length 17 mm. 

2. <A poor specimen. ‘The forehead very narrow anteriorly, 
about nine times as wide as it is broad, with a long narrow frontal 
callus. Length 164 mm. 


Tabanus tristis, Van der Wulp. 
Sumatra Exped. Dipt., 17, 5, pl. i, fig. 9 (1881). 
One female from Datar, Sumatra, in May. 


Black. Abdomen with lateral stripes and antennae ferru- 
ginous, the lateral borders of thorax aud posterior border of scu- 
tellum pale haired. Wings ashy grey, the costa darkened. @. 
Length rr mm. 

Eyes naked, divided by a narrow, posteriorly somewhat 
enlarged, frontal band, on which a stripe-shaped, below a little en- 
larged, black line appears; the remaining triangular forehead, be- 
sides the face and the cheeks, with grey-brown tomentum ; the beard 
pale grey. Antennae rust-coloured; the first joint triangular, the 
second projecting above spine-like; the third above at the base 


164 Records of the Indian Museum. [VoL. IV, 


with a small tooth and at the apex darkened (see plate). Abdomen 
black; the borders of the thorax and the posterior border of the 
scutellum with pale pubescence: on the fore breast and the fore 
coxae a very long dirty white pubescence ; breast sides with ashy 
grey tomentum; the first four segments of abdomen with a rusty 
coloured stripe on each side, which is broadest at the base, but 
posteriorly narrower, a pale pubescence is visible on them; on the 
dorsum are some traces of reddish segmentation ; on the under side 
they are distinct and light grey. Legs black-brown, somewhat 
shining; the femora by their pubescence greyish. Halteres brown. 
Wings with a grey-brown tinge, darkest on the fore border; 
stigma dark brown; veins black-brown, neuration normal. V. d. 
Wulp, Sumatra Exped. Dipt., p. 17. 

This species is unknown to me. 


Tabanus jucundus, 7 ? , Walker. 
CRs scitt ier ox 
List Dipt..,1,/p2Le7( 1848): 
Type @ and another specimen with no head, from Hongkong ; 


male and female from Pundaluoya, Ceyion (Green); females from 
Kohat, Punjab, and India; all in Brit. Mus. coll. 


In Howlett coll. females from Igatpuri, W. Ghats, Bombay 
Presidency ; from Mussoorie, N. India, about 7,000 ft. high. 


Walker described the male only. 


A well-marked small black species with two very distinct 
lateral grey stripes on the thorax and with a grey median stripe 
on the abdomen, starting from the third segment and continued to 
the sixth segment, two lateral stripes commence on the first seg- 
ment but terminate abruptly on the second segment. Wings clear. 
Legs reddish yellow. Antennae red. The large protuberant 
frontal callus. with the short linear extension distinguishes this 
species. It differs from Tabanus hilaris, Wik., by the presence of 
the lateral stripes confined to the first two segments, by the more 
distinctly striped thorax, and by the shorter linear extension of 
the frontal callus which is darker and larger, and by the narrower 
forehead anteriorly. 

Length males 13 mm., females 10-11 mm. 

9. Face covered with whitish tomentum and with white 
pubescence. Beard white. Pa/pi stout, ending in a short apex, 
pale yellow in colour covered with white hairs, a few black hairs 
are also visible. Antennae: the first joint pale with white pubes- 
cence, black at the upper edge, the second very small, pale red, the 
third red with very obtuse tooth, apex reddish brown. Forehead 
broad, narrower anteriorly, about four times as long as it is broad 
at narrowest end, covered with greyish tomentum and with some 
patches of black pubescence; the frontal callus is large, almost 
square, entirely filling up the space between eyes, tuberculous, 
rising above the plane of the head, dark blackish brown, shining 


d 


1gII.| G. Ricarpdo: Revision of the spectes of Tabanus. 165 


with a short fine line proceeding from it. Eyes with three cross- 
bands. 

Thorax with dark pubescence except on the broad grey stripes 
where it is whitish, sides grey, so that the thorax might be called 
four-striped, pubescence below black, breast covered with long 
white hairs. Scutellum: the two grey stripes are continued and end 
on its posterior border. 

Abdomen blackish, reddish brown anteriorly, the pubescence 
black, white on the stripes; sides grey, covered with white pubes- 
cence ; under side with white-haired segmentations. Legs reddish 
yellow, the fore femora darker, all with grey tomentum, the fore 
pair with black, the others with white pubescence, the fore tibiae 
with thick white pubescence on basal half, elsewhere black, apices 
of fore tibiae and tarsi black. Inthe specimen from Bombay all 
the femora are darker. 

Wings clear with small yellow stigma, veins dark. 

@. Identical. The eyes with large facets, the lower third 
consisting of small ones continued as a very narrow border to back 
of head; the cross-bands are visible in the type. 


Tabanus puteus, ~ ?,n. sp. 
(Pie xin ie, -Er\) 
Type o from Velverry, Ceylon, and another from Trincomalee. 


Type @ and another from Hot Wells, Trincomalee, one ? from 
Colombo, all from Yerbury coll.; ‘not uncommon” [note by 
donor]. 


A small brown species with a dusky mahogany-brown abdomen 
which has a straight median grey stripe. Antennae yellow. Legs 
brownish. Wings tinged with brown on the fore border reaching 
the third longitudinal vein. Forehead of 2? narrowest on the vertex. 
Length Ir mm. 

9. Face covered with yellow-brown tomentum and with a few 
dusky hairs. Beard scanty, brown. Palpi small, stout, the same 
colour as face, with black pubescence. Antennae: the first two joints 
pale yellow with some black hairs, the third bright yellowish red, 
narrow, with no tooth, only a small angle marking its usual 
position, the second joint half as long as the first one. Forehead 
broad, narrower at the vertex, about three times as long as it is 
broad anteriorly, the same colour as the face, frontal callus large, 
almost square, reaching the eyes with a thick short line proceeding 
from it, in all the specimens there is a deep depression on the 
posterior part of the rather prominent callus which is yellowish 
brown in colour. Thorax blackish brown with the sides reddish, 
dorsum with thick short pale yellow hairs (in type rubbed off), on 
sides with yellowish brown pubescence. Scutellum appears rather 
lighter in colour, with the same very short pubescence. Abdo- 
men narrow, no spots, the anterior segments more yellow-brown, 
the apex darker, the short yellowish hairs appear on the grey 


166 ' Records of the Indian Museum, [ViOr. IVs, 


stripe and on sides, but short black pubescence covers the dorsum. 
Legs dull brown or yellowish brown. 

Wings grey, the yellowish brown shading extending in a fainter 
shade along the veins but most intense on the fore border, stigma 
brown, veins yellowish. 

@”. Similar, the eyes with large facets, only the lower part 
bordering cheeks with small facets which do not extend to the 
vertex. The figure of head is taken from a hadly preserved speci- 
men, two ¢ specimens lately added to Brit. Mus. coll. have the 
whole forehead shining brown. 


Tabanus immanis, ? , Wiedemann. 


Auss. Zweifl. Ins., i, p. 123, 17 (1828); V. d. Wulp, Sumatra 
Exped. Dipts; 10521 (1881) -.7d Notes Weyden Museum) vito pajae 
22 (1885). 

Olive-green. Abdomen rusty brown with an indistinct paler 
stripe; legs black; @ 84 lines. From Java. 

Antennae black. Palpi grey brownish. Face mouldy greyish 
coloured ; forehead grey, with a brown shining stripe, very little 
wider below. Thorax olive-green, yellowish in certain lights. Breast 
sides mouldy greyish white. Abdomen in certain lights with a 
paler median stripe and almost square side spots; side borders 
themselves yellow; under side brownish, side borders broadly, the 
segmentations narrowly, yellowish. Wings a little tinged, fore 
border and border of veins, and apex yellowish. Halteres yellowish. 
Femora mouldy greyish, knees and base of fore tibiae rusty brown. 
In the Leyden Museum. Wiedemann, Auss. Zweifl. Ins., i, p. 123. 

A female from Silago (Middle Sumatra) in June. ‘This is 20 
mm. iong. The eyes are naked, and with very fine network; the 
frontal band is narrow, posteriorly a little broader, in the middle 
with a shining black-brown, below broader, stripe. The third joint 
of antennae has a distinct tooth. Palpi with light ashy grey 
tomentum, ending in a point. Beard as well as the pubescence of 
fore breast and fore coxae light grey. The pale median stripe of the 
abdomen consists of triangular spots; under side red-yellow, with 
a black-brown apex. The neuration of wings is normal, the first 
posterior cell at its opening a little narrowed. Perhaps Tabanus 
yubtcundus, Macq., Dipt. exot., Suppl. i, p. 32, 64, Suppl. iii, 
10, and Suppl. v, 27, is only a somewhat paler variety of this 
species. Van der Wulp, Sumatra Exped. Diptera, p. 16. 

Several specimens, females, all from Sumatra. ‘The eyes are 
bare, bronze-coloured, and after being moistened, without cross- 
bands. IJd., Notes Leyden Museum, vii, p. 71. 

This species is unknown to me. 


Tabanus bubali, Doleschall. 
Natuurkund. Tijd. Nederl. Ind., x, p. 407, pl. iii, fig. 3 (1856). 
A reddish brown Tabanus, eyes very black, antennae black, 


1g1r.] G. Ricarpo: Revision of the species of Tabanus. 167 


forehead like mouth of a horse (sic), greyish white, the abdominal 
dorsal stripe white. Wings hyaline. Length 12 lines. Habitat 
in Java, hostile to cattle. Doleschall, /.c., p. 407. 


It is doubtful whether it will ever be possible to identify the 
species from this description; the figure proves the description was 
from a female, but otherwise is of little use. 


Grour VIII. 


Forehead with one square, narrow, or oblong callus always 
prolonged towards the vertex as a more or less narrow line; four 
to ten times as long as it is broad. 

Species with median or lateral spots or both on abdomen, not 
usually forming a continuous stripe. The subgroup represented 
by Tabanus fumifer, Wlk., contains species distinguished by very 
natrow foreheads and frontal calli, usually reddish brown or red- 
dish yellow in colour : this is the most difficult group in the Oriental 
Region, the species being very nearly allied, and apparently dis- 
tinguished by small, often variable characteristics; they are rather 
large or medium-sized species. Tabanus signifer, Wlk., is the 
type of asmall subgroup easily distinguished by their prominent 
large abdominal pale spots. 

Tabanus rufiventris, Fabr., feldervi, V. d. Wulp, and leucosparsus , 
Bigot, are not included in the table being unknown to me. 

Tabanus triangularis, V. d. Wulp, in the Austro-Malay Fauna, 
must be nearly related to Tabanus brunneus, Macq., but the legs 
must be lighter judging from the description. 


Tabanus amaenus, ~, Walker. 
List Dipt., i, p. 163 (1848). 
Tabanus clausacella, 7, Macquart, Dipt. exot., Suppl. v, p. 
45 (1855). 
Tabanus administrans, @, Schiner, Reise der Novara Dipt., 
p. 83 (1868). 


Tahanus signatipennis, 2 , Portschinsky, Horae Soc. Entom. 
Ross., xxi, p. 180 (1887). 

Bellardia sinicus, % , Bigot, Mem. Soc. Zool. France, v, p. 629 
(1892). 

The description of Tabanus clausacella is as follows :—Black. 
Thorax with ashy grey pubescence. Abdomen with red segmen- 
tations, the second segment testaceous at the sides. Antennae 
with the third joint testaceous at the base. ‘Tibiae white, with the 
apex black. Wings with first posterior cell closed. 

Length 6 lines, ~. Palpi of a yellowish white. Face, beard 
and forehead white. Antennae: the two first joints and the small 


168 Records of the Indian Museum. [Vion Ve 


divisions of the third, black, first division of the third yellow, tooth 
moderate. Thorax with tomentum of a rather yellowish grey; 
sides of a whitish grey. Abdomen with fawn-coloured incisions, 
terminated by yellow; second segment with a testaceous lateral 
spot reaching the two borders; under side with tomentum and seg- 
mentations whitish. I,egs black; tibiae of a yellowish white, black 
at the apex. Wings clear, stigma brown, fitst posterior cell closed 
near the border. From N. China (M. Bigot). Macquart, Dipt. 
CXOLs, Vs Datos 

This description by Macquart of his species is unusually in- 
correct as he overlooked the grey triangular spots of the abdomen, 
yet there is no doubt of the identity of the type, which is 
however a very faded dirty specimen, a male measuring 15 mm. 
Schiner could hardly identify his male specimen from the above- 
mentioned description, and gave it anew name. His description is 
as follows :— 

@, type, from Hongkong. Closely related to Tabanus man- 
darinus, and identical in many respects with it; the differences are 
as follows. 

Abdomen at bottom dark brown, the two basal rings as in 
the above-mentioned species, the following three rings with white 
tomentum, leaving four almost triangular spots of the ground colour 
free on the anterior border ; or these segments might be described 
as brown, each with an anteriorly indented, posterior band; 
under the white tomentum, in some lights less distinct, the ground 
colour is reddish yellow, the sixth segment on the under side black- 
ish, at the sides reddish yellow, the tomentum as in Tabanus 
mandaryinus. Antennae rusty yellow, the third joint at the apex 
black-brown, the middle and posterior tibiae are almost wholly red- 
yellow. In every other respect identical with Tabanus mandarinus. 
6 lines. One male. 

Though this species is so closely allied to Tabanus mandari- 
nus I do not think it possible to consider it as merely a variety 
Of it 

I have before me also, from the same locality, a female speci- 
men that possibly may be the female of T. administrans, the 
identity of all plastic characters at least encourages this opinion, 
but the abdomen is differently marked (and besides is not well pre- 
served), and the first posterior cell is narrowed but not closed. 
The forehead is fairly broad, whitish grey above the antennae, the 
callus black, broad at the base, then becoming narrower and con- 
tinued to the vertex as a line, ocelligerous spot present, but no 
ocelli visible. Schiner, Reise der Novara, Diptera, p. 83. 

Portschinsky described his type from Chinese Mongolia thus— 

This species is similar to Tabanus autumnalis, from which 
it is quite distinct, the first posterior cell of wing being closed with a 
long petiole, the size of body is less and it is much narrower in 
shape. 

Black with cinereous or grey tomentum, sub-naked or very 
shortly pubescent. 


1giI.] G. Ricarpo: Revision of the species of Tabanus. 169 


Head wholly white and white haired, frontal callus narrow, 
oblong, ovate, continued in a black line. Eyes naked, not banded. 
Palpi white. Antennae reddish, at apex more or less widely black, 
the third joint at base with distinct tooth. Thorax with three 
wide stripes (anterior half divided by a narrow ashy grey line) in 
middle with very short black hairs, at sides with yellow hairs. 
Abdomen black with three grey or white rows or spots, the middle 
spot triangular as in Tabanus autumnalis, rather wider at apex ; 
under side grey with middle stripe brown less conspicuous, each 
segment at apex narrowly margined with yellow. Squamae dark. 
Halteres white, at base black. Length 7} lines. Portsch., Horae 
Soc. Entom. Ross., xxi, 180 (1887). 

@. ‘The abdomen is blackish markedas in the male, with grey 
tomentose triangular median distinct spots and with lateral less 
distinctly defined ones, the segmentations very narrowly lighter. 
Thorax similar. ‘Tibiae all yellowish with apices black. Palpi 
reddish yellow, long ending in an acute point, with black pubes- 
cence. Antennae with black pubescence on first two joints, third 
long rather slender with obtuse tooth. Forehead narrow, very 
slightly narrower anteriorly, about seven times as long as it is 
broad, covered with yellowish brown tomentum and some scattered 
pubescence. Frontal callus oblong, not touching eyes, reddish 
brown or brown with a short spindle-shaped extension. Length of 
specimens from 15—18 mm. 

Three females from Wei Hai Wei have the first posterior cell 
not closed but very much narrowed at the border, but are in all 
respects similar, with the exception of the greater size of the lateral 
spots which take up nearly the whole width of the segments. 

a. The colour ot the abdomen is yellowish brown or darker 
brown, the lateral white tomentose spots often vellowish, ill defined, 
the second segment sometimes almost wholly yellowish brown with 
three grey spots; the median triangular grey spots always weil 
matked. Thorax dark with grey tomentum and stripes, these last 
often with short scattered yellowish pubescence. ‘The fore tibiae 
are usually red-yellow as well as the others, but sometimes ob- 
scurely so, and always on only the basal half. 

Head large, the facets sharply divided, the small ones reach- 
ing half-way up the frontal triangle anteriorly and continued pos- 
teriorly as a narrow band to the ocelligerous tubercle. Tength 
from 15—18 mm. 

The Macquart type came from China (on the label is ‘* China 
boreal’’); the Walker type from Hongkong; the Portschinsky type 
from Chinese Mongolia; and Schiner’s type from Hongkong. In 
Bull. Ent. France, 1887, p. Ixxvii, Tabanus amaenus is recorded 
from near Yeso, N. Japan, by Bigot. 

In Brit. Mus. coll. are specimens from Foo Chow and Swatow, 
and Shanghai (Walker); Formosa; and Japan (Dr. Myajima, 1909). 
In the Kertesz coll. along series of males and females from Formosa. 

The species probably belongs to the northern part of the 
Oriental Region and the southern part of the Paleearctic Region. 


170 Records of the Indian Museum. [Voru. IV; 


Bellardia simicus, described by Bigot from one specimen from 
China, is from the description no doubt this species described by 
Walker and others, the type is probably now lost. 

This species has certainly a great resemblance to Tabanus 
autumnalis, L,., but may be distinguished from it by the closed 
first posterior cell, and even in these specimens with this cell not 
closed but very narrow, the narrower abdomen and smaller size 
of insect, the narrower forehead seven times as long as it is wide 
(in T. autumnalts it is only five times) and the frontal callus nearly 
reaching the eyes will serve to distinguish it. 


Tabanus mandarinus, «7, Schiner. 


Reise der Novara, Dipt., 83, 8 (1868). 


Tabanus trigeminus, 2 , Coq., Proc. U. States Nat. Mus., xxi, 
310 (1808). 

Brown reddish; thorax at bottom very dark with white 
tomentum which makes it in certain lights appear quite white, 
and with five stripes, the middle one narrow, rather yellowish, 
divided by a shining brown line, in certain lights very indistinct, 
the outer ones situated quite on the sides of thorax, and uniting 
pair-like with the inner ones before the scutellum. Breast sides 
nearly red-yellow, the shoulders red-yellow. Scutellum brown, 
in certain lights whitish, the posterior border red-yellow, the 
pubescence chiefly dark, on the breast sides white, and in certain 
places tuft-like. Abdomen black, the first and second segment 
thickly whitish tomentose, leaving of the ground colour only four 
spots apparent, on each of the 3—5 segments a whitish yellow 
triangular median spot which (?) takes up the whole hind border 
with its broad base, and reaches the fore border with its apex, 
the side borders. also whitish. extending on each segment a little 
inwards, the segmentations narrowly red-yellow ; under side black- 
ish with yellow sides and segmentations, in certain lights it 
appears wholly white with the exception of a broad middle stripe. 
Head very large, the eyes very convex, the facets below and at 
the side very small, the upper ones large. Face very concave in 
the middle, whitish, proboscis short, black, projecting horizontally, 
palpi white-yellow, the end joint mussel-shaped. Antennae brown, 
the first joint appearing grey, the second rusty yellow, both quite 
bare, the first above projecting triangularly, the second with an 
upper thorn-like point, the third above with a shallow indentation 
and an obtuse angle. Legs black-brown, coxae and femora 
appearing white in certain lights and white haired, tibiae red-yellow 
on their basal half, more so on their outer than their inner side, 
tarsi black. Wings hyaline, the longish stigma brown, the first 
posterior cell closed, no appendix. Length 9 lines; one o from 
Hongkong. 

A magnificent species. Schiner, Reise der Novara, Dipt., 
D203. 


1git.] G. Ricarpo: Revision of the species of Tabanus. ey 


Schiner thinks it is nearly related to Tabanus spolvaius, Walker, 
but not the same. It has in fact no kind of resemblance to the 
Walker species. 


Brauer in Denkschr. Akad. Wien, xlii, p. 191 (1880), remarks 
that this species is very similar to Tabanus rectus, Loew, from 
Spain and Marseilles, but that the large facets of the eyes reach 
to behind the ocelligerous tubercle, and the zone of small facets 
above is narrow and not so broad as in Loew’s species. 


This species and Tabanus administrans, Schiner (now equal to 
Tabanus clausacella, Macq.), are related to Tabanus autumnalis, L., 
and Tabanus rectus, Loew, but the closed first or very much nar- 
rowed posterior cell will serve to distinguish both species. This 
species is distinguished from Tabanus c/ausacella by the absence of 
spots on the sides of the posterior segments and by its larger 
broader abdomen, and forehead not so narrow anteriorly. 


In Brit. Mus. coll. specimens @ and @- from Tientsin; ? 
specimens from Wei Hai Wei (‘‘ biting cows’’); from Chinkiang, 
Yangtze River, Foo Chow, Shanghai; Formosa and Japan. 

Tabanus trigeminus is recorded from Japan, six @ specimens 
measuring 17-18 mm., the first posterior cell narrowed at opening. 
A long series from Dr. Miyajima from Japan have this same 
peculiarity. The species belongs to both zoological Regions. 
It is rather a blackish than a reddish brown species, with well- 
marked grey triangular median spot and lateral spots on the first 
two or three segments. ‘Th> first posterior cell of wings usually 
closed, but occasionally only narrowed at opening. Face greyish 
tomentose with yellowish white pubescence. Palpi large, not very 
stout at base, ending in an obtuse point, reddish yellow with 
black pubescence. Forehead in 2 narrower anteriorly, nearly 
six times as long as it is broad, frontal callus oblong, reddish 
brown, not reaching the eyes, with a thick linear extension, the 
forehead covered with grey or yellowish grey tomentum and grey 
hairs, in many specimens a dark patch is present in the middle 
with black hairs. Thorax brownish black with three white stripes 
on dorsum and sides white, breast sides covered with greyish 
tomentum. Scutellum blackish brown with grey tomentum bordered 
posteriorly with grey hairs, on dorsum of thorax the pubescence 
is white and black, at sides black, below white. Abdomen black, 
the first two segments might be described as black, the first with 
three spots and the sides grey tomentose, the second with a large 
median and two lateral grey spots, on the third segment in some 
specimens there appear traces of a third spot, the segmentations 
usually narrower than as described by Schiner but in the specimens 
from Japan they are broader, grey tomentose slightly reddish 
yellow below. Wangs with first posterior cell closed with a short 
petiole, or very narrow at the opening, the specimens from Japan 
have it open, not closed, but very narrow; the @ from Tientsin 
the same. 


Length of specimens 15—2I mm. 


172 Records of the Indtan Museum. [| VOL. IV, 


Tabanus varicolor, Ricardo. 


Tabanus variegatus, Rondani, Ann. Mus. Civ. Genova, vii, p. 
458 (1875) (nomen bis lectum). 

Lengthr4mm. Antennae with the first two joints and the base 
of the third reddish yellow, the apex blackish. Forehead greyish 
yellow, anteriorly paler, the lineal callus red, shining; head above 
and below white and white haired ; palpi very pale, black haired. 
Thorax on dorsum dark brown, the sides, posterior corners and 
margin of scutellum reddish ; pleurae greyish and white haired. 

Abdomen with the first two segments reddish yellow luteous ; 
the following ones streaked blackish brown and ferruginous ; the 
apex black, a pale dorsal stripe present on the four segments before 
the last one; under side with the basal segments blackish, or black 
spotted, posteriorly luteous, with white tomentum, and short 
white hairs. Wings sub-limpid, the costal border luteous. Legs: 
anterior ones black, the tibiae at the base widely luteous, the pos- 
terior tibiae and knees luteous reddish yellow, femora and tarsi 
more or less blackish. Calyptera ferruginous. Halteres with club 
outside whitish, inside and the stalk brown-red. Rondani, Ann. 
Mus. Civ. Genova, vii, p. 458. 

The name of variegatus being preoccupied by a Fabrician species 
from N. America, I propose the substitution of varicolor for this 
species, the type of which came from Sarawak, Borneo, and was 
seen by me at Genoa. 

It is a narrow-bodied specimen measuring 17 mm., with a 
yellowish shining abdomen irregularly marked with black on all the 
segments except the first one, and has besides three series irregular 
shaped greyish spots median and lateral, indistinct; the blackish 
markings are situated between the anterior and posterior borders 
of segments, the seventh segment is blackest, with black hairs on 
the outer border; the dorsum appears finely punctuated all over, 
pubescence black, thickest on the apical segments; under side 
similar but darker. Thorax blackish with grey tomentum, reddish 
at sides, with black hairs at the sides; scutelluim blackish, red 
round the outer border. Legs yellow, with black pubescence, the 
femora reddish brown with white pubescence, the tarsi reddish 
brown. Face with grey tomentum and whitish hairs. Palpz 
yellow, thickly covered with black hairs, slender, ending in an 
obtuse point. Antennae reddish, dusky on the third joint, the tooth 
distinct, the first two joints with black hairs. Forehead with 
yellow-brown tomentum, narrow, anteriorly barely half as wide, 
about eight times as long as it is broad, the frontal callus yellow 
mahogany, narrow, oblong, not reaching the eyes. 

I have not seen any specimens identical with this isolated 
type. 

Tabanus rubicundus, 7 @? , Macquart. 
(Pl. sanietiee 12.) 

Dipt. exot., Suppl. 1, py 26072846); 7d., Suppl i), p: 170 

(2847); 2d., Suppl. v, p. 47 (as55) 


IgII.] G. RicaARDO: Revision of the species of 'Tabanus. 173 


Tabanus internus, Walker, ° , List Dipt., i, p. 164 (1848). 


Tabanus monilifer, 2, Bigot, Mem. Soc. Zool. France, v, p. 
654 (1892). [Atylotus.] 

@. Testaceous, abdomen with dorsal white spots. Legs tes- 
taceous ; anterior tibiae white at base. Length 7} lines. 

Palpi of a pale yellow. Beard and face yellowish white. 
Forehead yellowish with a testaceous callus prolonged as a line. 
Antennae: the first two joints are testaceous, the third is wanting. 
Thorax and abdomen testaceous (denuded), with traces of a yellow- 
ish down ; an abdominal stripe of triangular whitish spots; under 
side with whitish down. Anterior tibiae whitish in front, brown at 
apex. Wings yellowish, veins normal. 

Brom India. Macq. Dipt. exot.. Suppl. 1/p2 160: 


@”. Wehave since describing the female observed the male 
which differs inthe dark colour of the thorax and abdomen. Macq., 
i.c., Suppl. ili, p.170. From Java. Length 6 lines. Palpi yellow- 
ish. Beard white. Forehead and face white, a little yellowish. 
Antennae testaceous, third joint with the usual tooth and the last 
divisions black. Eyes: upper part brown, tower part black. 
Thorax with a yellowish grey tomentum, a testaceous stripe with 
greyish tomentum, passing above the insertion of the wings ; sides 
with ashy grey tomentum. Abdomen testaceous; segmentations 
a little yellow (seen sideways), seventh segment brown ; under side 
the same. Legs fawn-coloured, femora blackish, a little fawn- 
coloured at apex; tarsi brownish. Wings clear, exterior border and 
stigma yellow: veins normal. From Java. Macq., /.c., Suppl. v, 
p- 47- 

Macquart’s types are in Mr. Verrall’s coll. 

Walker’s type from Sylhet, Assam. 

Bigots type from N. Khasi, in a very dirty condition, two 
other females with it are from Sibsagar, Assam, and India. 

In Brit. Mus. coll. specimens (females) from Sylhet ; Chargola 
Valley, Sylhet; Dibia, Khasi Hills District (Chennell), and Khasi 
Hills (Godwin-Austen, Sladen). In Indian Museum coll. females 
from N. Khasi and Assam. 

This species is distinguished from the species in Tabanus 
fumtfer subgroup by its smaller size, broader, almost parallel fore- 
head and reddish yellow colour. 

A medium-sized reddish yellow species, the median triangular 
spots on abdomen small and often indistinct, the pubescence on 
dorsum scanty. Antennae reddish, rather wide at their base. 
Legs reddish brown. Length 17-18 mm. 

Face covered with yellowish brown tomentum and with white 
pubescence. Beard yellowish white. Palpi reddish yellow with 
some grey tomentum and rather thick black pubescence, in shape 
long and slender, ending in alongapex. Antennae deep red, darker 
at the apex, the first two joints with some black pubescence, 
the tooth of third joint prominent. fovehead rather narrow, about 
six times as long as it is wide, very slightly narrower anteriorly, 


174 Records of the Indian Museum. [VO1zIVE 


the same colour as the face; frontal callus oblong, brownish, not 
reaching the eyes, the linear extension narrow. Thorax obscurely 
reddish brown, sometimes blackish, stripes indistinct, covered with 
some yellowish brown tomentum and a few hairs of the same colour 
and with scattered black pubescence, sides redder with black hairs ; 
breast black with grey tomentum and obscure yellow pubescence. 
Abdomen reddish yellow, the last three segments slightly darker, 
the small triangular grey tomentose spots with yellowish pubes- 
cence are present from the first to the fifth, sometimes reaching 
the sixth segment, most conspicuous on the third, fourth and fifth ; 
the dorsum with rather dense but inconspicuous black pubescence ; 
under side with lighter segmentations, and yellowish white pubes- 
cence. Legs: femora reddish brown with grey tomentum below 
and with yellowish pubescence, tibiae reddish yellow, the fore pair 
darker at apex with yellowish hairs on the paler part, the tarsi 
reddish brown with black pubescence. Wangs grey, tinged with 
brown on fore border and round veins, veins yellowish brown, an 
appendix is present only in the Bigot type and in a female from 
Khasi Hills. 

The male type (Macquart) has no sign of median grey spots 
on abdomen, and may possibly prove not to belong to this species, 
which seems otherwise confined to India and Assam. 


Tabanus stantoni, 2 , n. sp. 
CEA Say cast oes 2) 


In Brit. Mus. coll. a long series from Batu Tisa, Labuan 
Padang, Federated Malay States (Pratt, Stanton), others from 
Selangor, and Kuala Lumpur, Federated Malay States. 

A large reddish yellow broad-bodied species, distinguished 
from Tabanus fumifer, Wik., by the greater width of the forehead, 
anteriorly never more than one-third narrower than it is at vertex, 
in I’. funufer the forehead begins to get narrower half-way from 
the vertex, in this species it is hardly perceptibly narrower and 
only begins to be so just before it terminates. Thorax has a 
paler appearance. Abdomen more reddish yellow. Length 
19$—25 mm. 

Face with yellowish hairs. Palfi yellow with black pubes- 
cence, not very stout at base, ending in an acute point. Forehead 
eight to nine times as long as it is broad and barely a third 
narrower anteriorly, nearly parallel, frontal callus long and 
natrow, not touching the eyes, slightly club-shaped. Antennae 
reddish, with the third joint black. Thorax brownish with in- 
distinct stripes, covered with brownish yellow tomentum, 
appressed fulvous hairs and some black pubescence. Abdomen 
appearing reddish brown, more strictly reddish yellow, but the 
close black pubescence gives it the darker appearance and with 
some brownish yellow tomentum makes it appear not uniform in 
colour, the median spots often indistinct, are yellow haired, short, 


rgir.] G. Ricardo: Revision of the species of Tabanus. 175 


triangular; under side with thick yellow pubescence at the sides 
and on seginentations, leaving a broad dark median stripe. Legs: 
femora blackish with grey tomentum, the tibiae obscurely yellow- 
ish or reddish brown, apices and all tarsi black; the pubescence on 
coxae and femora yellowish, elsewhere black. Wings large, tinged 
yellowish brown or almost clear with no appendix, veins blackish 
on fore border, thin brown, the first posterior cell widely open. 


Tabanus indianus, 7 @,n. sp. 


2? Tabanus mentitus, 7 , W\k., List Dipt., i, p. 162 (1848). 
In Brit. Mus. coll. 


Type ? and a series from Kadra, N. Kanara, S. W. India 
(Bell). Others from Formosa in Dr. Kertesz’s coll. Type ~ and 
a series from Formosa in Dr. Kertesz’s coll. 


A reddish brown species with a long narrow abdomen, distin- 
guished from others of the fumifer subgroup by the distinct pale 
yellow fore tibiae with white hairs of the female, and by the longer 
narrow white-haired triangular median spots of abdomen. Femora 
with black pubescence. Length of type ? 20mm., others from 
17—22 mm. 

@. Head wider than thorax. Face covered with greyish 
tomentum, with a few white hairs. Beard white. Palpi very simi- 
lar to those of Tabanus malayensis, n. sp., with long obtuse apex, 
only slightly stouter at base, yellow with thick black pubescence. 
Antennae reddish brown, darker at apex, the first two joints and 
apex of tooth with black pubescence. Forehead covered with 
yellowish brown tomentum-eand with some black hairs, about eight 
times as long as it is wide, only slightly narrower anteriorly 
(barely a third) than at vertex: frontal callus reddish brown, 
long, narrow, not reaching the eyes, with long linear extension. 
Thorax blackish with some grey tomentum and with black pubes- 
cence and fulvous appressed hairs, sides with black hairs ; breast with 
reddish white pubescence. Abdomen reddish brown, the median 
triangular white-haired spots are present from the 2-5th segments, 
with traces of one on the first and sixth segments, their apices 
almost reach the anterior border of each segment; pubescence on 
dorsum black, thicker at apex; under side lighter reddish brown 
with white pubescence, which is also present on sides of abdomen 
except at the apex where it is black. Legs dark reddish brown or 
blackish brown, coxae with long white hairs, the femora with grey 
tomentum and black pubescence, white on the under side of the 
middle and posterior pairs, tibiae pale yellow, black at their apices, 
the fore tibiae appear whiter on account of their white hairs; the 
middleand posterior pairs have some white pubescence below, 
otherwise it is black, and on all the tarsi. Wings long, faintly 
tinged yellow-brown on fore border and along veins, stigma 
yellowish brown, veins brown, first posterior cell slightly narrowed 
at opening. 


176 Records of the Indian Museum. [ VoL. IV; 


The specimens from Formosa have the abdomen in females 
more reddish yellow, the segmentations paler with yellow hairs. 
Frontal callus a little broader and more pear-shaped. 

o. Redder in colour than the female, the spots on abdomen 
are smaller and the fore tibiae dull yellowish or obscurely red at 
base with only a few lighter hairs. Eyes large, the large facets 
occupy the upper half but do not reach the anterior border of the 
frontal triangle, nor the vertex where the small facets extend as 
a narrow border. Palpi yellow, with yellow pubescence and a few 
black hairs at apex. Length of males 17—20} mm. 

The type of Tabanus mentitus, Walker, a male from Foochow, 
China (G. T. Lay), is probably identical, judging from its similarity 
to the male from Formosa, but owing to its bad condition and the 
fact of its being a male, its identity must remain doubtful till the 
advent of more material. 

The blackish colouring sometimes surrounding the median 
abdominal spots is more distinct than usual, as oblong black spots 
on the second, third and fourth segments, and only on the third 
and fourth segments is a median grey spot visible, probably owing 
to denudation, the apex of the abdomen is blackish. 


Tabanus brunneus, Macq. 


Suites a Butions 1, p: 203-(1834)- 7d. Dipt. exot., 1, p. 132 
(1838). 

In Brit. Mus. coll. one female from Timor and one male from 
Java. 
In Indian Mus. coll. one female from Semangho, Selangor, ‘‘ on 
Catcles: 


Length 8 lines. 


Blackish brown. Palpi brownish. Face and forehead yellow- 
ish grey; the latter narrower than usual, velvety brown; elevation 
near the antennae longitudinal, and the usual line black. Antennae 
black, inserted a little tower than in other species. Thorax 
covered with a short thick, brown tomentum ; two brown stripes 
on the outer border; sides with yellowish hairs ; a yellowish space 
with black hairs under the base of the wings, abdomen with dorsal 
triangular white spots, the three first segments brown, the four 
others black ; a small yellowish spot on each side of the posterior 
border of the segments; under side yellowish, a hemispherical black 
spot on each segment. Legs black. Wings brownish, @. 

From Java, in my cabinet. Macquart, Suites a Buffon, 1, 
b.2203- 

Brown. Antennae black. Abdomen with triangular white 
spots. Legs black. Wings brown. Length 6 lines. 

2. Palpi brownish. Face and forehead greyish yellow, frontal 
callus narrow and little distinct from the line proceeding from it. 
Antennae black. Thorax black, with brown tomentum. Abdomen 
black; a dorsal triangular yellowish white spot as well as a spot 


) 


rgtt.] G. Ricarpbo: Kevision of the species of Tabanus. 177 


on the exterior border of each segment. legs black, with yellow- 
ish tomentum. From East India (M. Marc) Museum. An identi- 
cal specimen in my collection came from Java. Macquart, Dipt. 
Cxoty isp 1132: 

Type ¢ (the one from India I- believe) seen by me in Paris 
Museum. A medium-sized species distinguished from Tabanus 
tnternus, Wlk., with which I compared it, by the narrower forehead 
and frontal callus and by the spots on abdomen which are al- 
most hemispherical in shape, on the second to fifth segments, small 
and oblong on the sixth, abdomen dark brown in colour, the seg- 
mentations very narrowly yellowish, and the lateral margins more 
widely so, forming the spots mentioned by Macquart; under side 
brown with vellowish segmentations and yellow pubescence on them. 

Wings clear, tinged brown. Legs brownish, tibiae appearing 
obscurely reddish on middle and posterior legs. Forehead narrow, 
eight or more times as long as it is broad, narrower anteriorly, 
with a long narrow frontal callus and lineal extension. Antennae 
reddish brown, falp: rather broad, brownish yellow with black 
pubescence. Face with yellowish white pubescence. Beard the 
same colour. 

This is apparently a distinct species, distinguished by the 
almost wholly black legs, dark palpi and antennae and by the 
hemispherical spots of the dark blackish brown abdomen. 


Tabanus fumifer, 7 9, Walker. 
(Pl. xiv; fis. 142) 
Proc, linn. Soc., 1, p. 11-(1857). 
Types o and 2 from Sarawak, Borneo (Saunders coll.). 


In Brit. Mus. coll. specimens from Darien Tipus, Negui 
Sembilan, Federated Maiay States (Stanton coll.) ; from Sungei, 
Bessi, Selangor (Durham); from Batu Tiga, Selangor (Stanton 
coll.) ; Singapore (Ridley), Sumatra (Forbes). 

A somewhat variable species, in colouring, but distinguished 
by the extreme narrowness of forehead, becoming anteriorly only 
half the width of what it is at vertex. The abdomen varies from 
reddish yellow anteriorly, becoming darker at apex (type), to dark 
reddish brown with small median indistinct spots, sometimes not 
apparent, the black pubescence on dorsum always rather thick, 
sides with yellow hairs anteriorly (type), but some specimens 
have black hairs entirely. Legs blackish, the fore tibiae usually ob- 
scurely reddish, on their basal third or half with some yellow hairs 
occasionally reddish yellow on basal half, the middle and posterior 
pair obscurely reddish or reddish yellow, only black at their apices. 
Wings tinged brown all along the veins, but in some specimens 
only tinged very slightly. Length from 15—21 mm. ‘The female 
type 2I mm. 

9. Face covered with yellowish brown or grey tomentum and 
with some whitish yellow hairs. Beard pale yellow. Palpi large 


178 Records of the Indian Museum. DV oT. SEVe 


with obtuse apex, almost the same width throughout, yellowish 
red but densely clothed with black pubescence. Antennae reddish 
brown, the first two joints with black pubescence, tooth of third 
joint distinct. Forehead narrow, covered with yellowish brown 
tomentum, nearly eight times as long as it is broad, barely half as 
wide anteriorly as it is at vertex; frontal callus brownish, long, 
narrow, not reaching the eyes with a rather long linear extension. 
Thorax reddish brown or blackish with stripes hardly apparent, 
some greyish yellow tomentum on dorsum, the pubescence black, 
with some short appressed yellowish hairs, shoulders more red, sides 
with black hairs; breast covered with grey tomentum and with 
yellowish hairs. Scutellum identical with thorax. Abdomen red- 
dish brown, often more yellow-red anteriorly and darker at apex, 
spots usually present on each segment but often indistinct and 
indicated by a few yellow hairs only, usually small with short 
triangles ; under side reddish brown, the segmentations bordered 
with yellowish hairs ; sides of abdomen with black hairs and a few 
yellow hairs below or. with almost wholly yellow hairs. Legs: 
femora black covered with grey tomentum below, the pubescence 
black and short above, yellow and longer below, coxae black covered 
with grey tomentum and with thick whitish pubescence on the fore 
pair, the pubescence on the dark red tibiae black and rather thick, 
some yellow or white hairs on the fore tibiae at base which alone 
is reddish. Wings in type very distinctly tinged brown on all the 
veins, but this is fainter in many specimens and sometimes almost 
confined to fore border ; veins brown, stigma yellow. 

@. The male type has now lost part of abdomen, but other 
males in the Brit. Mus. coll. from Singapore and Sumatra appear 
identical. 

Beard and hairs on face in type brownish, in the other males 
whitish. The palpi yellowish red, with black pubescence. The 
frontal triangle reddish brown. Eyes with large facets on the 
upper part, not extending beyond the apex of the frontal triangle, 
reaching the vertex but gradually sloping away from it towards 
the centre of eye. Abdomen on its anterior half yellowish red, the 
apex darker, the sides bordered with a short fringe of black hairs, 
but on the posterior lateral angles of the segments are some yellow 
hairs ; under side reddish brown with yellow-haired segmentations. 
Wings in type deeply tinged yellowish brown, in other males 
yellow on fore border only, between the costal border and the 
first vein. . 


Tabanus malayensis, ? , n. sp. 


In Brit. Mus. coll. type @ and a long series from Darien 
Tipus, Negui Sembilan, Federated Malay States, collected by Dr. 
A. T. Stanton on a new road running through jungle in February 
and March, 1908, and others in 1909 by him at Batu Tiga, others 
from Kuala I,umpur, Selangor, Federated Malay States (Durham), 
and from Singapore (Ridley) 


IgiI.] G. RicarDo: Revision of the spectes of Tabanus. 179 


A species nearly allied to Tabanus fumifer, Wik., distinguished 
only by its colouring and usually smaller size (16$—-21 mm.), differ- 
ing in the bright yellow-haired median spots of abdomen which are 
larger and very distinct, with longer apices, and by the presence of 
lateral yellow-haired spots on sides of segments in form of isosceles 
triangles, usually present from the second to the fifth segments ; 
under side more yellow with yellow pubescence on the sides. The 
thorax and scutellum have a paler appearance with greyish yellow 
tomentum and more numerous appressed fulvous hairs. The 
tomentum on the femora is more yellowish grey. Forehead seven 
to eight times as long as it is broad, very narrow anteriorly as in 
Tabanus funufer with the same long narrow frontal callus. Antennae 
blackish brown, in some specimens reddish at base of third joint, 
the first two joints with some grey tomentum and black hairs. 
Palpi large, ending in an obtuse point, yellowish with dense black 
pubescence. Abdomen reddish brown, redder on the anterior seg- 
ments, the whole dorsum covered with thick short black pubes- 
cence, on the posterior borders of segments appear a few yellow 
hairs, the spots consist of a small almost square one on the first 
segment, on the following four they are triangular in shape with 
short apices which do not reach the fore border of segment, those 
on the third and fourth segments usually with the widest bases, 
on the fifth appear traces of a small triangular spot, all have a 
yellowish or greyish ground colour covered with bright yellow 
hairs, sides of segments with spots as described above ; under side 
covered with yellowish pubescence leaving a broad central black 
stripe apparent. Legs similar to those of Tabanus fumifer, W1k. 
Wings grey, very slightly tinged brown, chiefly on the fore border, 
stigma yellowish, veins brown, the first posterior cell slightly 
narrower at opening. 


Tabanus factiosus, 7 @ , Walker. 


Proc. Linn. Soc. London, iv, p. 102 (1859). 


Type ~ from Celebes. Type ? from Makessar, Celebes, both 
from Saunders coll. 


‘Two females from Negros, Philippines (Whitehead). 


@. This large reddish brown species is very nearly allied to 
Tabanus fumifer, Wik., indeed the female seems only distinguished 
from the latter species by its larger size and by the fore tibiae being 
more largely yellowish or white, for quite two-thirds of their length, 
and by the wings being nearly clear, even the space between costa 
and first longitudinal vein being only faintly yellow. The thorax 
appears lighter, having more grey tomentum and a few appressed 
yellow hairs. Abdomen large, broad, with grey triangular spots on 
every segment except the first and last ones, on the fourth, fifth 
and sixth the segmentations are lighter, colour of abdomen reddish 
brown. Length 23 mm. 


180 Records of the Indian Museum. [Vo.L. IV, 


@”. Male type is not mentioned in Walker’s description, but 
bears a label ‘‘ factious’’ in his handwriting. It is immediately 
distinguished by the eyes which have the facets almost the same 
size, those where the eyes join hardly perceptibly larger; this 
character at once distinguishes it from Tabanus fumifer. Beard 
blackish, with the same coloured hairs on face. Fore tibiae black. 
Wings deeply tinged yellowish brown. ‘Till further material is 
available, it is impossible to define this species more accurately. 


Tabanus dissimilis, 9, n. sp. 
(Pla xiveig <5.) 


Type ¢ from Selangor (Butler), a series from Singapore 
(Butler, Ridley), two @ from Silam, N. Borneo and Borneo. 


A species very nearly allied to Tabanus fumifer, Wik., but dis- 
tinguished from it in general appearance by the redder thorax, 
covered thickly with ashy grey tomentum, and with black pubes- 
cence so that it appears darker than in the Walker species, with no 
yellowish interspersed hairs, and by the clear wings, only the space 
between the costa and the first longitudinal vein being yellowish 
brown. In the less width of the forehead anteriorly with the frontal 
callus practically reaching the eyes lies the chief plastic difference. 

Length of type 21 mm., other specimens 19}—22 mm. 

Head large, wider than thorax. Palpi witha rather longer apex 
than in Tabanus fumifer. Forehead about ten times as long as it 
is wide, hardly more than half the width of the vertex anteriorly. 
Breast reddish with grey tomentum, and with black above and 
white pubescence below. Scutellum identical with thorax. Abdo- 
men very similar but anteriorly with traces of yellowish brown or 
grey tomentum, the spots indistinct, often absent, usually white 
haired ; under side reddish with grey tomentum and white pubes- 
cence : the lighter hairs on sides of abdomen below the black ones 
are white. Legs: the fore femora are reddish brown or black with 
grey tomentum, but with wholly black pubescence, only the 
middle and posterior pairs with white hairs below in some speci- 
mens, in the type they are black. No white hairs are visible on 
the fore tibiae, but a few on the hind tibiae below. 


Tabanus ignobilis, @ , Rondani. 
Ann. Mus. Civ. Genova, vii, p. 457 (1875). 
Length 15—20 mm. 


Antennae brown-red. Forehead fulvous, the linear callus 
reddish. Face white, with white hairs. Palpi very palely luteous, 
black haired. Thorax with the dorsum obscurely red and with 
darker stripes, the pleurae and breast whitish and white haired. 
Abdomen brown-reddish, on the dorsum with black spots, chiefly 
scattered posteriorly : the posterior margins of the segments 
narrowly pale; the median whitish stripe of spots; under side 


1g1t.| G. Ricardo: Revision of the species of Tabanus. 181 


brown-red the margins of the segments posteriorly with pale mar- 
gins and very short white hairs. 

Wings sub-limpid, the costal border brown yellowish, the 
stigma browner. Calyptera ferruginous. Halteres with the knob 
yellow, the stem red. Legs brown-red ; the apices of femora and 
tibiae and all tarsi blackish. Rondani, Ann. Mus. Civ. Genova, vu, 
p. 457. From Sarawak Province. 

Type ? in Nat. Hist. Museum,Genoa. Aspecimien @ in Brit. Mus. 
coll. from Sandakan, N. Borneo, compared with type, is identical. 

The type in good preservation, measures 20 mm., the black spots 
mentioned in original description seem accidental stains or marks, 
the median spots only commence from the third segment, the 
thorax and scutellum are covered with grey tomentum, being 
reddish in colour. 

A species nearly allied to Tabanus dissinulis, n. sp., distin- 
guished from it by its lighter redder thorax, abdomen and legs. 
The abdomen is shining, uniformly reddish with grey median spots 
from the second to the sixth segments, and with greyish narrow 
segmentations, the little pubescence present, chiefly black. 
Forehead, frontal callus and palpi similar to those of Tabanus dis- 
similis. Legs reddish brown, the tarsi and apices of tibiae darker, 
the fore femora on under side and hind tibiae at base with some 
white hairs, the pubescence otherwise black. The specimen from 
Sandakan measures 22 mm. ‘his species is not unlike Tabanus 
vubicundus, Macq., but is distinguished from it by the greater 
narrowness of the forehead anteriorly. 


Tabanus nexus, 2 , Walker. 

Proc: Winn Soc, 1, p.r10 (1875): 

Type 2 from Sarawak, Borneo (Saunders coll.). 

A large species very nearly similar to Tabanus ignobilis, 
Rondani, from which it is only distinguished by the wings being not 
quite clear, but deeply tinged with yellowish brown on the fore 
border along the veins, and in the discal cell, reaching beyond its 
borders. ‘The palpi are stouter, almost the same width throughout, 
so that their apices are obtuse. There are no white hairs on the 
hind tibiae, but the white pubescence on all the femora is distinct, 
thickest on the fore pair. 

Forehead about nine times as long asit is broad, and hardly more 
than a third narrower anteriorly, frontal callus not reaching eyes ; 
owing to its great similarity to Tabanus ignobilis, Rondani, this spe- 
cies is placed next to it though strictly speaking it should go under 
the heading of foreheads one-third narrower anteriorly in the table. 


Tabanus signifer, 7, Walker. 


Ins. Saund. Dipt., i, p. 452 (1856). 
Type o from China. 


182 Records of the Indian Museum. Von es 


A red-brown species easily distinguished by the prominent 
whitish tomentose triangular spots on the third and fourth abdo- 
minal segments, and with the same coloured spots on the lateral 
borders of the first four segments. Wings clear. Legs blackish. 
Tibiae reddish. Length 21 mm. 

Head large. Eyes with the large facets distinct, reaching the 
apex of frontal triangle, and the vertex, the small facets extending 
behind as a narrow border but not reaching the vertex apparently 
(eyes discoloured). Face covered with grey tomentum and with 
white hairs. Palpi yellow with black hairs. Beard white. 
Antennae incomplete, Walker does not describe them ; the first two 
joints reddish with black hairs. Thorax reddish brown with three 
grey stripes, sides with black hairs. Scutellum reddish brown. 
Abdomen conical, reddish brown, becoming darker towards the 
apex, the two triangular median spots have very short apices, and 
broad bases, the spots on lateral borders do not join them, a small 
whitish tomentose median spot is visible on the second segment at 
its base, pubescence black, but white on the spots; under side the 
same but with no median spots visible. Legs (incomplete) blackish 
with black pubescence, the tibiae reddish. Wangs yellow on costal 
border, stigma yellow, veins yellowish brown, appendix present. 

The species first named by Walker Tabanus apicalis, List 
Dipt., 1, p. 176, and later changed by him to Tabanus finalis (the 
first name being preoccupied), List Dipt., v, p. 258, is allied to this 
species, and is also a male, but differs in having three large white 
spots on the third, fourth and fifth segments and the cross-veins of 
wings are shaded ; it is a badly preserved specimen from unknown 
locality ; it may possibly belong to the American fauna. 


Tabanus significans, ?, n. sp. 


d 


Type 2 and another female from Kuala Lumpur, Federated 
Malay States (H. C. Pratt), in London School Tropical Medicine 
coll, and another female from Darien Tipus, Malay States, in 
Stanton coll. 

This well-marked species allied to Tabanus signifer, Walker, 
from China, is distinguished by the two well-marked grey tomen- 
tose median spots on the brown abdomen, by the white tibiae and 
wings slightly shaded on the cross-veins. Antennae red, forehead 
narrow. Length 18 mm. 

Face covered with whitish tomentum and with long white 
hairs, a band across the base of antennae and the subcallus on its 
outer borders with yellowish brown tomentum, between the 
antennae and the eyes the band bears some long brown hairs ; 
centre of subcallus covered with greyish tomentum. Beard white. 
Palpt pale yellow with black pubescence, stout, ending in a point. 
Antennae red, the first two joints paler and yellowish with black 
pubescence, the third joint broad but with a very slight tooth 
indeed. Forehead very uarrow, about ten times as long as it is 


tgiI.] G. Ricarpo: Revision of the species of Tabanus. 183 


broad, and a third narrower anteriorly, covered with greyish 
tomentum, the frontal callus reddish yellow, very narrow, almost 
cylindrical with a somewhat raised long lineal extension. Thorax 
reddish brown with short black pubescence, shoulders and sides 
covered with greyish tomentum which extends round the thorax 
as a narrow border posteriorly, pubescence on the shoulders black, 
on the sides white and long, extending round the scutellum which 
is the same colour as thorax but with grey tomentum, this with 
the white pubescence gives it a pale appearance, at least on its 
posterior half. Abdomen brown or red-brown, clothed with short 
black pubescence the grey tomentose spots are clothed with white 
pubescence and are situated on the third and fourth segments, the 
one on the latter is the largest, with a wide base and short apex, 
almost half moon in shape, not reaching the anterior border of 
segment, the other one is smaller, oblong, placed on the posterior 
border of segment, on the sides of the first and second segments 
are lateral similar spots, and on the fourth segment at side on 
posterior margin a small tuft of white hairs, traces of a few white 
hairs are visible in the middle of the second segment; under side 
brownish yellow with some whitish tomentum at sides and anteriorly. 
Legs: coxae and femora reddish with white pubescence on the 
former, and black on the latter, the middle and posterior femora 
are darker reddish brown, the tibiae white, black at their apices, 
with the pubescence to correspond, tarsi all deep black. Wengs 
slightly tinged with brown on the fore border and round trans- 
verse veins, stigma and veins brown. Halteres black 


Tabanus fuscomaculatus, 2, un. sp. 


Type female and four others from Sima, Myitkyina District, 
Upper Burma (Capt. Whitmore, I.M.S.), in Brit. Mus. coll., and 
another female from Sikhim in Indian Museum coll. 


A handsome stout dark reddish species, the abdomen with 
two large black spots on the second and third segments and the 
apex black. Antennae and palpi blackish. Legs blackish. Wings 
tinged with brown and with an appendix. Length of type 20 mm., 
others 1I97—214 mm. 

Face covered with greyish yellow tomentum, light yellow on 
the cheeks, some dark short hairs in middle of face and long soft 
pale yellowish hairs on cheeks. Beard pale yellowish. Palpi 
stout, much the same width throughout, ending in an obtuse point 
appearing black, but dark reddish, covered with greyish brown 
tomentum and with thick black pubescence, brighter red on the in- 
side. Antennae black, the first two joints red with black pubes- 
cence, the third joint broad at base with a distinct tooth. Fore- 
head narrow, about eight times as long as it is broad, slightly 
narrower anteriorly, about one third, forehead and subcallus 
covered with brownish yellow tomentum, the frontal callus black, 
narrow, club-shaped, not reaching the eyes anteriorly, with a long 
linear extension, 


184 Records of the Indian Museum. [Vor. IV; 


Thorax blackish, with brownish yellow tomentum, some ap- 
pressed yellow hairs chiefly along the lateral sides of dorsum and 
short black hairs scattered over dorsum; sides with yellow hairs, 
shoulders reddish with black hairs, no sign of stripes on dorsum. 
Scutelium similar to thorax, on posterior border with some long 
yellow hairs. Abdomen reddish, the last three segments wholly 
dull black, the median black spots very distinct, the one on 
the second segment largest, bluntly triangular, neither base 
nor apex reaching the border, the second one on third segment 
smaller and more irregular in shape, the segmentations on the 
first four red segments paler, yellowish ; pubescence on dorsum 
black, golden vellow hairs are visible on the lighter segmentations, 
chiefly at the sides, where they extend upwards, long and thick; 
under side reddish yellow with a broad median stripe and the 
apex black, pubescence the same but the golden yellow is more 
prominent. 

Legs appearing blackish, the coxae covered with yellowish 
tomentum and with long yellow hairs, the femora with some grey 
tomentum and yellow hairs on the upper and under side, otherwise 
the pubescence is black, tibiae obscurely reddish with thick black 
pubescence, tarsi with black pubescence but on the under side 
of the posterior tarsi it is deep red. Wings tinged on fore 
border and along the veins brownish, appendix short, not always 
present, all posterior cells widely open, stigma brownish yellow, 
veins brown. 


Tabanus leucopogon, 2 , Bigot. 
(PL esty si oes) 


Mem. Soc. Zool. France, v, p. 651 (1892). [Afylotus.] 

Type @ from India and another @ from Sikhim, in Indian 
Museum coll. A small species allied to Tabanus oxyceratus, Bigot, 
but distinguished from it by the large oval frontal callus with 
hardly any linear extension, and by the wings which are shaded 
round the veins and have an appendix. Length 15 mm. 

The type is a very faded poor specimen. 

Face with white tomentum and white hairs. Beard white. 
Palpi slender, long, not very stout at base, ending in a point, dull 
reddish appearing dark by reason of the thick black short pubes- 
cence. Antennae reddish brown. Subcallus and forehead covered 
with yellowish brown tomentum. Forehead short, wide, very 
slightly narrower anteriorly, practically parallel, about four times 
as long as it is wide, the frontal callus blackish, not reaching the 
eyes, oval or nearly square with hardly any prolongation, in the 
type with a fine short line about the length of the callus itself. 

Thorax brownish. Abdomen blackish brown, with white- 
haired median spots on each segment except the last two. Legs 
blackish, tibiae yellowish. Wings hyaline with brown shading 
round the veins, fore border and stigma yellow, veins brown. 


rgtt.] G. Rrcarvo: Revision of the species of ‘Tabanus. 185 


Tabanus discrepans, 9, n. sp. 
(PlR xiv, fie ize) 
Type ¢ and another from Ceylon (Yerbury coll.) 


A long narrow species in general appearance allied to the 
species of the fumifer subgroup and more especially to Tabanus 
indianus,n. sp., but distinguished from them by the wider forehead, 
parallel in this species, and by the larger less narrow frontal callus. 
Abdomen reddish brown with small median whitish spots. Legs 
and antennae reddish. Palpi yellow. Wings tinged yellowish 
brown. Length of type 20 mm. the other female 18 mm. 

Face covered with yellowish grey tomentum, and with some 
pale yellow short hairs, browner on cheeks. Beard whitish yellow. 
Paipi stout, ending in a point, yellow with black pubescence and a 
few white hairs intermixed. Antennae reddish, duskier at the tip, 
the first joint pale yellow with black pubescence, third joint of 
antennae with very slight tooth. Subcallus and forehead covered 
with darker yellowish brown tomentum, the latter with some 
black pubescence, parallel, about five times as long as it is broad, 
the frontal callus red-brown, oblong, not reaching eyes with a short 
but stout lineal extension. Thorax dusky brown, pale reddish 
brown at sides and on shoulders, dorsum with appressed short 
yellow hairs, sides with long black pubescence; breast covered 
with greyish yellow tomentum and with pale yellow pubescence. 
Scutellum same colour as thorax with some of the appressed 
yellow hairs on sides, in the centre with black hairs. Abdomen 
reddish brown with short black pubescence, on the paler segmen- 
tations some yellowish white hairs extending to the sides and up 
the lateral borders of segments, the grey tomentose median spots 
present from second to fifth segment, more or less triangular in 
shape; under side with wider whitish-haired segmentations, and 
with the black pubescence. Legs reddish, the coxae covered with 
greyish tomentum and with whitish pubescence, the fore and hind 
femora darker, the apices of fore tibiae and all tarsi blackish brown, 
femora and basal half of hind tibiae with vellow pubescence, else- 
where it is black. 

Wings tinged yellowish brown, the centres of the cells at apex 
and on posterior border lighter in type, not discernible in the other 
female, stigma and veins of fore border brown, other veins yellow- 
ish brown, type alone with rudiment of an appendix. 


Tabanus rufiventris, @ , Fabr. (mec Macq.). 


Syst. Antl., 96 (1805); Wied., Dipt. exot., p. 66 (1821) ; 2d., 
Auss. Zweifl. Ins., i, p. 118 (1828). 

Thorax dark, abdomen reddish, wings with a black costal 
border. Of the size of Tabanus bovinus. Head ashy grey haired, 
antennae black. Eyes golden; thorax dark, hardly ashy grey 
striped. Abdomen reddish, with a dorsal stripe of white spots. 


186 Records of the Indian Museum. [Vor. IV, 


Wings white, costal border black. Legs black; tibiae red. East 
India. Mus. Dom. Lund. Fabr., Syst. Antl., 96, 13. 

Thorax brown, white pruinose ; abdomen red, white spotted, 
wings with black costal border. 94 lines. From East India. 

Antennae brown, black at the base and apex. Head yellow 
haired. Forehead reddish with the usual shining stripe hardly 
wider below. Thorax with hardly perceptible linear-shaped grey- 
ish white stripes. Breast sides reddish. The white abdominal 
spots are situated on the posterior border of each segment, the 
side borders of these segments and the ventral segmentations are 
also white. Wings almost hyaline, brownish black on the border. 
Squamae brown. Halteres white. Inthe Royal Museum. Wied., 
Auss. Zweifl. Ins., 1, p. 118. 

This species judging from the description may be identical 
with Tabanus fumifer, Wik., but it is impossible to identify it with 
any other species in this group where the species are only divided 
from each other by very small differences, unless the type or the 
specimen determined by Wiedemann is seen. 


Tabanus dives, 2 , in parte, Rondani. 
Ann. Mus. Civ. Genova, vii, p. 457 (1875). 
? = Tabanus stantoni, n. sp. 


Type 2 in Genoa Museum seen by me, is the smaller species 
of two specimens belonging to different species, the larger one 
I believe to be the same as Tabanus univentris, W\k., and this one 
is either the same as 7abanus stantont, n. sp., or very nearly allied 
to it, only differing in the reddish brown abdomen having yellowish 
segmentations, the under side uniformly yellowish. Wings with 
lighter veins, yellow in colour, stigma the same, slightly tinged 
with yellow, no appendix. Face yellowish with yellowish grey 
hairs. Palpi large, stout, pale yellow with thick black pubescence, 
and thick yellow pubescence above. Forehead as in Tabanus 
stanton ,n. sp., the frontal callus yellowish mahogany and more pear- 
shaped, narrower anteriorly, about ten times as long as it is wide, 
the forehead with yellowish brown tomentum and black pubes- 
cence. Antennae incomplete, the first two joints yellow. Length 
18 mm. From Sarawak, Borneo. 


Tabanus felderi, 7 2, Van der Wulp. 

Notes Leyden Museum, vii, 78, 32; pl. v, fig. 7 (1885). 

? Tabanus bucolicus, 7 , Schiner, Reise Novara Dipt., 8£, 4 
(1868). 

Blackish; antennae, palpi, three first segments of abdomen 
(except the dorsal spots on segments 2 and 3), the posterior border 
of the following ones and the legs red; eyes naked; wings greyish 
red, with the angle of the third longitudinal vein omitting an ap- 
pendix. o@ @, length 22-23 mm. 


. 


tgtt.] G. Rrcarpo: Revision of the species of Tabanus. 187 

Head greyish white; face of the ~ ochraceous; on the front of 
the @ araised line, which is broader below. Antennae and palpi 
reddish yellow; third joint of the antennae on the upper part 
with a rather sharp tooth, slender towards the end, the tip brown. 
Palpi in the female as long as the rostrum, in the male much 
shorter. Eyes bare, bronze-coloured, even after having been 
moistened without cross-bands; in the male the facets of the unper 
portion large. Thorax and scutellum greyish black, with hardly 
any indication of longitudinal bands; pleurae cinereous with a 
yellow pile. Abdomen in the male pointed, in the female a little 
narrower towards the end, in both sexes reddish yellow; the first 
segment blackish below the margin of the scutellum, on the second 
il third segments a dark grey dorsal spot; the following segments 
fuscous, with pale reddish yellow hind margin; under surface of 
abdomen testaceous, with yellow hind margins of the segments. 
Legs reddish yellow; the tarsi a little darker; front coxae with 
a long pale yellow pile; hind tibiae in the male fringed with short 
hairs, the hairs of the outside black. Wings with a reddish grey 
tinge; the upper branch of the furcated cell angular at its base 
and with a recurrent vein. A single pair from Ningpo (near 
Shanghai) presented to Leyden Museum by Dr. Felder. V. d. 
Wulp, Notes Leyden Museum, vii, p. 78. 

Tabanus bucoiicus was described by Schiner from one male 
specimen taken at Hongkong, and from the description and the 
comparison of one male specimen in the Brit. Mus. coll. with no 
locality specified, it is possible that Wulp and Schiner described 
the same species, but as there are no females in any of the col- 
lections, the matter must be left doubtful for the present. I ap- 
pend all that Schiner says regarding his species. 

“Ts exactly the same as Tabanus intermedius, Egger, which, 
as the name 7. intermedius was used earlier by Walker for a 
North American species, I shell in future name Tabanus eggeri, 
[Brauer however retains the original name as he says Walker's 
species is doubtful; Kertesz in his last Cat. 1906 follows Schiner. | 
It will suffice to give the differences. The middle of the face in 
Tabanus bucolicus as well as the frontal triangle are snow-white, 
the remainder bright yellow, the beard is golden yellow, the legs 
are entirely bright reddish yellow, the tarsi hardly a little darker. 
In all other respects similar to Tabanus eggevi. 11 lines.” 

Brauer remarks at the end of his description of Tabanus inter- 
medius that Tabanus bucolicus has much larger facets of the eye 
(o) and these extend above to the ocelligerous tubercle, and thence 
almost entirely to hind border. 


Tabanus leucosparsus, ? , Bigot. 
Nouv. Archiv. Mus. Hist. Nat. Paris (3), ii, p. 203 (1890). 


Length 18 mm. 


Antennae incomplete, the first two joints brown; palpi, beard, 
face and forehead whitish grey, frontal callus and tubercle of 


188 Records of the Indian Museum. ~ [Vors ivy, 


vertex chestnut-coloured (ocelli indistinct), the former at the base 
wider and square, very much restricted in the middle, narrow, 
oval, prolonged above. Thorax of a pinkish or flesh-coloured grey, 
sides whitish. Abdomen black, with a median series of triangular 
spots, wide towards the bases, smaller and smaller towards the 
extremity, and on each side another series formed of narrow spots, 
all these spots are whitish; squamae brown; halteres pale fawn- 
coloured. Legs black, femora with a whitish tomentum below, base 
of tibiae widely whitish. Wings clear, no appendix. Eyes naked. 

Laos (Siam), one specimen, Bigot, Nouv. Archiv. Mus. Hist. 
Nat: Paris:(3), ai,-p) 2028: 

This species with black abdomen marked with spots may 
possibly be allied to Tabanus leucopogon, Bigot, from the descrip- 
tion of the frontal callus, but is a larger insect. 


Group IX. 


Forehead with one square, narrow, or oblong callus, always 
prolonged towards the vertex as a more or less narrow line, four to 
ten times as long as it is broad. Species with paler bands on the 
abdomen, spots are also sometimes present, large, medium-sized, 
or small species. 

Tabanus orientalis, Wied., Tabanus geniculatus and tncultus, 
V.d. Wulp, are not included in the table, being unknown to me. 
Tabanus geniculatus has a very narrow parallel forehead and narrow 
callus, Tabanus incultus the same but narrower anteriorly. 


Tabanus servillei, @ . Macq. 


Dipt. exot., i, p. 132, 13 (1838); Walker, List Dipt. Brit. Mus., 
Vv, p. 246 (1854). 

Tabanus pyrausta, 2 , Walker, Zoologist, viii, Appendix Ixv 
(1850)2 1d; Last Dipt) Bur. Mus. v. p2 246 (16524); 

Antennae black. Thorax fulvous. Abdomen black; with 
white segmentations. Wings red; the first posterior cell closed. 
Length ri lines. 

Palpi black, wide at the base. Face and forehead greyish yel- 
low ; the latter narrow with a black line, a little raised anteriorly. 
Thorax with greyish yellow tomentum. Abdomen black; the 
four first segments with a narrow yellowish white posterior band. 
Calyptera blackish, bordered with white. From E. India. M. 
Mare. Museum. One specimen from Javacccurs in M. Serville’s 
collection. _Macquart, Diptiexotas p. 132, 13: 

The specimen mentioned as from Java, is apparently the one 
with another female now in Mr. Verrall’s coll. in very bad condi- 
tion, but from the description and comparison of these with Walker’s 
types there is no doubt they are identical. Walker placed the two 


/ 


1g1I.] G. Ricarpo: Revision of the spectes of Tabanus. 189 


species one after the other in his List Diptera, v, without discover- 
ing their identity. 

His type (female) from Mr. Argent’s coll. and another male 
from Hon. EK. India Company’s Coll. are both from Java and well 
preserved, and from these the following notes are taken. 

Q. A large easily distinguished insect, the first posterior cell 
of the wing closed with a rather long petiole reaching the border ; 
blackish, the thorax covered with fulvous tomentum and the seg- 
mentations of the abdomen fulvous. Length 25 mm., the Mac- 
quart specimens 22 mm. 

Face covered with fulvous tomentum and with brownish pubes- 
cence. Beard brown, in one of the Macquart specimens it appears 
to be mostly golden yellow. Pali large, stout, black, with black 
pubescence. Antennae black, stout, the first two joints with black 
hairs, the third joint with a large tooth, and broad at base. 
Forehead and subcallus the same colour as face, the former very 
narrow anteriorly, at the vertex quite double the width, about ten 
times as long asit is wide, the frontal callus reddish brown, very 
long and narrow, almost reaching the eyes with a long stout lineal 
extension. Eyes as remarked by Walker with the facets a little 
larger above. The bright coloured thorax with some scattered 
yellow hairs, the sides with black hairs; breast same colour as 
thorax with long yellow pubescence and some brown hairs. 
Scutellum similar to thorax. Abdomen broad, the lighter segmenta- 
tion most apparent on the first four segments; under side with 
whitish segmentations. Legs wholly black, fore tibiae with some 
greyish tomentum, pubescence black. Wungs tinged deeply reddish 
yellow, stigma yellowish, veins yellow. 

@ identical. Eyes with the large facets hardly reaching 
beyond the apex of the frontal triangle, and the small facets not 
reaching the vertex. Length 21 mm. 


Tabanus crassus, 7 , Walker. 


Dipt. Saund., i, p. 50 (1850). 

Type male from E. India (Saunders coll.), and other males 
from Karwan, N. Kanara, India (Bell) (“ in bungalow ’’), Hong- 
kong (Walker), in Brit. Mus. coll. 

In Howlett coll. two males from Pusa, Bengal (‘‘ in bungalow’’) 
In Kertesz coll. males from Formosa. 

This species, of which curiously enough no females appear to have 
been collected, is an easily recognized blackish species, the first two 
segments of the abdomen covered with a hoary tomentum ; wings 
clear, the first posterior cell narrower at opening. Antennae reddish. 
Palpi and legs blackish. Length of type 19 mm., others I5—20mm. 

Face and subcallus covered with yellowish brown tomentum, 
the tormer with blackish brown hairs. Beard the same colour, 
thick. Palpi black with grey tomentum and blackhairs. Antennae 
dull reddish yellow, black on apex, the first two joints with black 


190 Records of the Indian Museum. [VoL. IV, 


pubescence, the third joint narrow, slender, with a small tooth. 
Eyes large, the large facets very distinct, the small facets hardly 
reaching the vertex. Thorax reddish brown or blackish brown, with 
yellowish brown tomentum and blackish pubescence. Scutellum 
the same. _Aldomen blackish brown or reddish brown, the first two 
segments reddish, densely covered with ashy grey tomentum, the 
segmentations very indistinctly paler, the pubescence on dorsum 
close, black, at sides black ; under side more reddish brown. Legs 
appearing black, but often dull reddish covered with thick close 
black pubescence, the femora and hind tibiae with longer black fringes. 
Wings quite clear, stigma hardly apparent, pale yellow, veins on 
fore border sometimes blackish, otherwise yellow, the first posterior 
cell in type slightly narrowed, in some others more distinctly so, no 
appendix. 

It isjust possible that Tabanus basalts, 2° , Macquart, may 
prove to be the female, but it is a very much narrower bodied 
insect with dark wings. 


Tabanus albocostatus, 2 , Bigot. 


Mem. Soc. Zool. France, v, p. 649 (1892). [Atvlotus.] 
Type ¢@ from India. 


No specimens of this species are in the collections mentioned 
in this paper. It is a medium-sized specimen distinguished by the 
lighter colour of the sides of thorax and of scutellum, the abdomen 
being a dull brown with very narrow pale segmentations, antenne 
(incomplete), palpi and beard dark, legs brown. Forehead rather 
narrow. Length 21 mm. 

Face and subcallus brown with yellowish brown tomentum, 
pubescence dark brown. Beard brown. Palpi dull reddish, 
covered with grey tomentum and with close black pubescence, long, 
not very stout at base, ending in an obtuse point. Antennae with 
the first two joints same colour as palpi, with black pubescence. 
Forehead brown with grey tomentum, raised, about eight times as 
long as it is broad, anteriorly reduced to almost half the width it is 
at vertex, frontal callus black, long, not reaching the eyes, its lineal 
extension stout, broad, ending in a finer point, not much longer than 
the callus itself which is oblong. Thorax reddish brown on dorsum, 
its sides redder, covered together with the shoulders with greyish 
tomentum which extends narrowly round the posterior border, 
scutellum covered with it also, pubescence of thorax now almost 
mil, a few white hairs present, on shoulders brown; breast brown 
with yellowish brown tomentum and brown pubescence. Abdomen 
dull brown, the segmentations very narrowly reddish yellow, with 
traces of a few white hairs in the middle, and white hairs are 
distinct on the sides of the first five segments, pubescence otherwise 
black; under side identical. Legs blackish with black pubescence, 
the tibiae more dull reddish brown. Wings tinged brown on fore 
border, veins and stigma brown, no appendix. 


tgtt.] G. Rrcarpdo: Revision of the species of Tabanus. IgI 


Tabanus explicatus, ? , Walker. 


List Dipt:, v, p. 328 (1854). 

Types, two females from Sikhim, E. Himalayas (Hooker) ; 
another female from Khasi Hills (Chennell coll.) ; two females from 
Coonoor, Central India. One male from Khasi Hills (Chennell coll.). 

A black species with narrow fulvous segmentations on the 
abdomen, yellowish white tibiae and hyaline wings with yellow veins 
and stigma. Length 21 mm. 

Face covered with greyish yellow tomentum and a few white 
hairs. Beard pale yellowish. Palpi reddish with thick black 
pubescence, almost the same width throughout, the apices obtuse. 
Antennae reddish, the third joint dusky, broad at base with a 
prominent tooth, the first two joints with black pubescence. 

Forehead and subcallus same colour as face, the former nearly 
seven times as long as it is broad and a third narrower anteriorly, 
the frontal callus blackish brown, shining, oblong, not reaching 
the eyes witha rather thick lineal extension. Thorax with some 
grey tomentum, reddish at sides, hairs on sides yellowish. Abdo- 
men blackish with narrow reddish segmentations on every segment, 
which in fresh specimens are covered with whitish hairs, often 
extending in the middle to almost a triangular spot, dorsum other- 
wise appears devoid of pubescence ; under side similar. Legs black, 
tibiae pale yellowish, black at apices, fore coxae densely covered with 
yellowish grey tomentum, femora and tibiae with white, apices of 
tibiae and tarsi with black pubescence. Wangs very slightly tinged 
yellow on second and third longitudinal veins. The specimens from 
Coonoor probably discoloured have more reddish brown colouring 
on the abdomen which has yellowish pubescence. ‘The male is 
identical with the female, the large facets occupy two-thirds of the 
eye, the small facets extending as a narrow border to the vertex. 


Tabanus nicobarensis, @ , Schiner. 


Reise Novara Dipt., p. 81 (1866). 


Black : thorax with some grey tomentum and with three paler 
stripes, only distinct in certain lights and then especially so anter- 
iorlv, sides also paler, the shoulders with white tomentum ; breast 
sides and scutellum ashy grey ; the short pubescence on the dorsum 
‘of thorax chiefly black or at least dark, but whitish on the sides of 
breast, with a black-haired stripe from the base of wings to the 
shoulders. Abdomen black-brown, the first segment whitish at 
the sides, all segmentations clothed with snow-white short hairs 
which are extended in the middle to small, blunt, triangular 
spots ; under sides identical ; viewed from behind towards the light 
a dark cinnamon-brown tomentum spreads over the whole abdomen 
which, in comparison with the thorax viewed in the same light and 
appearing very light grey, stands out in strong contrast. Head 
black, with whitish tomentum and shining white pubescence; the 


192 Records of the Indian Museum. [VoL. IV, 


forehead anteriorly likewise whitish, but with a bright brassy 
yellow glimmer ; the shining black callus situated rather far from 
the antennae, oblong, filling up the entire width of forehead, its 
posterior border with two incisions so that points appear at each 
side, but from the middle a shining black line proceeds towards the 
vertex, which is at first wider and from behind to the middle deeply 
incised fork-like ; in the place where usually the ocelli are situated 
appear two shining black raised little lines near each other; the 
remaining surface of the forehead is grey tomentose, as is the rather 
distinctly concave hind part of head; proboscis black and short, 
the labella long. Palpi black-brown, with paler tomentum, broad 
at the base then gradually ending in a point, curved, a little shorter 
than the proboscis. Antennae black, the first projecting above; 
the second very short with a thorny point, the third on the upper 
side with a shallow incision, the angle only slightly projecting. 
Eyes naked. Legs black, the tibiae by reason of the paler thick 
tomentum at the base, appearing whitish, the fore tarsi unusually 
broad and short. Wings tinged deep smoky brown, the centres of 
cells rather paler, fore border darker, the first posterior cell widely 
open, no appendix. Length 7 lines. Four females from Nicobar 
Islands, one from Nankauri, the others from Sambelong. The 
species could only be compared with Tabanus brunneus, Macq., but 
there is no trace of black spots on the abdomen, neither of median 
spots, and the sides are not whitish yellow. Tabanus brunneus 
is also larger—8 lines. Schiner, Reise Novara Dipt., p. 81. 

Two females from Nicobar Islands, 1904 (Bingham coll.; col- 
lected by G. Rogers, Esq., Deputy Surveyor of Forests), in British 
Museum coll. ; 

In these specimens the wings are clear, veins brown. ‘The 
forehead is about eight times as long as it is broad, and a third 
narrower anteriorly. ‘The frontal callus large, a little protuberant, 
reaching the eyes, with a lineal extension. Length 16-163 mm. 
The comparison with Tabanus brunneus appears to me mislead- 
ing, but no doubt Schiner had not had the advantage of seeing the 
type. 


Tabanus justorius, Rondani. 

Ann. Mus. Civ. Genova, vii, p. 455 (1875). 

From Province Sarawak, Borneo. . 

Length 15—17 mm. 

@. Body black; forehead brown-grey, callus linear, black, 
shining ; face white. 

Antennae black. Palpi obscurely brown. ‘Thorax, particularly 
at the pleurae, white, pruinose. 

Wings blackish, the two basal cells in the middle, and a stripe 


under the third longitudinal vein almost hyaline: costal border 


blacker ; stigma black, appendix present. Halteres blackish, with 
the knob largely whitish. 


Igtt.] G. Ricarpo: Revision of the species of Tabanus. 193 


Abdomen black, the posterior segments with white bands, 
which are a little dilated in the middle of dorsum ; under side with 
four white bands. Legs wholly black. Rondani, Ann. Mus. Civ. 
Genova, Vii, p. 455. 

Type female and another fron. Sarawak seen by me in Nat. 
Hist. Museum, Genoa. No specimens are present in the collections 
named in this paper, but an easily identified species. 

Blackish with white-haired bands on abdomen, legs and 
antennae black. Length 17 mm. 

Face grey with white pubescence. Palpi yellow, with thick 
black pubescence. Antennae now incomplete, the first joint with 
black pubescence. ‘The frontal callus long on the narrow forehead, 
with a short linear extension. Forehead quite seven times as long 
as it is wide and nearly two-thirds narrower anteriorly. Subcallus 
with grey tomentum. Thorax blackish brown with grey tomentum, 
and very indistinct stripes. Scufedlum brown with grey tomentum. 
Abdomen rather long and narrow, the first segment broadly whitish 
grey tomentose, second, third, fourth with whitish grey haired 
narrow bands enlarged in the middle, the fifth and sixth with white- 
haired median spots, colour of abdomen blackish brown with black 
pubescence ; under side similar. Legs brownish black, the pubes- 
cence black. Wings deeply tinged with brown, the discal, second 
basal cell and apex almost clear, appendix present. 


Tabanus khasiensis, 2 , Ricardo. 
Ann. Mag. Nat. Hist. (8), iti, p. 487 (1909). 
Type @ and two other females from Khasi Hills, Assam, 
T,000—3,000 ft. One female from Meerut (Indian Museum coll.). 


A black medium-sized species with white bands on the abdo- 
men. Antennae, palpi and legs blackish. Wings hyaline, tinged 
with brown. Length 15-16 mm. 

Nearly allied to Tabanus justorius, Rondani, from Borneo 
but distinguished by the more hyaline wings, by the forehead being 
the same width throughout, and by the darker palpi. 

Face covered with greyish tomentum in the middie, on the 
cheeks and subcallus with vellowish brown tomentum, the pubes- 
cence black. Beard black. Palpi on outside appearing blackish 
covered with grey tomentum and with black pubescence, on the 
inside obscure reddish yellow, long and pointed, stouter at base. 
Antennae dull reddish black or black, the first two joints with grey 
tomentum and black pubescence, the tooth of third joint small. 
Forehead same colour as subcallus, the same width throughout, 
with a few black hairs, nearly seven times as long as it is broad ; 
the frontal callus oblong, narrow, brown, not reaching the eyes, 
furrowed in the middle, with a long rarrow linear extension. 
Thorax, scutellum and abdomen brownish black, the former with grey 
tomentum and traces of two stripes, the pubescence black, a few 
appressed white hairs anteriorly ; sides with black hairs, a tuft 
of white hairs behind the base of wings; breast black with grey 


194 Records of the Indian Museum. Vor ay 


tomenttim and black and white hairs. Scutellum on outer border 
covered with grey tomentum, pubescence black, white on the outer 
grey border. Abdomen with distinct white-haired bands on every 
segment except the seventh, enlarging slightly in the middle to 
half-moon spots and wider at the sides; under side similar. Legs 
black or brownish black, with black pubescence, coxae with white 
pubescence, and a few white hairs on hind femora below, the tibiae 
are sometimes obscurely reddish brown at base. Wings clear, 
tinged faintiy with brown, in one specimen it is more distinct and 
borders the longitudinal veins. 


Tabanus sanguineus, 2? , Walker. 


Dipt. Saund., i, p. 54 (1850). 

Tabanus assamensis, 2 , Bigot, Mem. Soc. Zool. France, v, p. 
653 (1892). [Atvlotus.] 

Walker type @ from Java (Saunders coll.). 

Bigot type @ , faded, from Assam. 

In Brit. Mus. coll. females from Kelantan, Malacca; and 
Hongkong. 

In Howlett coll. from Pusa, Bengal; and Khasi Hills, 
I ,o00O—3,000 ft. high. 

Others sent for identification from Darien Tipus and Negui 
Sembilan, Federated Malay States; Kuala Lumpur and Jelebu, 
Federated Malay States ; Lushai Hills, Burma; and Formosa. 

In Kertesz coll. two females fron: Sumatra. 

This species has a wide distribution and is a handsome, easily 
recognised species, but very liable to vary in appearance owing to 
denudation on abdomen. In well-preserved specimens the abdo- 
men appears blackish or reddish brown, with well-marked whitish- 
haired triangular spots on each segment, and with white-haired 
posterior borders, widest at sides. Forehead wide with large club- 
shaped callus. All the tibiae white. Length of type 174 mm., 
others 18—21I mm. 

Face covered with light greyish tomentum and with rather 
thick silvery white pubescence. Palfi yellow or light reddish with 
white pubescence and some black hairs at apex, short, stout, 
ending in a point. Beard silvery white. Antennae reddish brown 
with distinct tooth, the first two joints with black pubescence, the 
third joint long and slender. Forehead broad, short, rather more 
than four times as long as it is wide, almost the same width 
throughout, darker than the face with chiefly black pubescence, 
the frontal callus red-brown, large, oblong, almost reaching the 
eyes with a stout spindle-shaped extension. Thorax reddish brown 
or blackish brown, with grey tomentum on dorsum and chiefly 
black pubescence, stripes on well-preserved specimens distinct, 
sides with black hairs, round root of wings tufts of white hairs. 
Scutellum same as thorax, bordered with a short fringe of white 
hairs. Abdomen reddish brown, often blackish brown, with white 


rg1I.] G. Ricarpo: Revision of the species of ‘Tabanus. 195 


haired segmentations on a grey tomentose border, enlarged in the 
middle as broad, triangular spots, and at the sides wider and more 
distinct ; under side blackish with the sides broadly white haired. 
Legs black, all tibiae with the exception of the apices pale yellow or 
white, the pubescence on coxae, femora, pale part of tibiae white, 
elsewhere black. Wings clear, stigma very pale yellow, veins 
brown, all posterior cells widely open. 


Tabanus oxyceratus, 2 , Bigot. 


Mem. Soc. Zool. France, v, p. 652 (1892). [A?ylotus.| 

Type 2? from India. 

In Brit. Mus. coll. females from Cheena Forest, Himalayas, 
and N. Chin Hills, Burma. 

In Indian Museum coll. from Mussoorie, India. A small 
black species with grey segmentations and grey median spots on 
the abdomen. Frontal callus large, protuberant, prolonged in a 
short, very thick line ending in a point. Forehead wide. Legs 
black, tibiae yellowish. Length of type 13 mm., others 11-123 mm. 

Face covered with grey tomentum and with whitish or yellowish 
hairs. Palpi long, not very stout at base, ending in an obtuse 
point, reddish yellow densely covered with black hairs. Antennae 
dull reddish, darker at apex, the first two joints with black pubes- 
cence, the second very small, the third wide at base with very 
slight angle. Subcallus and forehead covered with brownish yellow 
tomentum. forehead wide, slightly narrower anteriorly, about four 
times as long as it is broad, the frontal callus blackish, not quite 
reaching the eyes, the forehead with dark markings beyond the 
apex of callus and on vertex. Thorax with two distinct grey 
stripes, grey at the sides, breast with white pubescence. Scutellum 
in well-preserved specimen with a grey tomentose posterior border 
covered with white hairs. Abdomen short, the broad but short- 
ened triangular grey-haired spots are present from 2—5 segments, 
the segmentations reddish yellow but covered with grey hairs and 
grey tomentum; under side with reddish yellow segmentations, 
pubescence on dorsum of abdomen black Legs black, the tibiae 
on basal two-thirds reddish yellow with whitish hairs on the lighter 
part, the fore coxae and hind femora with white pubescence, else- 
where it is black. Wangs hyaline, stigma yellow, veins reddish 
yellow, appendix present. 

In the two specimens from Burma the hairs on the segmenta- 
tions are yellowish. 


Tabanus orientis, ¢ , Walker. 
(Pl. xiv, fig. 18.) 
fast Dipts, 1,.p.'152- (2848). 


Tabanus consocius, 2 , Walker, Dipt. Saund., i, p. 50 (1850). 
Tabanus perlinea, 2 , Walker, l.c. 


= 


196 Records of the Indian Museum. [Vou. IV, 


The type of Tabanus onentis is from Nepal, the types of 
Tabanus consocius and perlinea from India, Saunders coll. 

In Brit. Mus. coll. specimens from N. W. Province; Naini Tal 
and Thandiani, Himalayas; N. India ; W. Bhutan. 

In Indian Mus. coll. from Mussoorie, N. India, about 7,000 
feet ; in Brunetti coll. from Naini Tal, Kumaon, 6—7,000 feet 
(Lloyd); Minda, Nepal, and from Dongagali, Murree district, W. 
Himalayas. It appears to be a Northern Indian species in distri- 
bution. 

A medium-sized species with almost clear wings and a reddish 
yellow abdomen which, when not denuded, is clothed with yellowish 
or whitish triangular median spots and segmentations. Forehead 
moderately broad with a large callus, its linear extension almost the 
same width as itself. Legs black, the tibiae yellowish. Eyes with 
three bands. Length 15—17 mm. 

Face covered with yellowish or greyish tomentum. Beard and 
pubescence of face yellowish white. Palpi long, slender, only slightly 
stouter at base, ending in a rather obtuse apex, with scanty black 
pubescence and few white hairs below. Antennae red, often darker 
at apex, the first two joints and the apex of tooth with black hairs. 
Forehead slightly narrower anteriorly, about five times as long as 
it is wide, covered with yellowish or white tomentum and with 
some black pubescence, the frontal callus blackish or reddish 
brown, oblong, not reaching the eyes, its linear extension quite two- 
thirds its own width, though sometimes narrower at its lower end 
where it joins the callus. Eyes with three bands.. Thorax black, 
when undenuded covered with yellowish tomentum and_ with 
appressed pale yellow hairs, the pubescence black, no stripes 
visible ; the hairs at sides black: breast covered with yellowish 
white tomentum and with white pubescence; scutellum same as 
thorax. Abdomen reddish yellow on the three first segments with 
the exception of a large black median spot on the second and third, 
very distinct when insect is denuded, the fourth segment is very 
largely the same colour but often blackish on its posterior border, 
the remaining segments appear blackish but in the fresh insect the 
posterior segments are widely yellow, on all the segments appear 
yellow-haired triangular spots, and the segmentations are all yellow 
tomentose with yellow pubescence, elsewhere on the dorsum the 
pubescence is black, under side reddish yellow, only the extreme 
apex black, with yellow short pubescence. In some specimens the 
abdomen is darker with blackish markings on each segment, but 
the large callus of forehead distinguishes the species without diffi- 
culty. Legs black, the tibiae a rather transparent yellow, the fore 
pair black at the apex, the femora with yellowish white tomentum 
and pubescence, the tibiae and tarsi with chiefly black pubescence. 
Wings usually clear, sometimes slightly tinged, usually without an 
appendix. The eyes in life are described as greenish with three 
Violet-green bands. 

Tabanus orientalis, Wied., Auss. Zweifl. Ins., i, p. 145, from E. 
India, must be very nearly allied to this species, but Wiedemann 


tgtr.j G. Ricarpdo: Revision of the species of Tabanus. 197 


states the legs are ochre-brown, and calls the thorax yellowish with 
broad reddish brown stripes. It may possibly be identical with 
Tabanus consanguineus, Macquart, which has however yellowish 
legs. 


Tabanus fulvimedius, ¢ , Walker. 


List Dipt., i, p. 152 (1848). 

Tabanus melanopygatus, 2 , Bigot, Mem. Soc. Zool. France, 
p. 651 (1892). [Atylotus.] 

Tabanus pagodinus, 2 , Bigot, /.c., p. 654 (1892). [Atylotus.] 

The type of Tabanus fulvimedius is a 9 from Nepal (Hard- 
wicke Bequest), in bad condition, in the description the locality 
is given as doubtful: there is another @ in the Brit. Mus. coll. from 
N. Chin Hill, Burma. 

There is no doubt that both the Bigot species are identical, 
though the frontal callus in pagodinus varies slightly, having a 
narrower prolongation, described by Bigot as ‘‘ fortement retrecie au 
milieu,’’ he also mentions another callosity on the vertex—which, 
however, only appears owing to denudation—and the presence of 
black pubescence. The three bands on the eyes are just dis- 
cernible. 

In Dr. Kertesz’s collection there is a long series of females 
from Formosa, in very good preservation, with the pubescence on 
the abdomen perfect. 

The species is very nearly allied to Tabanus ortentis, Walker, 
the only apparent difference being in the more widely black apex, 
the last four segments being dark in this species. The wings are 
tinged with brown. 

A medium-sized reddish brown species, with a rather broad 
forehead about six times as long as it is broad, only slightly narrower 
at the vertex, with a large oblong callus prolonged as a thick linear 
extension. Antennae red, palpi slender, reddish yellow with black 
pubescence. Abdomen reddish brown on the first three segments, 
black on the remaining ones with narrow yellow segmentations, 
when undenuded with large greyish yellow haired more or less 
triangular-shaped median spots on each segment, forming a con- 
tinuous stripe. Legs black, the tibiae reddish yellow. Wings 
usually with a short appendix and slightly tinged brown. Length 
15-16 mm. 

The series from Formosa have the abdomen blacker, often only 
the first two segments being reddish yellow and not always entirely 
so. ‘The wings are very distinctly tinged brown often without an 
appendix. 


Tabanus consanguineus, ¢ , Macq. 


Dipt.exot.-1 (i), p. 127 (1838). 


Black. Abdomen with three white stripes, the lateral ones 
not reaching the apex. Legs red. Length 5$ lines. 


198 Records of the Indian Museum. [VOLEV: 


Palpi yellowish white, terminating in an acute point. Face and 
forehead of a pale yellowish grey; the latter with an elongated 
brown callus, continued as a rather wide band. Antennae; the 
first two joints ferruginous; the third wanting. Thorax brown, with 
whitish stripes and tomentum; sides ashy grey. Abdomen black; 
a dorsal narrow stripe of a rather yellowish white, very distinct, 
extends to the end of the sixth segment and is prolonged on the 
seventh by a greyish line; a similar stripe on each side only 
extends to the third segment, being faintly prolonged on the 
fourth by an oblong little spot; lateral borders whitish ; under side 
pale fawn with white tomentum. Legs with white tomentum. 
Halteres ferruginous. Alulae and, wings clear; stigma yellow. 

From the coast of Malabar. Collected by Mons. Dussumier. 
Museum. 

This species resembles Tabanus guinecensis, indicus, trilineatus, 
lineola, occtdentalis, trivittatus, dorsiger, quinquevittatus, costalis, 
obsoletus, but appears to us distinct from all. Macq., Dipt. exot., 
1 (Leper 27. 

From the above description I concluded the species was the 
same as Tabanus striatus, Fabr., but on seeing the type in the 
Paris Museum I find it is not so, but is nearly related to Tabanus 
orvientis, and fulvimedius, Walker, from which it seems only 
distinguished by its smaller size, and the legs being wholly 
yellow. 

The type is in a dirty condition, labelled from Malabar. 
Forehead broad, with the same shaped callus as in TYabanus 
ovtentis, reddish brown, extending almost as a straight stripe to 
the vertex. Palpi yellow, small and pointed. Thorax with grey 
tomentum. Abdomen blackish with pale white-haired segmenta- 
tions, red-yellow on the first two segments and on the sides of the 
third; no stripes or spots now visible; under side pale yellowish. 
Wings clear, veins and stigma yellow. Length 12 mm. 

Comparing it with a specimen of Tabanus orientis from the Brit. 
Mus. coll. it appeared similar allowing for denudation, with the 
exception of the lighter coloured legs and smaller size, so that 
for the present it seems best to keep it distinct from the Walker 
species. 


Tabanus orientalis, 7, Wiedemann. 


Anal. Ent., 21, 8 (1824) ; 7d., Auss. Zweifl. Ins., i, p. 145 (1828). 


Yellow; thorax with brown stripes; abdomen posteriorly 
almost with bands, with a stripe of white spots. 6% lines; ~ from 
East India. 

Antennae rusty yellow. Face mouldy grey; palpi colour of 
chamois-leather ; beard grey. Thorax yellowish with broad reddish 
brown stripes, the median one broadest, cut short behind, divi- 
ded longitudinally by a pale yellowish line; scutellum clear brown 
with a paler border; pleurae chamois-leather-colour. (Wiedemann, 
in Auss. Zweifl. Ins., i, p. 145, where he repeats this description, 


1gtI.] G. Ricardo: Revision of the spectes of ‘Tabanus. 199 


substitutes breast sides for pleurae.) Abdomen inclining to 
rusty yellow; on the posterior segments the base is clear brown, 
which appears as an indistinctly defined band; on the posterior 
border of each segment a triangular spot appears, and the border 
itself has whitish hairs. Wings more or less tinged yellowish, with 
almost leather-vellow veins. Halteres yellowish. Legs ochre- 
brown. Inthe Royal Museum. Wied., Anal. Entom., 21, 8. 


Tabanus incultus, 2 , Van der Wulp. 


Sumatra Exped. Dipt.; 17, 4, pl. i, fig. 8 (788). 


Two females in Lebong, Sumatra, in May. 

Brown. Antennae ferruginous. Abdomen with ochraceous 
segmentations ; anterior legs black, tibiae at the base yellow. @. 
Length 14-15 5 mm. 

Face yellowish, with ashy grey tomentum, towards the borders 
of the eyes lighter; forehead more rust-coloured ; frontal band 
very narrow, gradually widening towards the vertex; the black 
stripe lineal; pubescence of the face brownish. Beard tawny 
yellow. Eyes naked, with fine network. 

Antennae red-yellow, the first joint triangular, the second 
above with a little bristle, the third at the base with a sharp tooth, 
the apex of which is brown, the curved end of the joint black- 
brown (see plate). Palpi grey-brown; proboscis black. Thorax 
- and scutellum grey-brown with partly yellow, partly black-brown 
pubescence, which is longer on the sides of the breast ; on the thorax 
some traces of lighter stripes. Abdomen above black-brown, with 
ochre yellow segmentations which are distinctly marked, especially 
in the middle, by a short white pubescence; under side red-brown 
with very broad yellow bands on the segments. Fore legs black, 
the tibiae for wholly two-thirds at the base pale yellow ; middle legs 
brown-yellow, the coxae and femora at their base blackish, the 
tarsi with the exception of the base, black-brown; hind legs 
black-brown with the knees besides the inner border and apex 
of tibiae yellowish; the pubescence of legs pale and appressed, 
except on the outer side of the hind tibiae where it is very dark 
brown and upstanding. Halteres yellow. Wings tinged yellow- 
grey, more brown-yellow along the fore border; the stripe-shaped 
stigma tobacco-brown, the neuration normal. Although the above 
specimens have too much conformity with each other, for me to 
have the least doubt of their belonging to the same species, yet 
I must refer to single differences. The description is taken from 
the largest specimen; in the other the forehead and face are uni- 
formly ochre-yellow, also the pubescence of the face, of the beard, 
and of the segmentations of the abdomen, which are broadened 
out in the middle into triangular spots; the under side is darker; 
the hind tibiae are except at the apex wholly yellow, yet they have 
nevertheless black-brown pubescence on the outsides. V. d. Wulp, 
Sumatra Exped. Dipt., p. 17. 


200 Records of the Indian Museum. [Vor 


Tabanus geniculatus, ° , Van der Wulp. 

Sumatra Exped. Diptera, 16, 2, pl. i, figs. 7, 7a (1881). 

From Lebong (Sumatra) in May. 

Thorax dark ashy grey, substriated; abdomen brownish red, 
the fore borders of segments brown; antennae and palpi brown: 
legs black, knees reddish yellow. Length 21 mm. 

Related to the former species (7abanus tmmanis, Wied.) but 
somewhat larger and distinguished by the black legs, of which 
alone the knees are red-vellow, by the darker borders of the 
abdominal segments and by the darker beard. Frontal band 
very narrow, posteriorly not wider, in the middle with a shining 
black stripe-shaped line. Eyes naked, with very fine network, 
forehead, face and cheeks with reddish grey tomentum ; face with 
red coloured pubescence; beard black-brown. Antennae, proboscis 
and palpi black-brown ; the antennae on the inner side with grey 
tomentum, the first joint triangular, the second very short, above 
with a bristle; the third with an acute tooth and in consequence 
deeply indented. ‘Thorax and scutellum with dark ashy grey 
tomentum ; the thorax with indistinct stripes, between which in 
some lights appear pale brown-yellow stripes; breast sides dark 
grey-brown; the fore breast and fore coxae with darker pubescence. 
Abdomen brown-red, a little metallic, with dark brown fore borders 
to the segments ; under side black-brown with reddish segmentations. 
Legs black with red-yellow knees. Wings of a grey-brown tinge, 
along the border distinctly yellow-brown, stigma as a brown 
shadow at the end of the subcostal vein. V.d.Wulp, Sumatra 
mMxped. Dipt., 16, 2, pl.1, fig.7. 


GROUP X. 


Forehead with one square, narrow or oblong callus, always 
prolonged towards the vertex as a more or less narrow line, five to 
ten times as long as it is broad. Species with the abdomen uni- 
colorous or nearly so, sometimes darker at apex, but with no spots 
or bands, large, medium-sized or small species. 

The following species are not included in the table, being un- 
known to me, but probably belong to this group. 

Tabanus ardens, Wied. ; Macquart described the male, the type 
of which is probably lost. 

Tabanus laotianus, 2, and melanognathus, @, Bigot, the 
latter described as having a yellow abdomen and black legs, the 
former with legs greyish black, the tibiae greyish at base. 

Tabanus ixion, 7 , Osten Sacken, from Philippines. 

Tabanus fumipennis, Van der Wulp, must be nearly related to 
labanus caerulescens, Macq. 


Tabanus birmanicus, Bigot. 


Mem. Soc. Zool. France, v, p. 653 (1892). [Aévylotus.]| 


1g1t.] G. Ricarpo: Revision of the species of Tabanus. 201 


Type the only specimen, a female, from Burma. 

In Brit. Mus. coll. are females from Ningpoh, India 
(H. M. Lefroy). In Kertesz coll. one from ITushai Hills, Burma ; 
and another from Kuala Lumpur, Malay, London School Tropica! 
Medicine coll. 

A medium-sized dark brown species with lighter thorax, wings 
tinged brown, legs black, with white tibiae and a narrow forehead. 
Length of type 18 mm., others 16—19 mm. 

Face covered with bright yellowish brown tomentum and with 
brownish yellow hairs. Beard same colour as face. Awntennae 
bright reddish, darker at apex, first two joints with black hairs, the 
third long and slender and narrow, witha small tooth at base. 
Palpi yellow with black pubescence, stout ending in an obtuse 
point. Subcallus same colour as face. JLovehead a little darker, 
with black pubescence, about seven times as long as it 1s narrow, 
and a third narrower anteriorly, the frontal callus long, narrow, not 
reaching the eyes with a linear extension, red-brown in colour. 
Thorax covered with nearly the same coloured tomentum as the 
face but slightly darker, and with short appressed fulvous hairs and 
pubescence, shoulders and sides with black hairs; breast the same 
colour, with black pubescence. Scutellum same as thorax. <A bdo- 
men brown, yellowish brown on the sides of the first two segments, 
with black pubescence and some yellow hairs, which appear chiefly 
on the segmentations and as a median line which is, however, not 
always to be discerned; under side dark brown with black pubes- 
cence. Legs blackish, the knees yellowish brown, the fore tibiae to 
their extreme apex, and the others wholly yellowish white, the fore 
coxae covered with yellowish brown tomentum, the pubescence on 
them, femora and tarsi black, on the tibiae white, some yellow hairs 
on hind femora. Wings tinged yellowish brown, lighter on pos- 
terior border. 


Tabanus flavothorax, 9, n. sp. 


d 


Type and another female from Perak, Federated Malay States 
(Pratt), London School Tropical Medicine coll. In Brit. Mus. coll 
one female from Singapore (Ridley); one from Johore (Flower); 
one from Sungie Krudda, Sungie Siput, Perak (Pratt): others 
from Batu Tiga, Federated Malay States (Stanton coll.). 

A species nearly allied to Tabanus birmanicus, Bigot, but at 
once distinguished from it by the yellow middle and _ posterior 
femora, rather narrower forehead, and by the darker brown wings, 
with a distinctly marked clear apex ; in the Bigot species it becomes 
merely paler on the posterior part of apex. Length 16 mm. 

The forehead is quite eight times as long as it is narrow, and 
barely half as wide anteriorly, the frontal callus very narrow, 
reaching the eyes. Thorax more thickly covered with short yellow- 
ish pubescence over the yellowish brown tomentum, so that the 
dorsum appears yellower, the scutelluwm similar, no black pubescence 
on the shoulders, hairs at sides yellowish. Abdomen reddish brown 


202 Records of the Indian Museum. [VoL. IV, 


with black pubescence, long and narrow like the Bigot species; 
under side yellowish with a black median stripe, being densely 
covered with yellowish pubescence. Legs as in Bigot species but 
more largely yellow as specified above. 

Wings tinged dark brown, on the fore border not reaching the 
apex and extending almost as a band across the wing to posterior 
border, becoming smoky brown in colour, costal border yellow, 
stigma dark brown, veins brown. 


Tabanus tinctothorax, 2 , n. sp. 


Type ¢ and another female from Malay States, the type from 
Bukit Katu, Selangor, 3,500 ft. (Robinson), and the other from 
Kinta Valley, 5. Perak, Sept.-Oct. (Ridley). 

A blackish species with the thorax and scutellum reddish 
brown, covered with appressed red pubescence, under side of abdo- 
men with white-haired segmentations. Wings pale brown, yellow- 
ish on fore border and at base. Length 20 mm. 

Face black, with brownish tomentum, pubescence in middle of 
face brownish, on cheeks thick, black. Beard black. Palpi dull 
reddish, appearing black by reason of the thick black pubescence. 
Antennae dull black, the first two with thick black pubescence, the 
third long, the last four divisions short, not a third of the length of 
the whole joint, which is broad at base with a short tooth. Swub- 
callus covered with yellowish brown tomentum. Forehead a little 
darker with some black pubescence, about nine times as long as 
it is broad, almost half as narrow again at the vertex; frontal 
callus narrow, long, with a not very long linear extension, reddish 
brown, shining, furrowed in the type only. ‘The thorax and scutei- 
lum give the insect a distinctive appearance, some black hairs inter- 
mixed with the appressed red pubescence, sides of thorax with long 
black hairs; breast blackish brown with black pubescence. Abdo- 
men oval, blackish brown with close black pubescence, in the 
middle of the first segment some hairs similar to those on the 
scutellum ; under side similar, but with narrow white-haired segmen- 
tations on the second, third and fourth segments. 

Legs black with black pubescence, femora and tibiae with some 
yellowish brown tomentum, hind tarsi with red hairs below. 
Wings with costal border, extreme base and stigma yellow, on 
apical half and posterior border as far as anal cell tinged pale 
brown, the basal and anal cell and axillary angle almost clear, veins 
yellow, brown on apical half. Squamae pale brown. Halteres 
vellow. 


Tabanus nigrotectus, 2 , Bigot. 


Nouv. Archives Mus. Hist. Nat. de Paris (3), ii, p. 204 (2890). 
[Bellardia. | 

Length 20 mm. 

Antennae (incomplete) with first two joints reddish; face and 
forehead of a reddish grey ; frontal callus and vertex black, the 


I9g1t.] G. RricArDO: Revision of the species of Tabanus. 203 


former lineal, very narrow, very slightly dilated below; beard short 
and scanty, black, as are the palpi; the whole body dark black, 
abdomen a little shining. Wings black, with a long appendix, the 
first posterior cell closed with a long stalk. Eyes naked. One 
specimen from Laos, Bigot, /.c., p. 204. 

From Siam, not Island Laos as given in Kertesz Catalogue. 
Not known to me. . 


Tabanus caerulescens, ? , Macq. 


Dipt-exot, ,1:(2)p. 132 (1838). 

Black. Antennae testaceous. ‘Thorax with lateral testaceous 
stripes. Abdomen bluish black. Legs black. Wings brown. 
ienothromimes, <9) °~ 

Palpi and face black with scanty grey tomentum. Forehead 
at the base transversely furrowed, reddish grey, then blackish with 
a shining black spot club-shaped. Antennae brownish testaceous. 
Thorax black, with scanty grey tomentum: a lateral dull testa- 
ceous stripe above and in front of the wings. Abdomen black 
with blue reflections. Legs black; anterior femora hairy above; 
anterior and intermediate tibiae blackish brown, with scanty whit- 
ish tomentum. Squamae and halteres brown. Wings brown, 
less dark on the anterior border, the centre of the largest cells 
hyaline. Neuration normal. From Java Museum, Macq., Dipt. 
ExOce 7 in( Eh): 132: 

Type in Paris Museum seen by me. A black species with the 
wings tinged brown. Forehead very narrow anteriorly, a little wider 
at the vertex, the frontal callus narrow, with linear extension. 
Palpt stout, ending in a point with thick black pubescence. An- 
tennae (incomplete) reddish. Thorax black, red on the basal part. 
Abdomen black. Legs blackish, the posterior tibiae reddish. 
Length 18 mm. No specimens of this species occur in the collec- 
tions mentioned in this paper. 


Tabanus tenebrosus, ? , Walker. 


List Dipt., v, Suppl. i, pp. 242, 389 (1854). 

Type @ from Kanara, Malabar, and another female with the 
name on it but no locality. 

Tabanus obscuratus, Walker, from Celebes is nearly related to 
this species, but has a ‘onger narrower frontal callus, and the ab- 
domen with no sign of the greyish blue tomentum 

From Tabanus caerulescens, Macq., it may be distinguished 
by the dull obscure colour of its abdomen, face not black, legs 
with no white pubescence. 

A dull reddish brown species with a narrow abdomen, narrow 
forehead and frontal callus, blackish palpi; reddish yellow antennae 
and blackish legs. Wings brown, paler on posterior border and 
at extreme apex. Length 18 mm. 


204 Records of the Indian Museum. [VOE Ve 


Face and subcallus covered with yellowish brown tomentum, the 
former with rather long blackish brownish hairs in the middle and 
on the cheeks. Beard blackish brown. Palfi appearing black 
by reason of their very close black pubescence on yellowish brown 
tomentum, the insides yellowish, stout, ending ina point. Antennae 
reddish yellow, the first two joints with black pubescence, the 
third with the apex in the other specimen blackish, in the type 
partly broken off, the tooth at base acute, distinct. Forehead 
about eight times as long as it is broad, and a third narrower 
anteriorly, frontal callus dark brown, club-shaped, not reaching 
eyes, with a lineal extension. Thorax (damaged) covered when 
fresh probably with the same plum-coloured tomentum as abdo- 
men, in the type it is discoloured, in the other blackish; shoulders 
reddish, sides with black hairs. Scwtellum same colour as abdomen 
which is an obscure reddish brown, covered with greyish blue or 
plum-coloured tomentum, pubescence apparently black, under 
side the same. Legs blackish, the tibiae in type dull reddish, in 
the other female black, pubescence entirely black. Wangs brown, 
upper half of submarginal cell clearer and posterior border, in 
the other female many of the cells have clear middles or streaks. 


Tabanus perakiensis, 9 , n. sp. 


Type 2 from Kuala Lumpur, Federated Malay States, Stan- 
ton coll., Ig09, and another female from Perak (Pratt), London 
School Tropical Medicine coll. 

A black species nearly allied to Tabanus inobservatus, a new 
species from Sumatra, but distinct from it, having the face paler, 
the beard largely white, the antennae reddish yellow, the palpi 
paler, yellowish inside and on the outside yellowish but covered 
with grey tomentum, and the black pubescence is thick. ‘Tibiae 
dull reddish. Length 15 mm. 

Face covered with grey tomentum and with some short black 
hairs. Beard whitish with some black hairs. Palpi obscurely 
yellow, with some grey tomentum and with black pubescence, 
slender, a little stouter at base, ending in a point, base of first 
joint with long black hairs. Antennae reddish yellow, darker at 
apex, the first two joints paler with black pubescence, the tooth 
of third joint very small. Frontal triangle covered with reddish 
brown tomentum. Forehead narrow, a little greyer, with some 
black hairs, about eight times as long as it is broad and quite a 
third narrower anteriorly, the frontal callus reddish brown, long 
and narrow, not reaching the eyes, with a lineal prolongation. 
Eyes apparently with two or more cross-bands. Thorax brownish 
black, with some greyish tomentum and traces of appressed 
pale hairs, pubescence black, scutellum same as thorax. Abdo- 
men narrow, blackish or brownish black, with close-lying black 
pubescence, sides with black hairs; under side black with grey 
tomentum and white-haired segmentations. Legs blackish, the 
femora with grey tomentum, the fore tibiae on basal half and 


1gir.] G. Ricarpdo: Reviston of the species of Tabanus. 205 


the other tibiae wholly reddish yellow, the pubescence on legs 
black. Wings dark brown, paler on posterior border and at ex- 
treme apex, the centres of cells somewhat paler, veins and stigma 
blackish brown. 


Tabanus inobservatus, 2 , n. sp. 


B] 


Type @ and another from Kajoe-Tanan, Sumatra (Dr. 
Hagen), in Brit. Mus. coll. 

A dull blackish species. Antennae, palpiandlegs black. Wings 
brownish. Forehead very narrow, distinguished from Tabanus 
caerulescens, Macq., by its dull black not blue-black colour and 
its wholly black colouring, from Tabanus obscuvatus, Wlk., from 
Ceram by its dark wings, and the forehead is narrower anteriorly. 
The thorax also is black, not reddish brown as in the Walker 
species which has reddish antennae. Length 16} mm. 

Face blackish with some yellowish tomentum, its pubescence 
and beard brownish black. Palfi blackish, on inner border 
reddish yellow, with black pubescence, fairly stout ending in a 
point. Antennae blackish, the first two joints with black pubes- 
cence, the third slender with some grey tomentum, the tooth 
small. Subcallus covered with yellowish brown tomentum. 
Forehead nearly ten times as long as it is broad, anteriorly about 
half as broad as it is at the vertex, blackish with a little yellow- 
brown tomentum and some black hairs; frontal callus reddish 
brown, shining, long, narrow, almost reaching the eyes with a long 
lineal extension. Thorax, scutellum and abdomen brownish black, 
the former with a few golden brown hairs scattered over dorsum, 
pubescence otherwise black as it is on the abdomen and breast; 
abdomen conical in shape; under side similar to the dorsum. Legs 
black with black pubescence, the tibiae with some greyish tomen 
tum. Wangs deeply tinged with brown, paler at extreme apex 
and on the posterior border, where however all the veins are 
shaded, the discal cell paler in the centre ; rudiment of an appen- 
dix in the type, none on the other female, veins brownish black, 
stigma black, all posterior cells widely open. 


Tabanus fulvissimus, 2 , Rondaui. 


Ann. Mus. Civ. Genova, vii, p. 458 (1875). 

Length 20 mm. _ From Borneo. 

Body fulvous-reddish, unicoloured. Forehead brown fulvous, 
with the linear-shaped callus reddish, shining ; face paler. Antennae 
and palpi red. 

Thorax on dorsum reddish brown, the pleurae and breast 
fulvous haired. Abdomen on dorsum with the posterior borders 
of segments margined with short fulvous hairs; the ventral seg- 
ments at the base obscurely red, with the apices more or less 
paler. Wings pale brown-yellow, base and costal border fulvous. 
Calyptera brown-red. Halteres reddish with the knob paler. Legs 


206 Records of the Indian Museum. [VOL. 2V. 


ferruginous, the coxae reddish yellow, the tarsi pitchy black. 
Rondani, Ann. Mus. Civ. Genova, vii, p. 458. 

Type seen in the Museum at Genoa. Only the one specimen, 
from which I made the following notes :— 

A very large broad-bodied species. Abdomen reddish yellow, 
very broad. Antennae yellow. Legs reddish brown. Wings 
large, hyaline, tinged yellow. Length 22 mm. 

Face covered with yellowish grey tomentum and a few 
scattered grey hairs. Palpi reddish yellow, long, and a little 
stout at base ending in a long point, with black hairs, darker at 
the extreme apex. Antennae the same colour, darker at apex, 
the first two joints with black hairs, the third with a distinct 
tooth. Forehead the same width throughout, about six times 
as long as it is broad, the frontal callus mahogany-coloured, 
oblong with a linear extension, not touching the eyes, the fore- 
head and subcallus are covered with yellow-brown tomentum, 
some black hairs on the former. Thorax and scutellum the same 
colour as the abdomen with some grey tomentum, most noticeable 
on the anterior part of thorax. Abdomen with narrow yellow- 
haired segmentations; on the under side these are wider and 
whiter and the red-yellow colour is darker. Legs uniform reddish 
brown, the pubescence black. 

Four specimens sent me by Dr. Kertesz for identification 
from Borneo and Sumatra, agree with these descriptions with the 
exception of the wings which appear more brown than yellow, 
but the specimens are faded and dirty; and the palpi have yellow 
hairs. 


Tabanus univentris, 9 , Walker. 

List Dipt., i} p.. 15" (@848)s7d.5-Proc, Winns.soc., Wondonu. ar 
p. 9 (1857). 

Tabanus dives, in parte, 8 , Rondani, Ann. Mus. Civ. Genova, 

ii, p. 457 (1875). 

Type @ from Borneo. (Presented by the Admiralty.) 

In the Proceedings Linn. Soc. Walker refers to two varieties 
of the species from Mt. Ophir, Malaysia, which are not to be found 
in the Brit. Mus. coll. 

A large species easily recognised by the uniform yellow abdo- 
men, the colour of Tabanus par, Wik., the small African species. 
Length 20 mm. 

The Rondani type measures 22 mm. 

Face with yellowish grey tomentum and hairs. Palpz large, 
stout, ending in a point, yellow with black pubescence, some yellow 
hairs at base. Forehead and subcallus with yellowish tomentum 
and some black hairs on the former, which is narrow, about eight 
times as long as it is broad, only slightly narrower anterior.y ; 
frontal callus red-brown, oblong, with a linear extension. Antennae 
with the first two joints reddish with black hairs, the third blackish 
with a prominent tooth. Thorax brownish but covered with dense 


1gti.] G. Ricardo: Revision of the species of Tabanus. 207 


yellowish grey tomentum, and with two very indistinct grey stripes. 
Under side of abdomen yellow with blackish median stripe. Legs 
yellowish red, femora reddish brown, covered on the under side 
with dense yellow-grey tomentum, and chiefly whitish pubescence, 
the apices of tibiae, and tarsi reddish brown, some white hairs 
on the tibiae, otherwise the pubescence on tibiae and tarsi is black. 
Wings hyaline, tinged faintly with yellow, stigma yellow, veins 
yellow-brown. In Walker’s type the abdomen has segmentations 
and a faint median stripe, paler. 

Rondani labelled two specimens (females) from Sarawak as 
Tabanus dives, but on examining these specimens it was clearly 
evident they belonged to two different species, the one measuring 
22 mm. with a uniformly coloured yellowish abdomen marked with 
no spots is identical with Tabanus univentris, Wlk. ‘The des- 
cription was evidently drawn up from an examination of both 
specimens. The smaller specimen is probably nearly allied to or 
identical with Tabanus stanton, see page 174. 


Tabanus flavissimus, 2 , n. sp. 


Type @ from Pundaluoya, Ceylon (Green), and another ? 
likewise from Ceylon. 

A large bright yellow-bodied species nearly allied to Tabanus 
fulvissimus, Rondani, but distinguished from it by the narrower 
forehead, not parallel, and by the smaller narrower frontal callus. 
It is a larger species than Tabanus univentris, Walker, to which 
it is related in the shape of the forehead and frontal callus, but 
has wholly yellow legs and antennae, and the thorax is yellowish. 
Length of type 24 mm., the other is 20 mm. 

Face, palpi, antennae, forehead, whole body, and legs uniformly 
fulvous, the colour of Tabanus par, Wik., a South African species. 
Abdomen a brighter yellow. The pubescence of palpi black, they 
are long, rather slender, ending in a point. Beard brownish yellow, 
long. Antennae long and slender, the third joint narrow at base with 
hardly any tooth, its apex with some hairs, the first two joints 
with yellow pubescence. Jfovehead about seven times as long as 
it is broad, and barely a third narrower anteriorly, the frontal 
callus yellow, narrow, not reaching the eyes, with a lineal exten- 
sion. Thorax (denuded) with whitish yellow hairs at sides and some 
white pubescence. Abdomen with yellow pubescence, thicker on 
the apical half; under side similar. Legs with chiefly short black 
pubescence. Wings vellow on costal border, tinged narrowly yellow 
on all the longitudinal veins, appendix present, stigma yellow, 
veins yellow. 


Tabanus pauper, Rondani. 


Ann. Mus. Civ. Genova, vii, p. 456 (1875). 

Length 20 mm. 

@. Dull brown; the forehead subluteous, the linear callus 
pitchy red. Antennae brown-red, the apex, and the first two joints 


208 Records of the Indian Museum. [VOL LV 


partly, blackish. Head above and below white with white pubes- 
cence; palpi pale with very short black hairs. Thorax and scutel- 
lum subcoriaceous, with brown stripes; pleurae and breast greyish. 
Abdomen dull brown, with a dorsal intermediate series of whitish 
spots, but obsolete; the segmentations with short pale yellow hairs, 
on the under side with more distinct white hairs. Wings palely 
brownish, the veins bordered brown yellowish, the costal border 
browner, the stigma ferruginous. Legs, the anterior femora black, 
the anterior tibiae at the base widely reddish; posterior femora 
pitchy black, the tibiae red, the anterior tarsi wholly black, the 
posterior ones reddish at the base. MHalteres reddish, with the 
club pale subluteous. Rondani, Ann. Mus. Civ. Genova, vii, p. 456. 

Type @ seen in the Museum at Genoa. This large reddish 
brown species, length 23 mm., from Sarawak, is nearly allied to 
Tabanus stantoni, n. sp., but is distinguished from it by the absence 
of any median spots on the abdomen (the type is not in very 
good condition, but does not appear much denuded) which appears 
reddish brown, from the third segment onwards it is covered with 
yellowish brown tomentum, and with some short yellow pubes- 
cence, under side the same but with black pubescence. Thorax 
reddish brown covered with the same tomentum, and some greyish 
tomentum on the shoulders, the pubescence appears to have been 
chiefly black, the scutellum reddish with tomentum yellowish 
brown. Wangs hyaline, tinged pale yellowish brown along all the 
longitudinal veins, no appendix, veins yellow. Face with grey 
tomentum and white hairs. Palpi pale yellow, long, slightly 
stout at base, ending in an obtuse point. Jovehead narrow and 
about seven times as long as it is wide, and two-thirds narrower 
anteriorly, the frontal callus pear-shaped with a rather short 
linear extension. Antennae reddish, duskier at the tips, with a 
prominent tooth, pubescence black on the first two joints. 

No specimens of this species are in any of the collections 
mentioned in this paper. 


Tabanus leucohirtus, 2 , Ricardo. 

Ann. Mag. Nat. Hist. (8), iii, p. 488 (19009). 

Type @ and two other females from Kanara, Bombay, in 
Howlett coll. 

A medium-sized dull reddish brown species, not unlike 
Tabanus fumifer, Walker, in appearance, but distinguished from 
it by the absence of spots on the abdomen and by the dark beard 
and hairs on face. Wings tinged with brown. Legs blackish 
brown, tibiae obscurely red. Length 20—22 mm. 

Face covered with grey tomentum, pubescence brown, but 
some white hairs are visible on sides of face, beard is similar but 
very scanty. Palpi dull reddish, thickly covered with black 
hairs, large, stout ending in an obtuse point. Subcallus, fore- 
head and sides of cheeks covered with yellowish brown tomentum. 
Antennae bright reddish, the apex black, and the first two joints with 


Igtr] G. Ricarpo: Revision of the species of Tabanus, 209 


black pubescence, the third joint long, slender, the tooth near the 
base distinct, crowned with a few black hairs. Forehead narrow, 
about seven times as long as it is narrow, very slightly narrower 
anteriorly, the frontal callus dark reddish brown, narrow, oblong, 
not reaching the eyes, prolonged as a narrow raised line towards the 
vertex, hairs on forehead black. Thorax brown with yellowish brown 
tomentum and scattered black pubescence, a few appressed pale 
yellow hairs are visible, chiefly on the anterior half of dorsum 
and at sides. Scutellum similar. Abdomen dull reddish, appearing 
darker owing to the close short black pubescence, segmentations 
obscurely yellowish or lighter, a few white hairs at the sides of abdo- 
men; under side brown, but with grey tomentum on the sides 
and the segmentations, which latter are wider and more distinct, 
the pubescence black. Legs with black pubescence, but the fore 
coxae with long white pubescence, the fore tibiae reddish at base, 
the others reddish brown. Wings with brown veins and stigma, 
tinged with brown on the fore border becoming paler on the 
posterior border. 


Tabanus atrohirtus, ? , n. sp. 


In Brit. Mus. coll. 

Type ¢ and three other females from Ceylon (Yerbury coll.). 

This species is very nearly related to Tabanus leucohirtus, n. sp., 
but appears a distinct species differing in the following particulars. 

The face is covered with yellow-brown tomentum similar to the 
forehead and subcallus. The beard is wholly brown and thicker, 
and no white hairs visible on the face. The forehead narrower, 
being about ten times as long as it is narrow, and narrower 
anteriorly, the frontal callus longer and narrower. 

The thorax is more thickly covered with the short appressed 
yellow hairs, as is the scutellum. The abdomen appears darker, in 
the type almost black, owing to the thicker short black pubes- 
cence lying all over the dorsum, in the other specimens this pubes- 
cence is not quite so thick and shows the red-brown ground 
colour of the abdomen, the same colour as 7abanus leucohirtus ; 
under side has no light tomentum or pubescence but is similar to 
the dorsum. ‘The fore coxae have black pubescence. [Length 
18—20 mm. 


Tabanus pallidepectoratus, 2 , Bigot. 


Mem. Soc. Zool. France, v, p. 658 (1892). [Atvlotus.| 

Type female from Saigon, Cochin China. 

No other specimens of this species identified in the collections 
mentioned in this paper. 

Type in fairly good preservation is a robust reddish yellow fly, 
with a very narrow forehead and frontal callus, reddish antennae, 
yellow palpi, black legs, only the tibiae whitish at the base, and 
clear wings tinged yellowish brown. Length Ig mm. 

It is nearly allied to Tabanus jotdus, Bigot, from Assam, but 


210 Records of the Indian Museum. [Worstvs 


is distinguished from it by the narrower, longer frontal callus, and 
narrower forehead, by the broader base of third antennal joint, 
and wings less deeply tinged. The tibiae are less widely whitish at 
base. The abdomen is broader and shorter with no golden yellow 
pubescence on the segmentations, only a little appears at the sides 
of the segments. The thorax is a little lighter in colour. 

Face covered with greyish tomentum which becomes yellowish 
brown on the upper parts of cheeks, on subcallus and forehead, 
pubescence on face of white hairs, with darker ones on the cheeks. 
Beard white. Palpi yellow with thick black pubescence, large, 
stout, ending in an obtuse point. Awtennae long, slender, the first 
two joints pale yellow with thick black pubescence, the first joint 
with the upper angle produced, the third joint reddish with a 
small tooth, dusky at its extreme apex. Forehead about ten times 
as long as it is broad, and a third narrower anteriorly, the frontal 
callus reddish brown, long, narrow, not reaching the eyes, with a 
rather thick lineal extension. Thorax brown covered with yellow- 
ish brown tomentum, denuded, pubescence remaining is chiefly 
yellowish, on shoulders and at sides chiefly black ; breast covered 
with greyish tomentum. Scuteliwm similar to thorax. Abdomen 
fulvous, a little browner on the anterior borders of third, fourth 
and fifth, and almost wholly blackish brown on the last two seg- 
ments, pubescence short, golden yellow, scanty, thickest on the 
segmentations, black on “the last two segments ; under side darker, 
reddish brown with paler under segmentations, appearing banded on 
anterior segments, at apex ane dark, pubescence golden yellow 
only on segmentations, black elsewhere. Legs black, knees and basal 
third of all the tibiae whitish with some yellow hairs, fore coxae with 
pale hairs, elsewhere the pubescence is black. Wengs tinged yellow- 
brown on fore border, veins and stigma yellow. 


Tabanus nigropictus, @ , Macquart. 


Dipt. exot., Suppl. v, p. 44 (18535). 

Testaceous (denuded). Thorax with black stripes, breast 
black. Wings hyaline, the veins margined with brown. Length 8 
lines, @. Palpi yellow, long. Beard yellowish. Face and fore- 
head testaceous, frontal callus the same colour, prolonged in a line. 
Antennae; the first two joints testaceous, the third missing. Eyes 
naked. Thorax (denuded) testaceous with three wide black 
stripes ; sides with traces of yellowish tomentum; breast black. 
Scutellum testaceous. Abdomen (denuded) testaceous ; under side 
testaceous with the segmentations less dark, those of the fifth and 
sixth segments yellow. Legs testaceous, the last two joints of the 
tarsi brown. Wings brownish on the outer border, veins normal, 
bordered brownish, stigma reddish brown. From India. M. Bigot. 
Macquart, Dipt. exot. , suppl. v, p. 44. 

This type is much denuded as stated by Macquart, no other 
specimens yet met with by me. The species is distinguished by 
the uniform colour of the abdomen, which is however darker at 


rgti.] G. RicARDO: Revision of the species of Tabanus. 211 


the apex, and /egs, and by the long narrow palpi only slightly 
stouter at the base with a long obtuse point. Forehead almost 
parallel but anteriorly a little broader, about six times as long as it 
is broad, the frontal callus large, oblong, not reaching the eyes, 
yellow, shining, with a long lineal extension. Breast only black 
below legs, is probably reddish in fresh specimens and the same 
may be said of the thorax. Abdomen with short black pubescence 
on the dorsum. Legs with some white pubescence on the fore 
coxae and on the femora, otherwise black. Wzngs with all the veins 
tinged with a brown border, veins and stigma yellow. Tength 
18 mm. 


Tabanus joidus, 2 , Bigot. 


Mem. Soc. Zool. France, v, p. 657 (1892). [Atvylotus.| 

Type @ from India and another female from Sibsagar, Assam, 
this is the second specimen included in the original description. 

In Indian Museum coll. females with no locality specified, 
with the exception of one labelled Sibsagar ; two have MS. names in 
Bigot’s handwriting. 

A narrow-bodied species with a reddish brown abdomen and a 
darker apex, sometimes the third and fourth segments are darker. 
Wings deeply tinged yellowish brown, paler on posterior border. 
Legs black, tibiae white. Forehead narrow with an oblong narrow 
callus. Antennae slender, reddish. Length of type 174 mm., others 
from 16—20 mim. 

Face reddish brown with brownish yellow tomentum, pubes- 
cence appears brownish. Beard brown or yellowish brown.  Palpi 
yellow with thick black pubescence, stout ending in a_ point. 
Antennae pale red, the first two joints yellowish red with black 
pubescence, the third joint longer, slender, not very wide at base, 
with a distinct but small tooth. Forehead about six times as long 
as it is broad, and a third narrower anteriorly , covered with brown- 
ish yellow tomentum and some black pubescence, the frontal callus 
yellowish brown, narrow, oblong, furrowed in the middle, not 
reaching the eyes, with a lineal extension. Thorax brown with 
some short yellow or reddish yellow pubescence and greyish yellow 
tomentum, shoulders with long brownish hairs ; breast covered with 
greyish yellow tomentum and with yellow hairs ; scutellum same as 
thorax. Abdomen long and narrow, reddish brown, the last three 
segments in type black, segmentations fringed with golden vellow 
hairs, on the black part they only appear in the middle, pubescence 
on apex, and elsewhere black, but in one specimen it is almost 
entirely golden yellow on the ante ior segments, on one specimen 
the blackish colour extends on to the third and fourth segments : 
under side reddish brown with black bands commencing from the 
second segment and lighter segmentations, the extreme apex 
wholly black, pubescence black. Legs blackish, the coxae covered 
with greyish yellow tomentum, the fore pair with long vellow 
hairs at base, and at apex with shorter black ones, femora with 
black pubescence, the fore tibiae pale yellowish on their basal half 


212 Records of the Indian Museum. [VoL. IV, 


with white hairs, the others also pale yellow at base with white 
hairs extending only a third of length, then becoming a rusty dirty 
yellowish with thick short black pubescence ; in the specimens in 
Kertesz coll. the bases of tibiae are same colour as the rest, with no 
white hairs, all the tarsi blackish. Wangs with no appendix, veins 
vellowish brown, first posterior cell not narrowed at opening. 


Tabanus siamensis, ° , n. sp. 


d 


Type 2 and another from Siam, June 1906 (W. Palmer). Re- 
ceived from T. R. Bell, Bombay, per H. E. Andrews, in Brit. 
Mus. coll. 

A species allied to Tabanus joidus, Bigot, but distinguished 
from it by the shining black thorax, and reddish yellow abdomen 
with black apex, the under side identical. In shape of forehead 
and frontal callus, colour of antennae, palpi and legs and of wings 
it is very similar to the Bigot species. From Tabanus laotianus, 
Bigot, from Siam, it will differ in the colour of thorax, abdomen 
and legs. Length 17} mm., the other female 16 mm. 

Face, subcallus and forehead covered with yellowish tomen- 
tum, the former with some yellow hairs, and a few brown ones on 
the cheeks. Beard scanty, brownish. Palpi pale yellow with 
black hairs, some yellow ones towards the apex, stout, ending in an 
obtuse point. Antennae reddish yellow, the first two joints paler 
with black pubescence, the third one broader at base than in 
Tabanus joidus, with a distinct but small tooth. Forehead about 
seven times longer than it is broad and a third narrower anteriorly ; 
frontal callus oblong, not reaching the eyes with a long linear ex- 
tension, reddish brown, a large spot of the same colour on the ver- 
tex, probably owing to denudation. Thorax shining black, with 
traces of yellowish brown tomentum anteriorly and at the sides, 
and on the dorsum as stripes, shoulders reddish and breast black, 
both with the same coloured tomentum. Scwutellum similar to 
thorax. Abdomen reddish yellow, somewhat shining, the fifth 
partly. and the last two segments wholly, black, dorsum devoid of 
pubescence ; under side identical but the black segments fifth and 
sixth have distinct yellowish segmentations, hardly visible above, 
some yellow and black pubescence on the under side. Legs black, 
the tibiae reddish yellow on their basal half, pubescence black, the 
coxae covered with the yellow-brown tomentum. Wangs tinged 
brown, yellowish on the fore border, stigma vellowish brown, veins 
brown. 


Tabanus palpalis, 2 , Ricardo. 
(Pie xiv) figs: 21971203) 


Tabanus angustus, 2, Bigot (nomen bis lectum), Mem. Soc. 
Zool. France, v, p. 652 (1892). [Atylotus.] 


Type @ and another female from India, in the description 
one specimen only is mentioned. 


IgiI.] G. Ricardo: Revision of the species of Tabanus. 213 


The name is preoccupied by a Macquart species from 5S. 
America. 

A small narrow-bodied species, of which no specimens are pres- 
ent in the various collections dealt with in this paper, the type 
and other specimen are both ina faded dirty condition, but the 
species seems distinguished from any I have yet seen by its very 
narrow hairy palpi; the frontal callus is club-shaped with a short 
rather stout lineal extension; Bigot gives the colour of the abdo- 
men as light chestnut with a yellowish tomentum, which appears 
fairly correct. Legs uniformly dull yellow. Wings tinged yellow. 

Length of female specimen 13 mm., the type has the abdomen 
broken off and gummed on to the label, it appears about the same 
length. 

Face covered with grey tomentum which becomes on the upper 
part on cheeks near the eyes, and on subcallus yellowish brown, 
cheeks with some brown hairs. Beard brownish with some yellow 
hairs. Palpi narrow, almost the same width throughout, dull 
yellow, the first joint with long brownish hairs, which continue not 
quite so long on the lower border of second joint, which above is 
covered with the usual short black pubescence. Antennae reddish 
yellow, the first two joints with black pubescence, the third narrow 
but with a very prominent short tooth, at apex it is dusky in 
colour with hairs at the joints, apex of tooth also with a few black 
hairs. Forehead about five times as long as it is broad, and 
slightly narrower anteriorly, the frontal cailus reddish brown, pear- 
shaped, with a short linea! extension, not reaching the eyes. 
Thorax brownish. Abdomen apparently chestnut-brown the segmen- 
tations lighter with traces of golden yellow hairs, which are distinct 
on the sides and scattered on the dorsum; under side the same. 
Legs chestnut-brown, the tibiae yellowish, fore coxae covered with 
grey tomentum and with pale hairs, femora with some long soit 
pale hairs, pubescence otherwise black. 


Tabanus fiaviventris, ¢ , Bigot. 
Mem. Soc. Zool. France, v, p. 657 (1892). 


Type ¢ and two others from Sibsagar, Assam. 

In the description India only is mentioned, but the specimens 
are all labelled with the locality ; the fourth specimen included by 
Bigot under this species does not belong to it but is a new species 
now named Tabanus diversifrons. 

In Indian Museum coll. other females evidently of the 
original typical series from Sibsagar. In Brit. Mus. coll one male 
and two females from Ceylon (Yerbury), one of the females is 
marked as ‘‘ not common.’’ None of the specimens are in very 
good condition. 

A small yellow species easily recognised by its yellow abdomen, 
legs and antennae, clear wings and parallel forehead with a small 
indistinct yellow frontal callus. Length type 13 mm: others 
II—1I4 mm. 


214 Records of the Indian Museum. L ViOx 7 2EN, 


9. Face covered with yellowish tomentum and with some yel- 
lowish hairs. Beard pale yellow. Palfi pale yellow with sparse 
black pubescence and some longer yellow hairs below, stout, swollen, 
tapering suddenly to a slender point, so that the upper side is 
curved, the lower side straight. Antennae slender, reddish yellow, the 
first two joints paler, with a few yellow hairs, the tooth of third joint 
distinct. Forehead parallel, about six times as long as it is broad, 
same colour as face, the frontal callus small, not reaching eyes, yellow, 
shining, often very indistinct, with usually no linear extension 
visible. Thorax covered with greyish yellow tomentum and with 
fulvous appressed hairs, sides with yellowish hairs. Scutellum 
similar. Abdomen yellow, slightly darker at extreme apex, pubescence 
fulvous, and fairly thick, in well-preserved specimens ; under side 
similar. Legs wholly yellow, apices of tarsal joints a little 
browner, pubescence chiefly yellow. Wungs hyaline, veins and 
stigma yellow, appendix usually present. 

@. Similar, but the small facets of eye do not reach the vertex. 

There are males and females in the Indian Museum coll. from 
Calcutta and Tenasserim with the apices of fore tibiae, the fore tarsi 
wholly and the others partly brown, which may perhaps justify their 
being divided from this species, but the material is too poor for 
decision on this point at present. 


Tabanus diversifrons, 2 | n. sp. 


Dy 
[Atylotus flaviventris, Bigot, in parte.| 
Type @ and seven others from Shillong, Assam (H. M. Lefroy, 

Entomologist to Govt. of India), in Brit. Mus. coll. and Howlett 

coll., three females from North Khasi Hills, Lower Ranges (Chennell 

coll.), one female from N. Khasi Hills, in Bigot coil., which was 
placed under Tabanus flaviventyis, Bigot, and others from Buxar 

Duars, Bengal and Sylhet, Assam. 

A species very nearly allied to Tabanus fuscicauda, Bigot, but 
distinguished from it by the broader forehead nearly parallel, by 
the larger frontal callus, by the third joint of antennae being bright 
reddish yellow, black at apex, and rather wider at the base, by the 
more slender palpi and by the rather lighter coloured abdomen. 
Length 11-—1I5 mm. 

Face with hardly any brown hairs above. Pa/lpi vellow with 
black pubescence, only slightly stouter at base. Antennae: the first 
two joints yellow with black hairs. The subcallus in some specimens 
is almost bare owing to denudation. Forehead about six times as 
long as it is narrow, almost parallel, the frontal callus reddish brown, 
darker posteriorly and on linear extension, callus not reaching the 
eyes but oblong or pear-shaped with a rather stout linear extension. 
Thorax blackish with fewer appressed fulvous hairs, and the shoulders 
are not reddish. Abdomen reddish yellow, darker at apex, the 
black pubescence on dorsum not so thick, some grey tomentum on 
dorsum, segmentations narrowly paler with some yellow hairs; 
under side is similar. Legs black, tidiae reddish, fore tibiae black at 
apex. Wungs very slightly tinged brown. 


rorl.] G. Ricardo: Revision of the species of Tabanus. 215 


It is distinguished from Tabanus subcinerascens, n. sp., by the 
reddish yellow abdomen with black apex, and no grey tomentum, 
and by the antennae being black at apex. 


Tabanus subcinerascens, ? , n. sp. 
(Pl xiv. dig; 21) 


In Brit. Mus. coll. 

Type and three other females from N. Chin Hills, Burma 
(Watson), 1894. 

This species is very nearly related to Tabanus cinerascens , Bigot, 
but appears a distinct species, shape of the antennae, forehead and 
frontal callus being different. 

The third joint of the antennae is wider at the base and the 
whole joint shorter, the falpi a little stouter. The forehead is 
broader, almost the same width throughout, barely six times as long 
as it is broad, the frontal callus is larger, practically reaching the 
eyes, and is prolonged as a thick spindle-shaped line, reddish brown 
in colour. 

Thorax and scutellum reddish brown with black pubescence and 
some yellowish white hairs. Abdomen more uniformly reddish 
brown, a pale median stripe appears in some lights formed of greyish 
tomentum with a few scattered yellow hairs which also appear on the 
segmentations, sometimes forming a narrow fringe chiefly on the 
apical segments. Legs reddish yellow, femora frequently darker. 
Length 12-13 mm. 


Tabanus obconicus, Walker. 


Dipt. Saund., i, p. 54 (1850). 

Type o from Central India. 

One female from India (Wroughton) in Brit. Mus. coll. One 
female from Bombay and one from Belgatchia, Bengal, in Howlett 
coll. @. A-specimen with a yellowish brown conical abdomen, 
posterior borders of segments covered with grey tomentum, which 
in the middle reaches the anterior border of segment on the 
third and fourth segments at least, so that the anterior yellowish 
brown part of segment has a zig-zag posterior border, the last two 
segments blackish, the pubescence on dorsum black, longer at the 
apex, and on sides black ; under side yellowish brown covered with 
grey tomentum, and with scanty black pubescence. The /egs are 
reddish brown, the middle and posterior tibiae more yellowish brown, 
the fore coxae with grey tomentum and whitish pubescence. The 
thorax is reddish brown. ‘The wings hyaline tinged brown on ‘the 
veins, the cross-veins included. The eyes have the large facets on the 
upper part, reaching just beyond the beginning of the frontal 
triangle, and small ones occupy the lower portion, attaining the 
vertex as a narrow border. ‘The antennac are yellowish on the 
first two joints with black hairs, the third joint is wholly of an obscure 
black (whether this is owing to the bad state of preservation of the 


216 Records of the Indian Muscum. [VoL. IV, 


specimen, or is peculiar to it, is doubtful), tooth distinct but small, 
Face appears yellowish with chiefly yellow pubescence. ‘The 
palpi the same colour. Length 15 mm. 

@. Similar. Face with grey tomentum and white hairs, the 
subcallus and forehead with similar tomentum. Palpi yellow, 
stout at base, ending in a point. Forehead almost parallel, about 
five times as long as it is broad. Frontal callus large, oblong, not 
reaching the eyes, reddish brown with almost a spindle-shaped 
extension. Antennae wholly yellow. Thorax with some white 
hairs anteriorly and at the sides, otherwise the pubescence is black 
on dorsum and on shoulders ; breast with grey tomentum and white 
hairs. Abdomen in like manner has some white hairs on dorsum and 
at sides, the pubescence otherwise black as in @. Legs blacker, 
the fore tibiae at base obscurely yellow, the middle and posterior 
tibiae yellowish. 


Tabanus borniensis, 2 , Ricardo. 


Tabanus apicalis, 9 , Rondani, Ann. Mus. Civ. Genova, vii» 
Pp. 459 (1875). Name preoccupied by a species from Brazil des- 
cribed by Wiedemann in 1828. 


In Brit. Mus. coll. one female from Sarawak. (Purchased Stevens 
1857.) 

Length 12 mm. ¢. Body fulvous, with the apex of the 
abdomen blackish. Antennae widely red. Palpireddish. Forehead 
between the eyes dull brown, the lineal callus pitchy black, the 
exterior part behind the antennae fulvous. Head before and below 
pale grey yellowish and pale haired. Thorax with scutellum brown 
reddish, pleurae and breast grey yellowish. Abdomen with the 
three basal segments above and below fulvous, the two following 
especially at the base, streaked with black ; the two apical ones 
black-brown and brown haired; the preceding ones posteriorly 
shortly ciliated with pale yellow hairs. Wings subhyaline, the 
costal border brown yellowish, towards the apex brownish. Legs: 
the anterior pair dull piceous, the posterior ones yellowish red, all 
the tarsi blackish brown. Calyptera blackish. MHalteres red, the 
club pale vellowish, its apex paler. Rondani, Ann. Mus. Civ. 
Genova, vii, p. 459. 

Type @ seen by me in the Museum at Genoa. A small species 
from Sarawak, with a pale yellow abdomen, apically darker. An- 
tennae yellow with hardly any tooth. Legs yellowish. Wings 
clear. Length 12 mm. 

Face yellowish grey with scattered white hairs. Palpi yellow 
with black pubescence, slender. Antennae wholly yellow with 
yellow hairs on the first two joints. Forehead narrow, about eight 
times as long as itis broad, and nearly two-thirds narrower anteriorly, 
apparently dark coloured, with the frontal callus narrow not reaching 
the eyes, brownish, the lineal extension short, some yellow hairs 
onthe vertex. Eyes bronze-coloured, spotted black. Thorax yellow- 
ish somewhat shining, with grey tomentum, the scufellum yellowish, 
shining. Abdomen transparent yellow on the first three segments, 


rg1t.] G. Ricarpo: Revision of the species of Tabanus. 257 


the remaining segments blackish and reddish brown, with thick 
vellow ‘haired segmentations on third, fourth and fifth segment , 
less distinct on the second, the fifth is almost covered with yellow 
pubescence, the sixth and seventh with almost wholly black pubes- 
cence ; under side similar, the segmentations lighter coloured with no 
pubescence. Legs yellow, fore pair and all tarsi reddish brown. 
Wings slightly tinged vellow on the fore border and apex; veins 
and stigma yellow. 

The specimen in Brit. Mus. coll. has a few black hairs on the 
first joints of antennae. Beard scanty, pale yellow. Forehead 
one-third narrower anteriorly, brown, covered with grey tomentum, 
the lineal extension of callus reaches more than half way to the 
vertex, which is reddish brown. Sides of thorax with yellowish 
white hairs. Legs vellow, the fore femora reddish brown, the poste- 
rior ones paler. 

Another specimen from Sarawak has a label in Walker’s hand- 
writing ‘‘ hybridus’’ but I believe it to be aspecimen of this species ; 
the fore femora and tibiae are wholly vellow. 


Tabanus fuscicauda, 2 , Bigot. 


Mem. Soc. Zool. France, v, p. 658 (1892). [Atylotus. | 
? Tabanus andamanicus,@ , Bigot, /.c., p. 668. [Atylotus.| 


Type @ from Ceylon ; and 3 other females from Pundaluoya 
Ceylon (Green); and another from Peradeniya, Ceylon (Green) ; 
others from Kajoe-Tanau, Sumatra, June 1907 (Dr. Hagen). 

This species described by Bigot from a single specimen is a 
reddish brown medium-sized species with clear wings, blackish, 
legs and tibiae yellowish, with a very narrow forehead and long 
narrow frontal callus. Abdomen with no spots except an indistinct 
black median mark on the second segment, the segmentations 
narrowly paler. Length of type 15 mm., others from 14-15 mm. 

This species might possibly be confounded with small speci- 
mens of Tabanus fumifer, Wlk., but may be distinguished by the 
absence of any median pale spots on abdomen, by the lighter reddish 
yellow tibiae and by the paler segmentations of abdomen with white 
or yellow hairs. 

Face covered with grey tomentum, on cheeks and near antennae 
with yellowish brown tomentum, pubescence white. Beard white. 
A few brown hairs are present on the upper part of face and at sides 
of antennae. Palpi stout, ending in a fairly long point, yellow with 
black pubescence, a few white hairs visible at the base. An- 
tennae dull red, duskier at the apex, the first two joints and apex 
of tooth which is distinct, with black hairs. Subcallus and forehead 
yellowish brown with grey tomentum. Forehead very narrow, about 
nine times as long as it is broad, a third narrower anteriorly, the 
frontal callus very narrow, long, brown, not reaching the eves, 
with a linear extension, the pubescence on forehead black. Thorax 
blackish brown, no stripes visible, covered with grey tomentum 


218 Records of the Indian Museum. [Vior.sny< 


and with scattered short appressed yellowish pubescence and longer 
black hairs, shoulders reddish, sides with black hairs, a tuft 
of white hairs behind at base of wings. Breast covered with grey 
tomentum and with white hairs. Scwutellum similar to the thorax. 
Abdomen reddish brown, blackish towards the apex, and witha 
faint black median mark on the second segment, the segmentations 
yellowish, with yellow pubescence which is scattered and on some 
of the specimens is hardly visible, on the type it appears on the 
median line, the pubescence of dorsum thick, black ; under side very 
similar but appearing darker in the middle with grey tomentose 
sides. Legs black, the fore tibiae at their base, the others wholly, 
reddish yellow, pubescence on fore coxae and on posterior femora 
white or yellowish white, on the others a few whitish hairs appear, 
otherwise the pubescence on them and elsewhere is black. Wangs 
clear, veins yellowish brown, stigma yellow. In the specimens 
from Sumatra the black mark on the second segment is often not 
visible. 

Atylotus andamanicus 1s a very badly preserved female speci- 
men from the Andaman Islands, it appears to me to be identical 
with the above, the only difference appearing to be the presence 
of a round grey tomentose spot on each side of the second and 
third segments. 


Tabanus uniformis, 9 , n. sp. 


5) 


Type @ and a long series from Darien Tiptis, Negui Sembilan, 
Federated Malay States, collected February and March 1908 by 
Dr. A. T. Stanton, who informs me that the locality is a new road 
running through jungle country. 

A species with a uniformly coloured abdomen, reddish, with 
brighter reddish yellow haired narrow segmentations. Antennae 
and legs reddish yellow, the femora darker. Forehead narrow 
with a long narrow callus. Wings clear, tinged yellow on the fore 
border. Length of type 16 mm., others 14—16 mm. 

Face covered with yellowish grey tomentum, the upper part 
of cheeks bordering eyes, the subcallus and forehead with darker 
reddish brown tomentum, pubescence on face scanty yellow, on 
the cheeks browner and longer, beard dark yellowish with brown 
hairs intermixed. Palpi yellow, thickly covered with black hairs, 
stout, ending in an obtuse point. Antennae reddish, the first two 
joints yellow with black pubescence, the third moderately broad 
at base with a distinct tooth. Forehead about eight times as long 
as it is broad, and a third narrower anteriorly, frontal callus shining 
yellowish brown, very narrow, long not reaching the eyes, furrowed 
in the middle with a fine lineal extension, pubescence of forehead 
black. Thorax reddish brown as abdomen but with some greyish 
brown tomentum anteriorly and at sides, the pubescence on dorsum 
very largely of short fulvous hairs, but black hairs are intermixed 
especially in the middle of dorsum, hairs at sides blackish. Scwtel- 
lim 1s similar. 


1gtt.}| G. RicarDO: Revision of the species of Tabanus. 219 


Abdomen reddish brown with bright reddish yellow short pubes- 
cence, always present on segmentations and in type and well-pre- 
served specimens on the whole dorsum, black hairs intermixed, 
all pubescence is short but thick; under side the same but with 
paler segmentations, the black pubescence predominates. Legs 
reddish callow the coxae and femora reddish brown, on the middle 
and posterior pairs however this colour appears only in streaks, 
the fore coxae with grey tomentum and whitish hairs, the femora 
with short black pubescence, some yellow hairs intermixed at 
knees, and on outer border of the middle femora, in the tibiae it is 
black, largely yellow at the base, on the tarsi it is black. Wings 
clear, distinctly tinged yellow on the costal border, and more 
faintly so on the fore border, stigma yellow, veins yellow, a pale 
white streak is visible below the stigma. 


Tabanus subhirtus, 2? , Ricardo. 


Tabanus cinerascens, 2, Bigot, Mem. Soc. Zool. France, 
p. 669 (1892). [Atylotus.} (Nomen bis lectum.) The name is 
preoccupied by a species from Australia named by King in 1827. 


Type @ from Java. 

In Howlett coll. females from Belgatchia, Bengal, and from 
Bombay. 

A small olive-coloured or reddish brown species, the abdomen 
appearing dusky with yellowish brown tomentum and black and 
white pubescence, the thorax olive-brown with white hairs inter- 
mixed with the black pubescence. Antennae light red, palpi pale 
yellow with black and white pubescence, slender. Legs pale reddish 
yellow. Wings clear. Length 12 mm. 

Face covered with whitish grey tomentum and with white 
pubescence. Beard white. Palpz with a long pointed apex. Fore- 
head covered with yellowish brown tomentum and with black 
pubescence, narrow, anteriorly one-third narrower than at vertex, 
about seven times as ‘long as it is wide, frontal callus yellowish brown, 
long not reaching eyes, extended as a fine line beyond the middle 
of the forehead. Antennae long and slender, the first two joints 
with grey tomentum and some black hairs, the apex of third joint 
dark brown. Thorax: shoulders red with ‘black hairs, white hairs 
round base of wings, breast covered with whitish grey tomentum 
and with white hairs. Scutellwm same as thorax. Abdomen: seg- 
mentations very narrowly pale yellow, the apex appears darker 
dorsum sprinkled with white hairs; under side ashy grey with short 
white pubescence. Legs almost unicolorous, the femora and tarsi a 
little browner, pubescence black, on femora some white tomentum 
and on tibiae some few yellow hairs. Wzungs clear, veins yellowish 
and brown, stigma yellow. 

The specimens from Bengal have the abdomen rather redder 
in colour, and the very short white hairs are yellowish in colour 
and not so numerous. 


220 Records of the Indian Museum. [VoL IV, 


Tabanus formosiensis, 2 , n. sp. 


Type ¢@ and another from Formosa in Dr. Kertesz’s coll. 

A small species nearly allied to Tabanus cinerascens, Bigot, 
distinguished from it by the narrower shining black frontal callus 
and narrower forehead, and by its olive-coloured abdomen with 
short yellow pubescence scattered over the dorsum. Antennae 
and legs yellow. Wings clear. Length 11 mm. 

Face covered with whitish tomentum and white hairs, the 
upper cheeks, subcallus and forehead with yellowish brown tomen- 
tum, the former with black hairs. Beard white. Palpi pale yellow, 
with black pubescence, slender, not very stout at base, with a long 
slender apex. Antennae reddish yellow, the first two joints pale 
yellow with black hairs, the third with a very small but distinct 
tooth crowned with a few black hairs, narrow, not very wide at base. 
Forehead nearly ten times as long as it is broad, being only half as 
wide anteriorly as it is at vertex, frontal callus cylindrical, with 
narrow lineal extension, not reaching the eyes, black; some black 
hairs on forehead. Thorax blackish with some grey tomentum 
anteriorly, the dorsum with scattered short yellow pubescence, 
shoulders reddish with black hairs ; sides and breast ashy grey with 
white pubescence. Scutellum similar to thorax. Abdomen olive- 
brown, covered with yellowish brown tomentum and with black 
pubescence, short appressed yellow hairs are scattered over the 
dorsum ; under side black covered with ashy grey tomentum. Legs 
yellow, the coxae covered with ashy grey tomentum and with white 
pubescence, the femora with the same tomentum chiefly on their 
lower sides and with some white hairs below, fore tibiae on their 
basal half with yellowish pubescence, and the others with some on 
their lower sides, elsewhere the pubescence is black, fore tarsi 
black, the others more or less brownish. JWzngs clear, veins and 
stigma pale yellow. 


Tabanus tuberculatus, 9? | n. sp. 


From Indian Museum coll. 

Type @ and another from Companiganj, Sylhet district, 
Assam (B. Warren); and another female from Belgatchia, Bengal, in 
Howlett coll., and type o~ from Pusa, Bengal. 

A small yellowish species allied to Tabanus diversifrons, n. sp., 
but distinguished from it by the shining bare subcallus, forehead 
narrower anteriorly. Thorax covered with yellowish tomentum. 
Abdomen more red-yellow and narrower. Legs almost wholly 
yellowish. Length, male and female types 13 mm., others 12 mm. 

@. Face covered with greyish tomentum and with some short 
yellow hairs. Beard yellow. Palpi bright yellow, a little stout 
at base ending in a long point, pubescence of yellow and black 
hairs intermixed. Antennae (incomplete): the first two joints 
yellow. Subcallus shining, mahogany-yellow, rather protuberant. 
Forehead covered with greyish vellow tomentum, about seven times 


rgtt.] G. Ricarpo: Revision of the species of Tabanus. 221 


as long as it is broad, slightly narrower anteriorly, the frontal 
callus large, oblong, nearly reaching the eyes, red-brown, with a 
short rather stout lineal extension. Thorax black, but covered with 
greyish yellow tomentum and with yellow appressed hairs, sides 
and shoulders reddish yellow with pale yellow hairs; breast black, 
covered with greyish tomentum and with pale yellow pubescence. 
Scutellum same as thorax. Abdomen narrow, reddish yellow, the 
last three segments blackish with some grey tomentum, the pubes- 
cence on dorsum short, black, with bright yellow hairs intermixed 
which become thick at sides, on apical segments the black hairs 
are longer; under side the same, but the yellow pubescence 
predominates. Legs bright reddish yellow, the tibiae and tarsi pale 
yellow, the fore tibiae at apex and the fore tarsi black, the other 
tarsi brown on their apical joints, the fore femora with black short 
pubescence which is also present on apices of fore tibiae and on fore 
tarsi and apical joints of others, the pubescence elsewhere yellow. 
Wings hyaline with pale yellow fore border, stigma and veins. 

@. The males are similar. Eyes with the large facets distinct, 
occupying two-thirds of the eye, the small facets extending to the 
vertex as a narrow border. Face, subcallus and frontal triangle 
covered with greyish white tomentum, the pubescence of former 
and the beard white. Palfi pale yellow with yellow hairs and a 
few black ones at apex. 

Antennae light reddish yellow, the first two joints pale 
yellow, the third joint slender with a very slight tooth. Thorax 
blackish covered with greyish yellow tomentum, the pubescence 
consisting of long pale yellow hairs, sides and breast covered with 
grey tomentum and white pubescence. Scutellum the same with a 
fringe of long yellow hairs on its basal border. Abdomen the same 
as in female, conical, the blackish colouring begins on the fourth 
segment. Legs the same, but the tibiae have chiefly black 
pubescence. Wings clear, the stigma and veins pale yellow, the 
rudiment of an appendix present. 


Tabanus simplissimus, ? 7 , Walker. 


Proc, Linn: S0C.. 1p. Err (1857). 
Tabanus minimus, 2 , Van der Wulp, Sumatra Exped. Dipt., 
18, pl. i, fig. 11 (1892); 7d., Notes Leyden Museum, vii, p. 71 (1885). 


Walker’s types male and female from Sarawak (Saunders 
coll.) ; and others in Brit. Mus. coll. from Perak (Pratt), Kuala 
Lumpur, Selangor (Durham); Railway, Sungei Bessi, Selangor 
(Durham), and Selangor (Meade Waldo); Bidor, 5. Perak 
(Robinson and Annandale coll.) ; one male from Singapore (Ridley). 
V.d. Wulp described his species as follows :— 

‘« A female from Soeroelangoen (Sumatra) in August. Piceous; 
antennae, tibiae and posterior tarsi at the base red; frontal callus 
black, shining; wings subhyaline. @ , length 10 mm. 

Face and cheeks dark grey ; beard white ; subcallus above the 
antennae wholly taken up by a shining black tubercle, which has 


222 Records of the Indian Museum. [Vor. IV, 


a longitudinal furrow in the middle, and is prolonged above as a 
stripe-shaped line on the grey narrow frontal band. Eyes naked 
with very fine network. Antennae red-yellow, the two first joints 
projecting above tooth-like ; the third at the base with a small 
tooth ; at the apex brownish. Palpi grey-brown ending in a point, 
proboscis black-brown. Thorax, scutellum and abdomen brown- 
black, slightly shining. Thorax with short pale pubescence, the 
shoulders and the triangle at base of wings brown ; breast sides 
black-grey with whitish pubescence ; abdomen with some trace 
of reddish segmentations ; under side with distinct grey segmenta- 
tions. Coxae and femora black, the fore coxae with whitish grey 
tomentum and pubescence of same colour; knees and tibiae pale 
red-yellow ; on the fore legs the apex of tibiae as well as the whole 
tarsi black, on the hind legs the first tarsal joint red-yellow with 
the apex and following joints black-brown. MHalteres yellow. 
Wings hardly a little greyish; the stigma marked by an almost 
unnoticeable brown-yellow shadow ; veins normal. This species 
seems related to Tabanus pusillus, Macq. (Dipt. exot., i (1), 127, 12), 
from China, which has also a black subcallus, but the palpi are 
described as pale vellow and the legs as uniformly red-yellow.”’ 
V.d. Wulp, Sumatra Exped. Dipt., p. 18. 

‘“T need only add to my original description that the eyes 
are bare and of a bronze colour, with small facets; even after having 
been moistened they show no trace of cross-bands. In these Javan 
specimens the yellow colour of the legs is so pale that it may be 
called whitish. Two females from Ambarawa in Java (Ludeking).” 
V. d. Wulp, Notes Leyden Museum, vii, p. 71. (1885). 

Van der Wulp evidently re-described the same species as 
Walker had described years before, but the colour of the abdomen 
as stated by him hardly applies to the specimens in Brit. Mus. 
coll. and in London School Tropical Medicine coll., which 
range from yellowish at the base becoming darker towards the 
apex, to reddish brown becoming blackish at the apex, the pubes- 
cence on dorsum black but not very thick, the segmentations in 
fresh specimens fringed with yellow hairs. The tibiae whitish or 
very pale yellow. The wings hyaline but very faintly tinged 
yellowish on fore border, the costal border yellow. 

The forehead is narrow nearly ten times as long as it is broad, 
about a third narrower anteriorly, the frontal callus narrow, long, 
blackish, reaching the eyes with a linear extension, forehead covered 
with grey tomentum. The palpi vary from usually yellow with 
black pubescence to almost black with grey tomentum, swollen 
at base, ending in a rather fine point. Length from 9—12 mm. 
The male type (now headless) has the abdomen almost wholly 
yellow, underneath dark at the apex. 

The male from Singapore has the large facets of the eyes distinct, 
reaching across almost as far as the posterior border of the subcallus, 
the small ones are continued as a narrow border to the vertex. 

This species may usually be distinguished from those specimens 
of Tabanus ceylonicus, Schiner, which have the abdomen almost 


rg1I.| G. RIcaArDO: Revision of the species of Tabanus. 223 


wholly reddish brown (not merely somewhat reddish brown at the 
sides as Schiner describes), by its whitish or yellowish beard and 
lighter palpi which even when darker have grey tomentum, appear- 
ing greyish, but on examining a long series of specimens of both 
species, I found that some with the reddish brown abdomen had 
yellowish beards but the palpi dark, or blackish beards and the palpi 
rather reddish. I can find no other specific difference to divide 
the species, and it appears doubtful whether they can be maintained 
as two distinct species, both in shape of palpi, antennae and frontal 
callosities are almost identical, and both occur in the same localities 
but Walker’s species is not so numerous in the collection. Van der 
Wulp was mistaken in thinking Tabanus pusillus, Macq., had a 
black shining subcallus, his species has no relation to the Macquart 
type which I examined in the Paris Museum. 


Tabanus ceylonicus, 2? , Schiner. 
(Pla xty tise 22.) 


Reise Novara Diptera, p. 93 (1868). 

Tabanus mtidulus, 2, Bigot, Mem. Soc. Zool. France, v 
p. 679 (1892). 

Bigot type from Java and other females in Brit. Mus. coll. 
from Trincomalee, Ceylon (Yerbury), with note from donor ‘‘ Rare” ; 
from Dinding, Siam (Ridley) ; from Singapore, Straits Settlement 
(Falshaw), with note from donor ‘‘ Found in Singapore and Federa- 
ted Malay States, bites cattle and horses, found in town and jungle; 
very rarely seen and not so common as other Tabanidae’’; from 
Tapah Road, Perak, Federated Malay States (Durham); anda long 
series from Bidor, S. Perak (Robinson and Annandale coll.) ; 
from Kuala Lumpur (Stanton coll.) ; from Kajoe-Tanan, Sumatra, 
Dr. Hagen, ‘“ on cow.’’ One male from Trincomalee, Hot Wells, 
Ceylon {Yerbury). 

Shining black, very bare, the thorax at the sides and especially 
the triangle at base of wings as well as the abdomen at the sides 
somewhat brown-red, on the sides of breast just under the root of 
the wings is a rusty brown small spot ; the under side of abdomen 
is likewise but not very distinctly reddish brown, the short hairs on 
the sides of abdomen black. Head black, the face in certain lights 
dark cinnamon-brown, the forehead narrow, somewhat broader 
behind, the shining black callosity just above the antennae large, 
occupying the whole space between the eyes, beyond it a rather 
long square less shining spot, from which three fine shining stripes 
reach to the vertex, the space between it dull bluish grey. 
Antennae bright red-yellow, the third joint black at the apex, the 
first joint cylindrical, not wider above, the second short with no spine 
above, both bare, the third hardly concave above so that the angle 
projects but little, the joints ringed as in other genuine Tabanus 
species. Proboscis black with long sucking discs. Palpi black- 
brown, little shorter than the proboscis, broad at the base, ending 
in a point, somewhat curved; the scanty beard black. Legs black, 


) 3 


224 Records of the Indian Museum. [Vor. IV, 


the knees and tibiae white-yellow, as well as the metatarsi of the 
middle and posterior legs, and to a certain extent the succeeding 
tarsal joints. Wings wholly hyaline, the subcostal vein pale yellow 
and similarly shaded, the first posterior cell open, no appendix, but 
the angle is acute, 6 lines. 

Two females from Ceylon. 

A species deviating very much from all Tabanus species, and 
which might be considered as the type of a definite group of species. 
Schiner, Reise Novara Dipt., p. 93. 

The Bigot specimen agrees with above description and is no 
doubt the same. ‘The species is easily recognised by its shining 
subeallus, black beard and palpi, black legs with white tibiae and 
almost clear wings, but the abdomen varies from wholly black to 
almost wholly brown-red, as in the specimens from Sumatra in 
which only the last four segments or only the apex are blackish, 
it is always shining but in undenuded specimens with unnoticeable 
short black pubescence. ‘The thorax is blackish, sometimes with the 
shoulders and sides reddish, appressed fulvous pubescence appears 
on fresh specimens, but not in the wholly black specimens, sides with 
black hairs. 

Q. The face is black covered with cinnamon-brown tomentum, 
the palpi always blackish, though yellowish on the inside. The 
antennae are rather narrow and slender, the third joint especially 
so, the first two joints have black pubescence which is thickest on their 
upper angles. The forehead is narrow, about eight times as long 
as it is broad, from a third to half narrower anteriorly, the frontal 
callus sometimes long and narrow, not square, but always reaching 
the eyes, linear extension not usually reaching the vertex. Forehead 
covered with the bluish grey tomentum, but posteriorly with 
darker brown tomentum and with black hairs. The eyes have two 
more cross-bands. ‘The fore tibiae are black at their extreme apex. 
Wings with yellow veins and stigma and costal border. Length of 
Bigot type 12 mm., of others 1o—12 mm. 

o. The male is similar to the female. ‘The hairs on the face 
are much thicker and very black, as is the beard and the hairs on 
the palpi. Subcallus black, shining. Eyes with the large facets 
very distinct, reaching across, just beyond the apex of the frontal 
triangle, the small facets are continued as a narrow border to the 
vertex. The abdomen in this one specimen is reddish brown, black 
at the apex. Length 1o} mm. On the relationship of this species 
to Tabanus simplissimus, Wl\k., see under that species. 


Tabanus ardens, ? , Wiedemann. 


Dipt: exots 1,78; 31 ((1821))-ds) Auss.. Zweitl lus de epae5 4. 
68 (1828). 

Macquart, Dipt. exot.,i (1), p. 129, 16 (1838). 

Thorax glauco, fulvo pubescente ; abdomine ferrugineo ; alis 
infumatis ; pedibus fulvis. Thorax mouldy grey with golden yellow 
pubescence. Abdomen rusty yellow. Wings smoke-grey. Legs 
golden yellow, 6 lines. 9%. From Java. 


1giI.] G. RicarDo: Revision of the species of Tabanus. 225 


Antennae very bright rusty vellow, black at their extreme apex. 
Face and beard golden yellow. Palpi light brownish ; forehead 
ochre-brown, a little narrower than usual with a brown shining 
stripe very little broader below. Thorax rusty yellow striped. 
Breast sides golden yellow and mouldy grey. Abdomen with the 
first segment at the sides broadly, and the following ones only on 
the side-borders, waxy yellow with golden yellow pubescence, other- 
wise the abdomen above is almost rusty yellow, clothed with very 
short black small hairs, in the middle of the posterior border of the 
first segment a golden vellow haired spot. Wings on fore border and 
towards the base, also in the middle a little yellowish. Halteres 
golden yellow. ‘Tarsi brown. In Westermann’s coll. Wied., 
Auss. Zweifl. Ins., i, p. 154. 

@. Wiedemann only described the female. We refer to this 
species a male specimen from the collection of M. Serville. It 
differs from the female by the antennae, the third joint of which is 
blackish, vellow at the base. The face and the base of the forehead 
are of a dirty light yellow. The thorax is brown without distinct 
stripe. From Java. Macquart, Dipt. exot., i (1), p. 129. 

This species is not known to me. 


Tabanus fumipennis, Van der Wulp. 


Sumatra Hxped. Diptera, p: 18, 6, pl. 1, fig. 10 (1882). 


One female from Alahan (Sumatra), caught in October. 

Bluish, black, subglabrous. Antennae and legs piceous, the 
knees of anterior legs and base of tibiae red; wings brown. Tength 
16 mm. 

Face and forehead with light ashy grey tomentum ; moustache 
dirty white. Eyes naked. Forehead narrow, posteriorly some- 
what broader, with a stripe-shaped, shining, black line. Antennae 
black-brown, the first joint cylindrical, the second very short, with 
a small spine above, the third at the base with a hardly perceptible 
tooth (see figure). Proboscis shining black. Palpi dark brown 
with a little ashy grey tomentum. Abdomen unicoloured, bluish 
black, a little shining; the thorax and the scutellum covered with 
a greyish bloom ; the sides of breast, at least the part near the fore 
legs light grey, with pubescence of the same colour; under side with 
light grey segmentations. Legs black-brown, but because of the 
grey pubescence partly greyish ; the knees of fore legs and the basal 
half of tibiae red-brown. Wings of a tobacco-brown tint, which is 
darker along the veins and on the fore border ; the veins and the 
stripe-shaped stigma black-brown; neuration normal TJ abanits 
caerulescens, Macq. (Dipt. exot.,i(1), p. 128), might be noticed in 
the determination of this species, were it not for the antennae 
which are there described as brown-yellow, and the shining black 
spot on the forehead of which Macquart speaks is wanting in 
this species in the specimen lying before me. Van der Wulp, 
Sumatra Exped. Diptera, p. 18. 


226 Records of the Indian Museum. (Vor. 


This species is not known to me; it would appear from the 
description that its chief differences from the Macquart species, to 
which it must be very nearly allied, lies in the black-brown not 
reddish antennae; thorax with no reddish stripes or colour; legs 
with the base of the tibiae red, not black as in the Macquart species, 
the posterior tibiae of which only are reddish. The shape of the 
forehead and frontal callus appears identical in both species. 


Tabanus ixion, o , Osten Sacken. 
Berlin. Ent. Zeit., xxvi, p. 99 (1882). 


Dark brown, abdomen and legs black ; wings brown, the tips 
and a part of the discal cell hyaline. Length 15 mm. 

Palpi and face dark brown, with some black pile; head,o. 
flattened from above ; the division between the large and the small 
facets very distinct. Antennae dark brown or reddish brown ; the 
third joint long and narrow; its upper angle projecting squarely, 
the excision of its upper side very shallow. Thorax dark brown, 
with black pile, especially on the pleurae ; abdomen black, shining; 
legs black ; halteres with a yellow knob. Wings brown, axillary 
angle paler, apex hyaline within the second submarginal cell, the proxi- 
mal half of which is brown, a hyaline spot in the middle of the dis- 
cal cell both ends of which are brown ; on both sides of that hyaline 
spot the brown ground colour of the wing is somewhat yellowish ; 
the first posterior cell open although somewhat coarctate towards 
the tips; no stump of a vein. Two@. 

From Philippine Islands. Ost. Sacken, Berlin. Ent. Zeit., xxvi, 
Pp. 99. j 

This male is unknown to me. The females I have described 
as Tabanus inobservatus from Sumatra must be nearly allied to it. 


Tabanus melanognathus, o , Bigot. 


Nouv. Archiv. Mus. Hist. Nat. Paris (3), ii 
[ Atylotus. | 


, Pp. 204 (1890). 


Rene ther 7mm, 

Antennae dull yellow, the third joint black at its apex, deeply 
incised above with a considerably projecting tooth; face, beard 
and palpi yellowish grey, the former very short, the lower half of 
the composite eyes blackish ; the whole body yellow, thorax paler ; 
pleurae brown, halteres yellow; legs entirely black; wings pale 
yellow at the base, the posterior third hyaline (apex widely hyaline 
in the Latin description); no appendix; eyes naked. Laos, one 
specimen. Bigot, /.c., p. 204. This species is not known to me. 


Tabanus laotianus, ? , Bigot. 


Nouv. Archiv. Mus. Hist. Nat. Paris (3), ii, p. 205 (1890) 
| Atylotus.| 


1g11.] G. Ricarpo: Revision of the species of Tabanus. 229 


Length 16 mm. 

Antennae dull yellow, blackish at the base and at the apex, the 
third joint deeply incised above with a fairly prominent tooth ; palpi 
dull yellow (in the Latin description flesh-coloured). Face, beard 
and forehead grey; frontal callus and spot at vertex blackish ; 
the former very narrow, elongated, suddenly dilated below. ‘Thorax 
dull yellow, a little reddish, with four wide stripes indistinct, abbrevi- 
ated posteriorly, blackish (dull chestnut in the Latin description) ; 
sides greyish ; scutellum brownish yellow ; abdomen pale yellow, 
a little brown at the apex; pleurae brownish as well as the stem 
of the halteres, the club of latter whitish ; wings very pale grey, 
narrowly tinged brownish at the base, stigma the same colour, no 
appendix ; legs greyish black, tibiae pale grey at the base, the pos- 
terior ones longitudinally tinged with brown behind ; eyes naked. 
From Laos. One specimen. Bigot, /.c., p. 205. 

This species is not known to me. 


Group XI. 
Subgenus Therioplectes, Zeller. 


Isis, 1842, xi, 819 (1842). 

In this group are comprised all species with distinctly pubes- 
cent eyes, and ocelligerous tubercle on vertex. The heads of the 
males not differing much in size or shape from those of the females. 
The facets of the eyes in the males are uniform in T. hivtus, Wlk., but 
in T. subcallosus, n. sp., they are very distinctly different in size ; 
the other males are not known. Eyes with cross-bands. 

There are only four species from the Oriental Region, forehead 
broad, three to four times as long as it is broad. Species rather 
pubescent. Length 11—17 mm. 


Tabanus (Therioplectes) subcallosus, ~ 2 , n. sp. 


Types @ and @ and two other females from Mussoorie, 
India (Brunetti coll.); others from N.-W. Provinces and Dalhousie 
in Brit. Mus. coll. 

A small black species with white bands on abdomen, easily 
recognised by the shining, yellowish brown subcallus, by the two 
separate transverse calli of forehead, and by the slightly incrassate 
white fore tibiae. Length 11} mm. 

2. Face covered with greyish tomentum and white pubescence. 
Palpi obscurely yellow, thickly covered on the outside with grey 
tomentum, with thick long white pubescence at base and with some 
short black hairs elsewhere, stout, ending in short acute point. 
Antennae reddish, the first two joints yellowish with black pubescence, 
the tooth of third joint very slight. Subcallus convex, brownish 
vellow. Forehead broad, about three and a half times as long as itis 
broad, almost the same width throughout, covered with grey and 


228 Records of the Indian Museum. [VoEs ive 


brown tomentum, frontal callus black, transverse, not reaching eyes, 
the fore border near subcallus irregular, the median cailus black, 
smaller, irregularly heart-shaped, ocelligerous tubercle smail, pubes- 
cence of forehead black. Eyes hairy ; ‘‘ green black, with central 
bright green band edged each side with purple’’ (from note by 
collector), thus withtwe stripes, @ with large facets above. Thorax 
black with grey tomentum and with black pubescence and some 
appressed fulvous hairs ; at sides and base of wings with long white 
pubescence. Scutellum black covered with grey tomentum. 4 bdo- 
men with white-haired bands on each segment, not produced in the 
middle, under side with the white bands broader. Legs black, tibiae 
yellowish white, black at apices, pubescence on coxae, under part of 
femora, light parts of tibiae white, elsewhere black. Wangs hyaline. 

Male identical, with the exception of the second and third 
segments of the abdomen which are yellowish red at the sides. 

The hairiness of eyes is not very apparent in the specimen from 
N.-W. P. and the palpi are pale yellow with black hairs. 


Tabanus (Therioplectes) hirtus, ~ , Walker. 


Ins. Saund., p. 52 (1850). [Tabanus.] 


Type male from India (coll. Saunders), and another male from 
India; females from Naini Tal, Himalayas (Lt.-Col. R. Moore) ; 
near Muktesar, U. P., India (Capt. J. EK. Holmes), and N-W. Prov- 
inces (C. Horne), all in Brit. Mus. coll. 

In Indian Museum coll. females from Phagu, 9,000 ft., Simla 
hills (Annandale). | 

This species bears a superficial resemblance to Tabanus ortentis , 
WIlk., but is at once distinguished by its hairy eyes. A robust 
species with black abdomen largely reddish yellow at the sides. 
Forehead almost parallel with the frontal callus, broad continued 
as a thick spindle-shaped line towards the vertex. Legs blackish. 
Length of male type 15 mm., of females 16-17 mm. 

9. face and subcallus covered with greyish tomentum, the 
latter hairy, with long yellowish hairs, beard the same colour 
as hairs of face. Palpi dull reddish, covered at the base with long 
yellow hairs, elsewhere with chiefly short black pubescence, stout, 
ending in a short point. Antennae blackish, the first two jcints with 
black pubescence, the third joint with hardly a perceptible angle 
in place of a tooth, broad at base. Eyes with three cross-bands, 
densely pubescent. Forehead rather darker than face, clothed with 
blackish pubescence, longer at vertex, almost parallel and barely 
four times as long as it is broad, frontal callus black, shining, 
triangular, broad at base almost reaching eyes, its apex continued 
in a spindle-shaped line, ocelligerous tubercle present on vertex. 

Thorax blackish with three indistinct narrow greyish or yellow- 
ish stripes broadly separated, pubescence of black long hairs with 
fulvous short often appressed hairs intermixed, shoulders reddish 
with long black hairs, tufts of white hairs above wings on sides, 


rgtr.} G. Rrcarpo: Revision of the species of Tabanus. 229 


breast greyish with yellow and black hairs intermixed. Scutellum 
same colour as thorax, posteriorly reddish, with long black pubes- 
cence. Abdomen black, broad and stout for size of insect, the red- 
dish yellow colour largely predominates on the first two segments, 
and sometimes on the third segment , the segmentations yellow haired, 
often produced in the middle as triangular spots, the whole dorsum 
in well-preserved specimens to a great extent covered with short 
vellow pubescence, intermixed with longer black hairs at the apex 
and on sides ; under side blackish covered with grey tomentum and 
lighter segmentations. Legs blackish, the fore tibiae at base and the 
others more largely so, dull reddish, pubescence on coxae yellow, 
femora with some grey tementum and chiefly black pubescence, 
some yellow hairs present, the pubescence on tibiae and tarsi almost 
wholly black. 

Wings clear, tinged brown along veins, and on cross-veins, 
stigma and veins brown, appendix present, first posterior cell widely 
open. 

x. Similar, but the yellow on sides of abdomen is less, and the 
pubescence on dorsum chiefly black, the segmentations vellow 
haired. Legs darker. Eyes with three cross-bands, densely pubes- 
cent, facets uniform in size, hairs on cheeks long, blackish, in middle 
of face yellowish. Palfi reddish with long black hairs, some yellow 
hairs below. 


Tabanus (Therioplectes) wyvillei, 2 , n. sp. 


Type female and two others from Narkanda, Himalayas, 9,000 
ft., July 1907 (Lt-Col. F. Wyville Thomson, I.M.S.); and another 
female from Kasauli, India, from the same donor, who says of 
those from Narkanda, ‘‘ Very common in the Himalayas ; annoys 
man and animals by biting.’’ He also notes the eyes are red with 
three transverse green bars. 

A smaller species than Therioplectes hirtus, Walker, distinguished 
from it by the slender palpi, and by the golden yellow pubescence 
on abdomen, which in well-preserved specimens almost wholly covers 
it, whereas in the Walker species the same coloured pubescence 
is chiefly confined to the segmentations ; the sides are very much 
less reddish yellow, only appearing so on the sides of the second 
segment. ‘he tibiae are much lighter reddish vellow. Length of 
type 13 mm., others from 13-14 mm. 

Face covered with greyish yellow tomentum and with long 
chiefly black fine hairs. Beard yellowish. Palpi reddish yellow, 
slender, stouter at the base ending in a long acute point, the 
pubescence long, black, a few yellow hairs at base below. 

Antennae reddish, darker at apex, the first two joints with black 
pubescence, the third with hardly a perceptible angle in the place 
of a tooth, subcallus and forehead covered with grey tomentum, 
forehead almost parallel, very slightly narrower at the vertex, barely 
three times as long as it is broad, the frontal callus blackish, very 
indistinct in all the specimens, in the type it appears as a small 


230 Records of the Indian Museum. [ Von.) LV; 


irregularly shaped spot, perhaps in well-preserved specimens the 
forehead may prove to have no real callus, ocelligerous tubercle 
small. Eyes hairy, not very thickly covered with pale yellowish 
hairs, with three cross-bands. Thorax black, covered with grey 
tomentum and with pale yellow hairs, shoulders with long black 
hairs, breast blackish with grey tomentum and yellowish pubescence. 
Scutellum same as thorax. Abdomen brownish black, appearing red- 
dish yellow on sides of second segment, the whole dorsum almost 
wholly covered with short golden yellow pubescence, and the sides 
with identical pubescence ; under side blackish almost devoid of hairs 
but with yellowish grey tomentum. Legs blackish, the femora 
with chiefly yellow pubescence, the fore tibiae except at apices, 
and the others wholly, reddish yellow, with black pubescence, but 
some yellow hairs intermixed, fore tarsi brownish, the others red- 
dish yellow. Wings hyaline, shaded brown on transverse and 
longitudinal veins, stigma and veins brownish, no appendix present. 


Tabanus (Therioplectes) albilateralis, 9 , Macquart. 


Dipt.exot., 1 (i), p- 120; 17 (1838). | Labanusy| 

Black. Eyes hairy. Thorax with sides and stripes white. 
Abdomen with golden yellow dorsal spots and segmentations, the 
two first segments testaceous at sides. Legsred. Lengths lines, 2. 

Palpi whitish yellow, elongated, pointed, the apex appearing 
to form a separate joint. Face and forehead fawn coloured with 
whitish tomentum, the latter wide, with a broad callus, brownish 
yellow, posteriorly with three points. Antennae are wanting. Eyes 
pubescent. Thorax black, with long brown hairs and _ whitish 
stripes ; above the wings is a longitudinal stripe of long white hairs 
bordered above by yellow hairs ; sides with long white hairs. Ab- 
domen wide ; the first two segments testaceous with a dorsal trian- 
gular spot ; the others black with dorsal triangular spots ; posterior 
borders testaceous covered with golden hairs ; under side of a dirty 
fawn colour. Legs yellowish, anterior pair wanting. Halteres 
yellowish. Squamae brownish. Wings slightly brownish, darker 
at the base and on outer border, the first posterior cell a little 
narrowed. From Java. Macquart, Dipt. exot., i (I), p. 129. 

Type in Paris Museum, distinguished from Therioplectes hirtus, 
Wlk., with which I compared it by its broader forehead with a 
broad yellowish callus, smaller size and yellow legs. A small species 
with a black abdomen, reddish yellow on the first two segments and 
partly so on the third segment, median spots and segmentations 
yellowish haired ; under side pale yellowish white. Thorax with 
yellowish white hairs at sides. Legs yellow. Wzungs tinged brown, 
the first posterior cell much narrowed. 


Ne oe ee 


a ee eee 


IgiI.] G. Ricarpo: Revision of the species of Tabanus. 231 


LIST OF SPECIES OF TABANUS FROM THE ORIENTAT, 
REGION, INCLUDING ALL SYNONYMS. 


[Note. — All synonyms are printed in italics. | 


TABANUS abbreviatus, Bigot. 

abscondens, Walker. 

administrans, Schiner == amaenus, WIk. 

agricola,! Wied., an Arabian species recorded by Schiner 
from Hongkong; identification doubtful. 

albilateralis, Macq. (Therioplectes.) 

albimedius, Walker. 

albocostatus, Bigot. 

albofasciatus, n. sp. 

alboscutatus, Rondani = optatus, WIk. 

albulus, Walker = virgo, WIk. 

amaenus, Walker. 

andamanicus, Bigot ? = fuscicauda, Bigot. 

angustus, Bigot = palpalis, Ricardo (nomen bis lectum). 

annamitus, « , Bigot. 

apicalis, Rondani = borniensis, Ricardo (nomen bis 
lectum). 

ardens, Wied., assamensis, Bigot == sanguineus, WIk. 

atrohirtus, n. sp. 

auricinctus, Macq. = barbarus, Coquebert. A species 
from S. Europe and N. Africa. 

auriflamma, Walker. 

auristriatus, 1. sp. 

aurotestaceus, Walker. 

basalis, Macq. 

bicallosus, Ricardo. 

bicinctus, n. sp. 

birmanicus, Bigot. 

bituberculatus, Bigot = fulvus, Macq. 

borniensis, Ricardo. 

brunneus, Macq. 

brunnipennis, n. sp. 

bubali, Doleschall 

bucolicus, # , Schiner ? = feideri, Wulp. F 

calidus, Walker ? = albimedius, Wlk. Type recorded 
from Asia with a query. 

ceylonicus, Schiner. 

chinensis, Thunberg ? = striatus, Fabr. Delete from list. 


cinerascens, Bigot = subhirtus, Ricardo (nomen bis 
lectum),. 
clausacella, Macq. = amaenus, Wik. 


|! Macquart records his species as from the islands of the Indian Ocean, 
but this large black species with golden haired bands on abdomen has not since 
been recorded from the Oriental Region, which makes it appear doubtful whether 
Macquart’s locality is correct, so that for the present it is not included in the 
species from the Oriental Region,—see Macq. Dipt. exot., i, p. 134. 


232 


Records of the Indian Museum. [ Vor. IV, 


caerulescens, Macq. 

conicus, # , Bigot ? = abbreviatus, Bigot. 

consanguineus, Macq. 

consocius, Walker = orientis, Wlk. 

costalis, Lichtenstein. Delete from list, description un- 
recognisable. 

crassus, ” , Walker. 

discrepans, n. sp. 

dissimilis, n. sp. 

ditaeniatus, Macq. 

diversifrons, n. sp. 

= univentris, WIk., 2” parte. 

= stantoni, n. sp., 7m parte. 

dorstlinee, Wied: = striatus, F. 

equestvis, Wulp = optatus, Wik. 

explicatus, Walker. 

factiosus, Walker 

felderi, Wulp. 

[fervidus, Walker — conformis, Wlk., a South African 
species, therefore does not belong to the Oriental 
Region. | 

flavicinctus, n. sp. 

flavissimus, n. sp. 

flaviventris, Bigot ( == diversifrons, n. sp., 77 parte). 

flavothorax, n. sp. 

flexilis, Walker, from Celebes. 

formosiensis, n. sp. 

fulvimedius, Walker. ‘wpe described from unknown 
locality. 

fulvissimus, Rondani. 

fulvus, Mg. 

fumifer, Walker. 

fumipennis, Wulp. 

fuscicauda, Bigot. 

fuscicornis, n. sp. 

geniculatus, Wulp. 

hilaris, Walker. 

hirtipalpis, n. sp. 

hirtistriatus, n. sp. 

hirtus, Walker. (Therioplectes.) 

hybridus, Wied. 

ignobilis, Rondani. 

immanis, Wied. 

incultus, Wulp. 

indianus, n. sp. 

[indicus, Fabr., a South American species not occurring 
in Oriental Region as stated by Walker. ] 

inobservatus, n. sp. 

imscitus, Walker. Delete from list, type lost. 

internus, Walker = rubicundus, Macq. 


dives, Rondani 


rgit.| G. Ricarpo: Revision of the species of Tabanus. 


No 
Oo 
Oo 


ixion, Osten Sacken. 

javanus, Fabr. 

joidus, Bigot. 

jucundus, Walker. 

justorius, Rondani. 

khasiensis, Ricardo. 

lacrymans, Bigot — albimedius, WIk. 
laotianus, Bigot. 

leucocnematus, Bigot. 

leucohirtus, Ricardo. 

leucopogon, Bigot. 

leucosparsus, Bigot. 

macer, * , Bigot = striatus, Fabr. 
malayensis, n. sp. 

mandarinus, Schiner, 


manilensis, Schiner ? — striatus, Fabr. 
megalops, #7 , Walker = striatus, Fabr. 
melanognathus, o, Bigot. 

melanopygatus, Bigot = fulvimedius, WIk. 
mentitus, o , Walker? = indianus, n. sp. 
minimus, Walp = simplissimus, WIk. 
moniifer, Bigot = internus, WIk. 


monotaeniatus, Bigot. 

negativus, n. sp. 

nemocallosus, Ricardo. 

nephodes, Bigot. 

nexus, Walker. 

nicobarensis, Schiner. 

nigromaculatus, n. sp. 

nigropictus, Macq. 

nigrotectus, Bigot. 

nitidulus, Bigot = ceylonicus, Schiner. 

non-optatus, n. sp. 

obconicus, @ , Walker. 

optatus, Walker. 

orientalis, Wied. 

orientis, Wied. 

oxyceratus, Bigot. 

pagodinus, Bigot = fulvimedius, WIk. 

pallidepectoratus, Bigot. 

palpalis, Ricardo. 

partitus, Walker = striatus, Fabr. 

pauper, Rondani. 

perakiensis, n. sp. 

perlinea, Walker = orientis, WIk. 

pictipennis, Wulp = flexilis, Wlk., from Celebes. 

pratti, n. sp. 

priscus, Walker = albimedius, Wlk, Type described 
from unknown locality. 

puella, Walker = virgo, Wied. 


234 Records of the Indian Museum. PVOrSEV. 


puteus, n. sp. 

pyrausta, Walker = servillei, Macq. 

pyrrhus, Walker = ditaeniatus, Macq. 

rarus, n. sp. 

rubicundus, Macq. 

rubidus, Wied. 

vubiginosus, Walker. Delete from list, type lost. 

rufiventris, Fabr. Delete from list, impossible to identify 
from description. 

vufocallosus, Bigot = striatus, Fabr. 

sanguineus, Walker. 

serville1, Macq. 

sexcinctus, n. sp. 

slamensis, n. sp. 

signifer, Walker. 

significans, n. sp. 

simplissimus, Walker. 

simicus, # , Walker = striatus, Fabr. 

speciosus, n. sp. 

stantoni, n. sp. 

striatus, Fabr. 

subcallosus, n. sp. (Therioplectes.) 

subcinerascens, n. sp. 

subhirtus, Ricardo. 

sumatrensis, 7, Macq. Delete from list, type lost, des- 
cription being of » only, not sufficient for identifica- 


tion. 
tenebrosus, Walker. 
tenens, Walker = striatus, Fabr. 
tinctothorax, n. sp. 
trigeminus, Coquillet = mandarinus, Schiner. 


tristis, Wulp. 

tuberculatus, n. sp. 

wumbrosus, Walker — albimedius, Wlk. 

uniformis, n. sp. 

univentris, Walker. 

vagus, Walker — albimedius, WIk. 

vagus, Walker ,from Hongkong. Delete from list, type lost. 

vanderwulpi, Osten Sacken. 

varicolor, Ricardo. 

variegatus, Rondani = varicolor, Ricardo (nomen bis 
lectum). 

virgo, Wied. 

wyvillei, n. sp. (Therioplectes.) 


GEOGRAPHICAL DISTRIBUTION OF SPECIES. 


India, including Assam and Burma, 59 species. 


Tabanus /Jurtipalpis, bicallosus, flavicinctus, leucocnematus, 
bicinctus, sexcinctus, ditaeniatus, virgo, nemocallosus, optatus, 


rgtr.]| G. RicarDo: Revision of the species of Tabanus. 235 


non-optatus, nephodes, albofasciatus, basalis, aurtflamma, striatus, 
jilaris, rubicundus, abbreviatus, rubidus, albimedius, speciosus, 
monotaentatus, brunnipennis, abscondens, aurtstriatus, hybridus, 
jucundus, indianus, brunneus, mgromaculatus, leucopogon, crassus, 
alboscutatus, explicatus, khasiensis, sanguineus, oxyceratus, orients, 
fulvimedius, consanguineus, orientalis, bitmanicus, nigrotectus, 
tenebrosus, leucohivtus, nigropictus, joidus, palpalis, flaviventris, 
diversifrons, subcinerascens, obconicus, sublurius, tuberculatus. 
Therioplectes subcallosus, lirtus, wyvitlet. 


Ceylon, 12 species. 


Tabanus ditaeniatus, striatus, albimedius, speciosus, jucundus, 
puteus, discrepans, flavissimus, atrohirtus, flaviventris, fuscicauda, 
ceylonicus. 


Nicobar Islands, I species. 


Tabanus nicobarensis, Schiner. 


Siam, Federated Malay States, Annam, and Cochin China, 29 species. 


Tabanus varus, bicinctus, pratti, annamitus, striatus, abbre- 
viatus, rubidus, /irtistyiatus, brunnipennis, hybridus, stanton, 
brunneus, fumifer, malayensis, dissimilis, s¢gnificans, leucospar- 
sus, sanguineus, birmanicus, flavothorax, tinctothorax, perakiensis 
pallidepectoratus, siamensis, umjormis, simplissimus, ceylonicus, 
melanognathus , laotianus. 


Sumatra, 16 species. 

Tabanus optatus, basalis, striatus, rubidus, ¢vzstis, immanis, 
fumifer, sanguineus, zzcultus, geniculatus, inobservatus, fulvissimus, 
fuscicauda, simplissimus, ceylonicus, fumipennis. 

Java, 17 species. 

Tabanus javanus, optatus, striatus, rubicundus, abbreviatus, 
rubidus, albimedius, immanis, bubali, brunneus, servillei, sangui- 
neus, caerulescens, subhirtus, ceylonicus, ardens. 

Therioplectes albilateralis. 

Borneo, I4 species. 


Tabanus optatus, pratti, hybridus, varvicolor, fumifer, dis- 
similis, gnobilis, nexus, justorius, fulvissimus, wniventris, pauper, 
borniensts , simplissimus. 

Philippines, 4 species. 


Tabanus vanderwulpi, striatus, factiosus, txion. 


South China and Formosa, 18 species. 


Tabanus fulvus, negativus, fuscicornis, striatus, abbreviatus, ab- 
scondens, hybridus, awrotestaceus, jucundus, amaenus, mandarinus, 


2306 Records of the Indian Museum. [Vor. IV, 


indianus, signifer, feldert, crassus, sanguineus, fulvimedius, formo- 
SUeNStS. 

The species peculiar to each district, vzz., not as yet recorded 
from elsewhere, are printed in italics. They number from Indian 
district 39, and two common to India and Ceylon (faviventris, Bigot, 
and speciosus, n. sp.) ; from Ceylon 4 species, all new ; from Nico- 
bars I species ; from Malay district 14 species, chiefly new species ; 
from Sumatra 5 species; from Java 7 species; from Borneo 7 
species ; from S. China district 8 species : from Philippines 3 species. 


SPECIES OF TABANUS FROM N. CHINA AND JAPAN. 


The following is a list of species and synonyms occurring in 
China above the line dividing the Oriental Region from the Palae- 
arctic Region, roughly about 35° lat. N., and in Japan. The des- 
criptions of those species which occur in both Regions are included 
in those of the Oriental Region. ‘The description of Tabanus 
cordiger, Mg., and Tabanus fulvus, Mg., are to be found in Brauer’s 
monograph of the species of the Palaearctic Region in Denkschr. 
Akad. Wien, xlii. The descriptions of the other species are given 
here, with the exception of Tabanus hoang, 7 , Macq., which should 
be deleted from list being apparently destroyed. 


TABANUS amaenus, Walker. 
angusticornis, Loew. (? Atylotus.) 


bituberculatus, Bigot = fulvus, Mg. 

buddha, Portsch. = chrysurus, Loew. 

chrysurus, Loew. 

clausacella, Macq. = amaenus, WIk., occurs in Palaearctic 
and Oriental Region. 

confucius, Macquart = yao, Macq. 


cordiger, Mg. 

ditaeniatus, Macq. 

fulvus, Mg., occurs in Palaearctic and Oriental Region. 
hirticeps, Loew. (Therioplectes.) 

hoang, Macq. Delete from list, type lost. 

humilis, Coquillet. 

japonicus, Bigot. (? Atylotus.) 

lama, Portsch., [| Region 
mandarinus, Schiner, occurs in Palaearctic and Oriental 
pulchriventris, Portsch. (Therioplectes.) 


pusillus, Macq. ‘‘ China ’’ given as locality. 

pyrrhoceras, Bigot 2? = chrysurus, Loew. 

pyrrhus, Walker = ditaeniatus, Macq., a South African 
species. 

rufidens, Bigot ? = yao, Macq. 

signatipenms, Portsch. = clausacella, Macq. 

simicus, Bigot [Bellardia] = amaenus, WIk. 


‘tataricus, Portsch. 
! This species is included here, though not strictly recorded from China, but 
being very near its borders, is likely to be found there. 


Ig1I.] G. Ricardo: Revision of the species of Tabanus. 237 


tenebrosus, Coquillet (mec Walker). 
trigeminus, Coquillet = mandarinus, Macq. 
trigonus, Coquillet — yao, Macq. 

vao, Macq. 

yokoamensis, Bigot. 


Tabanus angusticornis, 2 , Loew. [? Atylotus.| 


Wien. Entom. Monatschr., ii, p. 104 (1858). 


From Japan. 

Ashy grey to yellow. Abdomen darker at base, on each side 
with luteous spots. Eyes short haired. Forehead very wide, 
with no smooth callus. Antennae ochraceous, narrow, the third 
joint angulated above. Legs obscurely luteous, base of anterior 
femora, apical half of anterior tibiae, anterior tarsi wholly and four 
last joints of posterior tarsi black. 

Wings brownish cinereous, stigma and veins dark brown, no 
appendix. Length of body 5#lines. Length of wing 5 lines. 

Face yellow pollinose and yellow haired. Palpi yellowish, 
clothed with pale yellowish and black hairs. Antennae: first joint 
produced, black haired, second black above, third ochraceous, 
natrow, above angulated, hardly forming a tooth. Forehead very 
wide, yellow pollinose, on vertex and as far as the anterior angle 
of eyes with very short black hairs, and with no shining callus. 
Eyes short pilose. Thorax yellow pollinose and yellow haired, 
intermixed with black hairs, at the base of the wings thicker. Abdo- 
men blackish, slightly yellow pruinose and with a very large Iuteous 
spot on each side from the exterior angle of the first segment, and as 
far as the posterior margin of the third segment, a blackish spot 
lying between, which is extended contiguous to the anterior margin 
of the third segment ; first segment with the borders of the following 
segments except the last, and lateral borders of abdomen yellow 
haired, rest of abdomen black haired. Under side blackish, with 
yellowish ashy grey tomentum, borders of apical segments luteous, 
pubescence of abdomen for the most part black, at the base, sides 
and posterior border of anterior segments yellow haired. Legs 
obscurely luteous, black haired, anterior femora at base, anterior 
tibiae on apical half, extreme apices of posterior tibiae, whole of 
anterior tarsi, and posterior tarsi from apex of the first joint, black. 
Halteres luteous. Wings brown cinereous, yellow before the first 
longitudinal vein, stigma and veins dark brown, with an appendix 
(!), first posterior cell at apex subdilated. Loew, Wien. Entom. 
Monatschr., ti, p. 104. This species is not known to me. 


Tabanus chrysurus, 2 , Loew. 
Wien. Entom. Monatschr., ii, p. 1035 (1858); Portschinsky, 


Horae Soc. Entom. Ross., xxvi, p. 202 (1891) ; Coquillet, Proc. U. 
States Nat. Mus., xxi, p. 309 (1898). 


238 Records of the Indian Museum. [VoL.1V, 


2 Atylotus pyrrhoceras, 2, Bigot, Ann. Soc. Entom. France 
(6), vii, Bullet. Ixxvii (1887). 

Tabanus buddha, ° , Portschinsky, Horae Soc. Entom. Ross., 
xxi, 181 [Budda] (1887) ; 1d., xxvi, p. 202 (1891). 

Black brown. Eyes naked. Antennae broad, ferruginous 
red ; palpi luteous, with reddish yellow hairs ; face, forehead and 
dorsum of thorax with reddish yellow hairs ; abdomen with reddish 
yellow and reddish yellow haired segmentations, the posterior 
segments wholly reddish yellow haired ; femora black-brown, the 
apices yellowish; tibiae yellow, the anterior pair blackish at the 
apex, anterior tarsi black, the posterior one dark yellow ; wings 
brown. Length of body 92 lines, of wings 8% lines. 

Black-brown, but not wholly dark. Head with reddish yellow 
pubescence. Eyes naked. Antennae ferruginous red, the first 
joint arched (fornicatus), the third broad, the broad joints with 
short black hairs. Palpi yellow, clothed with very short reddish 
yellow hairs. Forehead narrow, the callus oblong below, continued 
upwards as a long stripe. Thorax on dorsum reddish yellow haired, 
more frequently with hairs intermixed towards the posterior 
margin. Scutellum with black hairs. Pleurae the same. Abdo- 
men black-brown, anteriorly with short black hairs, the posterior 
margin of the first segment dull reddish yellow, at the sides reddish 
yellow haired, the posterior margins of the remaining segments 
reddish yellow and with reddish yellow pubescence ; with the fourth 
segment almost wholly and the following ones wholly reddish yellow 
haired ; under side black-brown, with short black pubescence, with 
ali the segmentations except the first one reddish yellow, and with 
pale reddish yellow pubescence. Femora brownish black with black 
hairs, the apices yellowish and with ferruginous hairs, the tibiae 
luteous, with pale yellow hairs, the anterior ones with blackish 
apex and ferruginous pubescence; anterior tarsi brown-black, the 
posterior ones dullluteous. Halteres black-brown. Wings brown, 
the first longitudinal vein yellowish, the third one with no appendix, 
stigma hardly distinct, the discal cell longer than is usual in most 
large European species. From Japan. Loew, Wien. Entomol. 
Monatschr., 1i, p. 103, 5. 

Labanus buddha is recorded from Chinese Mongolia and des- 
cribed in Latin in Horae Soc. Entom. Ross., xxi, and the following 
was published in the twenty-sixth volume in Russian, the transla- 
tion of which I procured through a friend from Mr. Aylmer 
Maude: 

‘“ The well-known dipterologist V. v. Réder, in a_ private 
letter among other things, kindly informed me that in his opinion, 
the above-named. species described by me, is no other than the 
Tabanus chrysurus described by Loew from Japan. ‘There is not 
the least doubt of the great resemblance of the two species ; yet it 
seems to me certain, nevertheless, that they are different species. 
Not having a Tabanus chrysurus at hand, I can only point out the 
differences between the two species ; basing myself on a comparison 
of my specimens of the Tabanus buddha, with Loew’s description 


Igit.| G. RicaRDO: Reviston of the spectes of Tabanus. 239 


of the Labanus chrysurus. ‘The chief difference is that the basal 
joints of the antennae of the Tabanus chrysurus are covered with 
short black hairs, the frontal callus is oblong and stretches upwards 
in the form of an elongated stripe ; the thighs are of a brownish 
black, and the front tibiae blackish at the ends, the front metatarsus 
brownish black, and the discal cell of the wings is longer than that 
of large species of the European Tabanus. 

The basal joints of the antennae of Zabanus buddha are quite 
bare ; the frontal callus of the forehead is quite round, and does 
not extend upwards at all, all the legs are of a reddish yellow, only the 
thighs at the base are vellow-brown, and the discal cell of the wings 
does not ditfer from that of large species of our Tabanus.’’ 

These small differences hardly seem sufficient to keep Jabanus 
buddha as a distinct species, the lighter colour of the femora may 
perhaps allow it to be considered as the Chinese subform of Tabanus 
chrysurus. 

Coquillet records seven specimens from Japan. In Brit. Mus. 
coll. are females from Yokohama District (Prior), and Hakodate, 
Japan (Fletcher); and in Kertesz coll. from Japan. 

The frontal callus is large, reddish brown, shining, club- 
shaped, the linear extension usually short, rather stout, or nearly 
non-existent. The forehead about five times as long as it is broad, 
almost parallel. The legs in all of the specimens have the femora 
black, reddish yellow at their apices, in some the apices of fore 
tibiae are hardly darker, or very slightly so. Length 25—30 mm. 

Atylotus pyrrhoceras was described from one female from 
north of Yeso, Japan; judging from the description it is probably 
a denuded example of Tabanus chrysurus, which is distantly related 
to Tabanus barbarus, Coq., which occurs in $. Europe and N. Africa, 
but easily distinguished from it by the wholly yellow-haired abdo- 
minal apex, by the yellow beard and darker femora. Tabanus 
auricinctus, Macq., Dipt.exot.,i (1), p. 134, pl. xvii, fig. 2, described 
from the islands of the Indian Ocean, is apparently a synonym of 
Tabanus barbarus, asgiven by Pandellé in Revue d’Entom., i, p. 
201, i (1883), the older specimen is recorded from South Europe 
and N. Africa, and has therefore an extended range if Macquart’s 
locality is correct, which appears doubtful from the absence of any 
specimens in the collections from the Oriental Region which I have 
had access to (see ante, p. 491). 


Tabanus cordiger, Meig. 


Syst. Beschreib., ii, p. 47 (1820),—see Kertesz’s Cat. 1906 for 
full references. 

In Brit. Mus. coll. four temales and two headless specimens 
from Japan (Dr. Miyajima, 1909). ‘The subcallus is almost entirely 
black, shining, perhaps from denudation as the insects suffered 
much damage on the journey; otherwise they are identical 
with European specimens. ‘This species has thus a very wide 


240 Records of the Indian Museum. | Vorsine 


distribution over Europe and Asia, having previously been recorded 
from Asia Minor. 


Tabanus hirticeps, 2°, Loew. (Therioplectes.) 
Wien. Entom. Monatschr., 11, p. 105 (1858). 


From Japan. 

Ashy grey. Thorax white striped. Abdomen marked with 
a double series of ashy greyish spots. Eyes hairy. Antennae 
black, the third joint brown, apex black. Forehead wide, the 
lower callus duplicated, large, the middle one smaller, tuberculated, 
the third wart-like at the vertex, all black. Legs black, tibiae 
brown. Wings ashy greyish, stigma and veins black brownish, no 
appendix. Length of body 6 lines. Length of wing 54 lines. 

Face white ashy grey, with white hairs. Palpi ashy greyish, 
clothed with longer white and black hairs. Antennae curved, the 
basal joints black, the first arched a little, the third joint brown, 
at apex black, wide, above angulated, the angle obtuse. Fore- 
head very wide, ashy grey ; lower callus duplicated, the anterior 
part wider-and contiguous to antennae, upper part filling the whole 
width of the forehead, middle callus smaller, remote from margin 
of eyes, less smooth, below tuberculated, vertical callus minute, 
wart-like. Eyes hairy. Thorax and abdomen very much denuded, 
blackish, but with traces of whitish grey pollinose and ashy grey 
colouring, stripes of thorax whiter. Abdomen with double series 
ashy grey spots, the median stripe of abdomen composed of hardly 
apparent triangular spots. Hairs of thorax white, mixed with 
black at base of wings. Posterior margin of abdominal segments 
dull white. Legs black, tibiae brown, toward apices blackish, 
femora mostly white haired, tibiae black haired, the posterior ones 
at base white haired, tarsi black haired. Halteres black. Wings 
somewhat ashy grey, transverse veins very slightly shaded with 
brown, stigma and veins dull brown, no appendix. Loew, Wien. 
Entom. Monatschr., ii, p. 105. 

This species is not known to me. 


Tabanus humilis, ? , Coquillet. 


Proc, U.S, Nat. Mus:; so0l, 311 (1808): 

Female from Japan, differs from Tabanus trigonus, as follows : 

Head above antennae yellow and brightly polished, antennae 
wholly reddish yellow, the tooth of the third joint very small, hairs 
of the palpi mixed white and black, those of the occiput, lower side 
of the head and on the pleurae, white. Scutellum noticeably 
lighter grey than the mesonotum. Abdomen somewhat polished, 
black, marked with grey as in ¢vigonus, the median triangles ex- 
tremely small on the last two segments ; venter black, light grey 
pruinose. Extreme apices of the middle and hind tibiae brownish 
black. Wings hyaline, stigma yellow, base of anterior branch 
of the third longitudinal vein not appendiculate, first posterior cell 


IgtI.]_ G. Ricarpo: Revision of the species of Tabanus. 241 


not narrowed at its apex. Length 12-13 mm., two specimens. 
Coquillet, Proc. U.S. Nat. Mus., xxi, 311. 
This small species with shining subcallus is not known to me. 


Tabanus japonicus, 9? , Bigot. 


Mem. Soc. Zool. France, v, p. 632 (1892). [Dzchoelacera. | 
Ricardo, Ann. Mag. Nat. Hist. (7), xiv, p. 368 (1904). 


Antennae yellowish, the apex black; palpi yellowish; eyes 
tomentose ; forehead grey, callus narrow, elongated, black ; beard 
white; thorax and scutellum black, with scattered yellowish tomen- 
tum, sides with yellowish grey hairs, abdomen reddish yellow, the 
base of the segments tinted with reddish brown, those of the last 
ones blackish; legs black, tibiae yellowish ; wings almost hyaline, 
the outer border tinged pale yellow, stigma pale reddish. 15 mm. 
From Japan, one specimen. Bigot, Mem. Soc. Zool. France, v, 
p. 632. I have not seen any specimens of this species, which ap- 
parently has eyes hairy, and may possibly belong to the subgenus 
Atylotus. 

Tabanus lama, 2? , Portsch. 


Horae Soc. Entom. Ross. 
(eyes). 


Spel p20 by (LOOn). ply 1, fig... ¥ 


From Chinese Mongolia. 

Oblong, ovate, luteous. Eyes naked, ornamented with four 
bands, the middle ones dark purple with red margins. Head with 
white tomentum and hairs. Forehead grey, callus subquadrate, 
black, shining. Antennae luteous, the third joint testaceous, at 
apex darker. Palpi white and with white hairs. Thorax black, 
white pruinose with four dark grey stripes. Scutellum black, white 
or grey pruinose. Abdomen luteous, each segment in the middle 
with a white triangular spot girdled on each side with an oblique 
black line and with two small black side spots, under side and all legs 
wholly luteous, all hairs of abdomen white or whitish, only the last 
segment of abdomen at apex clothed with some black hairs. Wings 
subhyaline, shining white, nervures pale ferruginous with a long 
appendix. 45—6 mm. Portsch., Horae Soc. Entom. Ross., xxvi, 
Dp: 20D. 

This species is not known to me, 


Tabanus pulchriventris, 2? , Portsch. (Therioplectes.) 

Horae Soc. Entom. Ross., xxi, p. 179 (1887). 

From Etschjin-Choro, Chinese Mongolia. 

In size, stature and colour of body this species is similar to 
Tabanus tropicus, but is easily distinguished frem it by the abdomen 
being entirely and densely covered with long fine red golden hairs, 
brown palpi, etc. 

Black, with rather long pubescence. Head black, ashy grey 
pruinose, underneath with white beard, round eyes with spare 
black pubescence. Frontal stripe rather wide, ashy grey, and with 
black hairs on vertex. Frontal callus not large, subrotund, black, 


242 Records of the Indian Museum. [Vor. IV’, 


continued as a narrow black line. Eyes shortly but distinctly pale 
haired, with three bands ; ocelligerous tubercle small.- Palpi brown, 
on apical half narrow with obtuse apex, above and a little below 
channelled, on basal half incrassate, and at base with proboscis 
with rather long black hairs. Antennae black, basal joints with 
rather long black hairs, second joint wholly, and third at base, 
widely red, third joint at base with no tooth. Thorax black with 
rather long black pubescence and numerous yellow hairs intermixed. 
Pleurae long and densely covered with snowy or white hairs. Scutel- 
lum black with rather long yellow pubescence. Abdomen black, 
the three first segments widely yellow at sides and all densely 
covered with red golden hairs, under side black, each segment in the 
middle black haired, but at apex narrowly margined with yellow 
hairs. Wings subinfuscate, darker round the transverse veins, 
stigma brown. Squamae ashy grey, margined with brown. Hal- 
teres brown. Legs black and black haired. Tibiae yellow testaceous 
with yellow hairs, and apex with tarsi brown or darker. Length 
7 mim “Portsch.. Horace sete. ps k70: 
This species is not known to me. 


Tabanus pusillus, @ , Macq. 

Dipt: CxOtl (Ch) p.13 (ko 38) 

Black. Antennae red. Abdomen with yellow hairs. 

Legs red. Length 4% lines, ?. 

Palpi of a pale yellow. Face and forehead brownish grey ; 
the latter with a black roundish callus. Antennae fawn coloured. 
Thorax black, denuded ; a brownish spot on each side. Abdomen 
black, with yellow hairs; segmentations fawn coloured. Legs 
fawn coloured. WHalteres yellow. Wings a little yellowish, pale 
yellow at base and on outer border. 

From China: Museum, Macq., Dipt. exot.,1(1), p. 131. 

Type @ in very bad preservation seen by me in Paris Museum. 

A small species with clear wings, the veins yellow. Legs 
wholly yellow. Abdomen apparently dark brown with yellowish 
pubescence. Eyes apparently bare. Lorehead with only one 
callus, nearly square, not reaching eyes, no linear extension, the 
same width throughout, about four times as long as it is wide. 
Antennae yellow. Face hairy. Beard white. Palpfi slender, a 
little stout at base with a slender point and rather long black hairs. 

I have not found any specimens identical with this species. 
There is another 7. pusillus named by Macquart from New Holland. 
This is left for the present in the Palaearctic Region, but as only 
‘“ China ’’ is given, it may belong to the Oriental Region. 


Tabanus tataricus, 2 , Portsch. 


Horae Soc. Entom. Ross., xxi, p. 178 (1887). 
From Btschan, Central Asia; Mt. Altai,—see Verrall, British 
Flies, v, p. 10, 1909. 


1g1I.} G. Rrcarpo: Revision of the spectes of Tabanus. 243 


Related to 1 abanus astur, Erichs. 

Body ovate, black with rather long and dense pubescence. 
Head comparatively rather small, ashy coloured, pruinose and with 
the wide frontal stripe long and densely pale yellow haired. Fore- 
head above antennae and the narrow band below, each side of eyes, 
black, shining. Frontal callus black, shining, short (but equal in 
width to the front), eyes shortly but distinctly yellow haired with 
three wide bands, ocelligerous tubercle small. Palpiblack and black 
haired, with apex moderately acute. 

Antennae reddish brown, at base black haired, the third joint 
at base with no tooth, brown testaceous, apex and upper side 
more or less widely black. Thorax, pleurae, breast and scutellum 
all wholly black with long and densely yellowish (shining white) 
hairs. Abdomen short, wide, ovate, stout, black, the three first 
segments at sides fulvous, the fourth with golden or reddish hairs, 
with shining luteous rather long and dense pubescence, the other 
segments with luteous pubescence ; under side black, shining, the 
last segments alone with very narrow pale incisions, the first seg- 
ments mostly with black incisions, the three last alone luteous 
haired. Wings subhyaline, stigma brown, all veins black, middle 
transverse veins clouded, the third vein with an appendix, clouded 
at its base, halteres brown, apices white. Squamae ashy coloured, 
margined with white. Femora all black and black haired, anterior 
tibiae yellow, anterior half and tarsi brown, intermediate tibiae 
and tarsi and all posterior ones wholly yellow with white hairs. 
G7 -) sPactseh; Horace setc..p. 178. 

This species apparently differs fron. Tabanus astury which was 
described from Siberia in the absence of the small tooth, the frontal 
callus being black, not grey, the thorax and abdomen more yel- 
low haired, the femora black haired instead of yellow haired, the 
wings subhyaline, not black-brown. 

It must be nearly allied to Tabanus pulchriventris, described 
on the next page by the author, from Chinese Mongolia. 


Tabanus tenebrosus, ° , Coquillet. 


Proc. U. S. Nat. Mus., xxi, p. 310 (1898) ( mec Walker). 

Female. Differs from Tabanus trigonus asfollows. Head above 
the antennae yellowish grey, pruinose, tooth of third joint of 
antennae very small, hairs of occiput, of under side of the head and 
on the pleurae, white. 

Abdomen opaque, black, sometimes a reddish spot on sides of 
the second segment, wholly greyish pruinose, the median triangles, 
posterior and lateral margins of segments 2-6 a lighter grey than the 
remainder of the abdomen, the darker markings consist principally 
of pairs of spots bordering the median triangles ; venter black, 
brownish grey pruinose in the middle, the sides a lighter grey. 
Wings hyaline, stigma pale yellowish. Length 18—20 mm. 

Four specimens from Japan. 


244 Records of the Indian Museum. (HE VOL, 2Ve 


This name is preoccupied by a Walker species from India. 

The species is very probably the same as the specimens I have 
identified from Japan as Tabanus cordiger, Mg., but the description 
is too short to make a decision possible. 


Tabanus yao, 2, Macquart. 


Dipt. exot., v, p. 44 (1855). 

Tabanus confuctus, @ , Macquart, /.c., p. 46. 

Tabanus irugonus, 2 , Coquillet, Proc. U. S. Nat. Mus., xxi, 
p. 309 (1898). (Japanese subform.) 

? Alylotus rufidens, 9° , Bigot, Ann. Soc. Entom. France (6) 
vii, Bullet. Ixxviii (1887). | Atylotus. | 

Both the Macquart types in Mr. Verrall’s coll. are from N. 
China, and with them are a series of females from Japan which 
are almost identical and may perhaps be considered as the Japanese 
subform 7. trigonus. 

In Brit. Mus. coll. are females from Wei Hai Wei, China 
(Muat), “* biting horses;’’ and from Shanghai; and males from 
China (Walker coll.), and males and females from Japan (T. trigonus). 

Black. Thorax with ashy grey pubescence. Abdomen blackish, 
with dorsal white spots and red segmentations. Antennae red with 
the apices black. Legs red. Wings with an appendix. 

Length g lines,2. Palpi yellow. Beard white. Face with 
white tomentum, sides with yellow tomentum. Forehead anteriorly 
with white tomentum, then with yellowish grey tomentum ; an- 
terior callus testaceous, prolonged by a black line. Antennae 
yellow; the last four divisions of the third joint black, this joint 
with the usual tooth. Eyes naked. Thorax with ashy grey tomen- 
tum, and yellow hairs. Abdomen blackish, with dorsal triangular 
spots of a greyish white tomentum, as well as the posterior borders 
of the segments at the sides; sides of the three first sometimes of 
brownish yellow ; under side yellow; the three last segments blackish 
with yellow posterior borders. Legs yellow, anterior coxae with 
yellow hairs. Wings a little yellowish, veins normal with the 
exception of the appendix. From N. China. M. Bigot. This 
Tabanus resembles T. confucius strongly ; and as I have only ob- 
served males in this latter and females in the former, it appears 
that they belong to the same species. However the appendix, 
which is not found in 7. yao, is a character that we have never 
observed to be sexual, and this determines us to consider these 
two species as distinct. Macq., Dipt. exot., v, p. 44. 

On page 46 Macquart gives the description of Tabanus confu- 
ctus, as follows :— 

Black. Thorax with white stripes. Abdomen with the sides 
and segmentations testaceous. Antennae red with the apices black. 
Legs red. 

Length 8 lines, ». Palpi brownish, the last joint oval. Beard 
yellow. Face with white tomentum, sides with yellow tomentum. 


) 


Igit.} G. Ricardo: Revision of the spectes of Tabanus. 245 


Forehead anteriorly with white tomentum, sometimes traversed 
longitudinally by a brownish stripe. Antennae yellow, the last four 
divisions of the third joint black, this joint with the usual tooth. 
Eyes naked. ‘Thorax with indistinct yellowish grey tomentose 
stripes ; sides with grey tomentum and yellow hairs ; a small black 
stripe sometimes indistinct ; breast with whitish tomentum and hairs. 
Abdomen: the testaceous sides reach more or less towards the mid- 
dle, sometimes only leaving a black dorsal mark on the second and 
third segments ; the last four segments with their posterior borders 
of a bright red testaceous ; with a posterior yellow edging ; under 
side testaceous ; the three last segments black, with a testaceous pos- 
terior border. Legs of a dull testaceous ; anterior coxae pale, with 
yellow hairs; posterior tibiae brownish on the posterior side. 
Wings almost clear, a little brownish on the outer border ; veins 
normal, as those of 7. bovinus. 

From N. China. M. Bigot. 

A large species allied to Tabanus bovinus, L., but distinguished 
from it by the larger tooth of the antennae which are bright reddish 
yellow, usually black at the apex, and by the wholly yellow legs. 
2. Abdomen reddish yellow with apex blackish, and a large median 
series of grey tomentose, triangular spots on a black median stripe 
and grey tomentose segmentations, the anterior borders of segments 
are toa great extent reddish brown ; under side almost wholly yellow. 
Legs yellow, but the tarsi reddish brown. Wengs often tinged with 
brown along the veins, or nearly clear. Forehead very slightly 
narrowed anteriorly, about eight times as long as it is narrow. 
Frontal callus pear-shaped, large, hardly touching the eyes with 
a linear extension. Palpi reddish yellow, long, aimost the same 
size throughout. Length 24 mm. 

‘This species may be considered as the Chinese form of Tabanus 
bovinus. 

Tabanus confucius,* , Macquart, type (male), was distinguished 
from this species by the absence of an appendix to wing, which 
of course is not a trustworthy character, some of the specimens in 
Brit. Mus. coll. possess one and some have none ; I believe the male 
type of confuctus is the same species as Tabanus yao but so denuded 
that the black median stripe is not covered with the triangular 
median grey spots, though traces of them can be seen. Tabanus 
trigonus, Coq., is identical, with the exception of the colouring of the 
legs, with the Macquart species, and can only be considered as a sub- 
form so far peculiar to Japan. Abdomen darker. The femora and 
coxae are blackish or reddish brown with grey tomentum and 
bright yellow pubescence on the coxae and under part of femora, 
on the upper sides of the latter it is shorter and black, tibiae yel- 
low, appearing blackish at apices owing to the thick black pubes- 
cence which elsewhere is largely yellow, tarsi blackish with black 
pubescence. 

Atylotus rufidens, Bigot, may possibly be a denuded specimen 
of the subform ¢ryigonus, described from one female, collected north 
of Yeso, Japan. 


246 Records of the Indian Museum. FVOL.AEV; 


Tabanus yokoamensis, ? , Bigot. 
Mem. Soc. Zool. France, v, p. 678 (1892). 


Antennae black, base of the third segment reddish, this last 
joint very concave above with a projecting tooth ; palpi very pale 
yellow ; face and beard white ; forehead grey, callus black, narrow, 
pointed above, much enlarged below and nearly square, on the 
vertex is a very indistinct tubercle and no ocelli ; thorax blackish 
with three wide greyish indistinct stripes ; scutellum blackish ; 
sides grey with grey hairs; the two first segments of abdomen grey, 
with a large blackish spot on each side, the others black narrowly 
bordered reddish, squamae whitish bordered with brown, halteres 
yellow, the club brown with its apex whitish ; legs black, base of 
tibiae widely whitish ; wings almost hyaline, stigma pale yellowish. 

Var.? Abdomen black with a dorsal stripe formed of triangular 
greyish spots, smaller in size. 

I place this species in the genus Tabanus (proprie dictus) but 
with some doubts, owing to the very mediocre development of the 
vertical tubercle. Length 14 or 17mm. Yokohama, Japan, three 
specimens, Bigot, Mem. Soc. Zool. France, v, p. 678 (1892). 

This type and species is unknown to me. 


SPECIES OF TABANUS FROM PERSIA, TURKESTAN, 
AFGHANISTAN, BALUCHISTAN, ASIA MINOR 
AND SYRIA. 


The following is a list of the species with all synonyms from 
the above localities in the Palaearctic Region, including three new 
species here described. Tabanus infuscatus, Loew=apricus, Mg., 
Tabanus fulvicornis = montanus, Mg., var.,and Tabanus bromius, L., 
are given on the authority of Bigot who inserts them in a Cata- 
logue of species of Tabanus from N. Persia and Caucasus, in Ann. 
soc. Ent. France (5),-x, p: 140:(1880)- 

Tabanus abazus, 2, Bigot, Ann. Soc. Ent. France (5), x, 
p. 146 (1880), was described from two specimens as black, with some 
greyish reflections. Eyes hairy. Antennae black. Frontal callus 
cordiform with linear extension. Abdomen with segmentations 
narrowly white and three series small indistinct grey spots. Legs 
black, tibiae pale testaceous. ? Persia, Caucasus. 

Tabanus niveipalpis, 2 , Bigot, Mem. Soc. Zool. France, v, p. 645 
(1892) [Atylotus], described from one specimen from ? Persia. Eyes 
naked. Abdomen blackish with the segmentations narrowly yellow- 
ish, and a large oval spot on the sides of second and third segment 
reddish vellow. Legs blackish, anterior tibiae whitish, the apices 
black, the other tibiae testaceous white. Wings hyaline. Length 
12mm. Bigot used this name later for a species from the Cape of 
Good Hope which is however a synonym of Tabanus albilinea, W1k. 

Tabanus canipalpis, Bigot, Mem. Soc. Zool, France, v, p. 649 
(1892) [Atylotus], from Persia was described from one specimen. 
Abdomen black with yellowish segmentations and red on the first 
two segments. Length Ir mm. 


Igit.}| G. RICARDO: Revtsion of the species of Tabanus. 247 


Tabanus polyzonatus, Bigot, Mem. Soc. Zool. France, v, p. 648 
(1892) [Atylotus|, from Persia. Abdomen black with three series 
greyish spots and segmentations greyish. Length 17 mm. 

These types are not known to me. 

Tabanus glaber, Bigot, polygonus and pulverifer, Wik., sabule- 
torum and quadrifarius, Loew, not included in Brauer’s monograph 
of species of Palaearctic Region, are therefore given here, with the 
descriptions of the four new species, one from Japan, one from 
Perso-Baluchistan Frontier, one from Cyprus and Persia and one 
from Persia. 


TABANUS abazus, Bigot (? Atylotus). 
adjacens, n. sp. 
alexandrinus, Wied. 
apricus, Mg. 
autumunalis, L. 
bifarius, Loew (Atylotus). 
bromius, L. 
canipalpis, Bigot. 
cordiger, Mg. 
cyanops, Brauer. 
cyprianus, n. sp. 
decorus, Loew (Therioplectes). 
eggeri, Schiner. 
fulvicornts = montanus, Mg., var. 
gigas, Herst. (Atylotus). 
glaber, Bigot. 
graecus, F. 
infuscatus, Loew = apricus, Mg. 
ispahanicus, Rond. [aspahanicus] = spectabilis, Loew. 
lunatus, Fabr. (Atylotus). 
montanus, Mg., var. 
muhlfeldi, Brauer (Therioplectes). 
miyajima, n. sp. 
nigrita, Fabr. 
niveipalpis, Bigot. ? Persia. 
persis, n. sp. 
polyzonatus, Bigot. 
polygonus, Walker. 
pulchellus,’ Loew. 
pulverifer, Walker. 
quadrifarius, Loew (? Atylotus). 
sabuletorum, Loew (? Atylotus). 
solstitialis, Mg. 
spectabilis, Loew. 
tricolor, Zell. (Atylotus). 
umbrinus, Mg. (Atylotus). 
unifasciatus, Loew. 


| Occurs also in Algeria, see Ricardo, Ann. Mag. Nat. Hist. (7), Xvi, p. 200 (1905), 


248 Records of the Indian. Museum. {[VoyL. IV! 


Tabanus adjacens, ¢ , 0. sp. 


Five females. Co-types from Perso-Baluchistan Frontier in 
Indian Museum coll. 

A small species with reddish yellow or yellow abdomen and a 
large broad black median stripe, yellowish legs and antennae. 
Forehead with two small callosities. Length 8}—10 mm. 

Face brown covered with grey tomentum. Beard very scanty, 
white. Palpi whitish, large and nearly the same width throughout. 
Antennae vellow, the third joint with little more than an angle, 
broad at base. Forehead parallel, about three and a half times as 
long asit is broad, the frontal callus small, heart-shaped, not reaching 
the eyes, the median one about the same size, irregular in shape, 
both reddish brown. Forehead and subcallus same colour as face. 
Thorax shining black, no trace of stripes, but with thick grey 
tomentum on the anterior border, and on sides which are yellowish. 
Breast blackish covered with grey tomentum. Scutellum black. 
Abdomen bare, shining, reddish yellow, the broad black median 
stripe with irregular lateral borders, frequently extending on the 
posterior borders of segments to the sides of abdomen as narrow 
segmentations, and on the last three segments occupying their 
whole surface with the exception of the sides ; under side yellowish. 
Legs yellowish, the fore tibiae at apices, and fore tarsi brown. Wangs 
very hyaline, the veins brown, yellow on the fore border and at 
base, with a long appendix, the stigma vellow. None of the speci- 
mens are in good condition and are probably denuded. 


Tabanus cyprianus, @ | n. sp. 


’ 


Type ¢ from Kelopside, Cyprus (Dr. G. A. Williamson), and 
two females frotn Seistan, Persia (A. S. Leese); these last in the 
Howlett coll., with label ‘‘ from horse, very common.’’ 

This species Atylotus-like in the shape of head, though no hairs 
are visible on eyes with a lens, does not appear to have been des- 
cribed betore, though it may be related to Tabanus agricola, Wied. ; 
it is related to the group containing Afylotus ditaeniatus, fulvus, 
rusticus, latistriatus and fuscipes. 

Seistan is a province in Eastern Persia bordering Afghanistan, 
so that the species presumably has a wide range. 

A small yellowish grey species with clear wings, yellow legs and 
antennae, broad forehead with two small callosities, abdomen yellow- 
ish at base and on segmentations, covered with grey tomentum and 
with four series small black spots. Length 12-13 mm. 

Face covered with greyish white tomentum and with white 
pubescence. Beard white. Palpi yellowish white with white pubes- 
cence, almost straight, not so curved as usual, stouter at base 
ending in a point. Antennae pale yellow, the third joint more 
reddish yellow, the first two joints with chiefly white hairs, but a 
few black ones are visible, first joint small, not cap-shaped, second 
very small, third with hardly any tooth but only an angle, broad at 


Igti.] G. Ricarpo: Revision of the species of Tabanus. 249 


base for its size. Horehead and subcallus same colour as face, the 
former about four and a half times as long as it is broad, parallel, 
the frontal callus small, square or oval, not reaching eyes, the median 
one smaller, both shining, mahogany-brown, hind part of head with 
white hairs, pubescence on forehead scanty, yellowish. Eyes with 
no bands. 

Thorax, scutellum and abdomen blackish but covered with grey 
tomentum appearing greyish, and clothed with short yellowish or 
white pubescence, some black hairs on shoulders. Abdomen reddish 
yellow on sides of first two segments, and segmentations the same 
colour, hardly appearing so in the type, owing to the pubescence, 
four round black dots appear on the fore border of each segment 
except the last one ; under side a little paler with same pubescence, 
the black spots not present. Legs pale reddish yellow, apices 
of fore tibiae and tarsi blackish brown, the other tarsi brownish, 
pubescence on coxae and femora white. on tibiae mixed white and 
black, and on tarsi black. Wangs clear, veins brown, stigma pale 
yellow, long appendix present. 


Tabanus eggeri, Schiner. 


Reise Novara Dipt., 81 (1868). 

Tabanus intermedius, Egg., Verh. zool-bot. Ges. Wien, ix, 
389 (1859); Brauer, Denkschr. Akad. Wien, xlii, 182, 42, pls. iii 
and vi, fig. 42 (1880); Gobert, Mem. Soc. Linn. Nord. France, 

, 6 (1881); Pandellé, Revue d’Entomol., ii, 202, 4 (1883). 

This species is as yet only recorded from South Europe, Egypt 
and Asia Minor. 

Two females in Mr. Howlett’s coll. identified as this species 
from Seistan, Persia, ‘* on horse, uncommon’’ (A. S. Leese). 

The males have the first posterior cell open, not narrowed, 
the females have it closed or very considerably narrowed, in these 
specimens the latter is the case. 


Tabanus glaber, @ , Bigot. 
(Plesxivy, fo2i ae) 
Mem. Soc. Zool. France, v, p. 678 (1892). 


Type 2 from Afghanistan (in description India is given) in Brit. 
Mus. coll. Six females from Helmund, Afghanistan, Indian Bound- 
ary Commission (Aitchison), 1889, and another female labelled T. 
glaber in Bigot’s handwriting from Afghanistan, in Indian Museum 
coll. These appear to have been the original series of specimens 
from which Bigot made his type, all having a small yellow-papered 
‘“g’’ attached to them. 

One 2 in Brit. Mus. coll. from Kashgar, I. ‘Turkestan (Lansdell), 
1889, and two 2 from Seistan, Persia, in Howlett coll., °* taken from 
horse’’; these are identical though the-frontal callus differs a little 
from that of type. This species does not appear to have been 


250 Records of the Indian Museum. [VoL IV; 


described under any other name among the Palaearctic species to 
which Region it belongs. 

A species easily recognised by the broad black stripe on the 
red abdomen, with the thorax black, the wings clear, the antennae 
black, the legs black, the tibiae reddish yellow. 

Length type 14 mm. 

Face covered with grey tomentum and with white pubescence. 
Beard white. Palpi pale yellow, almost devoid of pubescence, 
with only a little short white pubescence, and an occasional black 
hair, large, ending in a fairly long apex. Antennae wholly black. 
Forehead of medium breadth, narrower anteriorly, about five times 
as long as it is wide at its narrowest point, covered with grey tomen- 
tum and with a few white hairs at vertex. Frontal callus blackish 
brown, square, reaching the eyes, at a short distance beyond it 
appears a spindle-shaped callus in the middle of forehead, united 
posteriorly to a broader irregular shaped callus on the vertex. 
Eyes with cross-bands. Thorax with grey tomentum, stripes hardly 
distinct, shoulders reddish, pubescence on dorsum consists of black 
hairs with some scattered pale yellowish hairs intermixed. 

Abdomen: the black stripe almost entirely covers the first seg- 
ment, on the next three segments it is about a third of the width 
of abdomen, and on the remaining segments almost entirely covers 
them, leaving only their sides and narrow segmentations red ; 
under side reddish, the stripe not so well marked, the pubescence on 
dorsum black with some yellow hairs, and grey tomentum on the 
red parts. Legs black, the knees of femora and the tibiae pale yellow 
or reddish yellow, the apices of fore tibiae black. Wangs clear, veins 
and stigma yellowish brown, the first posterior cell slightly narrowed 
at its opening. 


Tabanus miyajima, 7 o, n. sp. 


Type (male) from Japan (Miss Pascoe’s coll.), 1896. Co-types, 
four females from Tokio, 1909, and Japan sent by Dr. Miyajima 
in a small collection of Japanese Diptera to the British Museum ; 
unfortunately the collection was much damaged by the journey, but 
these females are sufficiently well preserved and of such a striking, 
easily distinguished appearance, as to justify their description as 
a new species. 

2. A black species, the abdomen with whitish-haired bands, 
most prominent on the fourth and fifth segments where they are 
produced in the middle as triangular spots; scutellum grey. Legs 
black, the tibiae largely yellowish white. Wings with all posterior 
cells widely open. Frontal callus club-shaped with stout linear 
extension. Length of male 18 mm., of females 18-19 mm. 

The species has a general resemblance to Tabanus significans, 
n. sp., of the Oriental Region. 

Face covered with greyish tomentum and with some silvery 
white pubescence, the sides of cheeks and the subcallus and centre 
of face immediately round base of antennae covered with yellowish 
brown tomentum, the hairs on sides of cheeks short, brown. Beard 


1g1t.| . G. Ricardo: Revision of the specics of Tabanus. 251 


silvery white. Palpi obscurely reddish, with some grey tomentum 
and with thick black pubescence, stout, ending in an obtuse point. 

Antennae incomplete, the first two joints dull red with black 
pubescence. Forehead narrow, about seven times as long as it is 
broad, very slightly narrower anteriorly, black, with greyish 
tomentum, and some black hairs, the frontal callus brownish black, 
club-shaped, a little protuberant, long, stout, not quite reaching the 
eyes, prolonged in a stout short linear extension. Thorax black, 
with grey tomentum, disposed as indistinct stripes, pubescence 
sparse, black, with some appressed fulvous hairs intermixed, shoul- 
ders with longer black hairs, at base of wings some tufts of white 
hairs. Scutellum thickly covered with grey tomentum and with 
some white pubescence on its outer border. Breast black, covered 
with grey tomentum and with white pubescence. Abdomen black, 
the first segment with only some white hairs at sides, the second 
and third with indistinct grey tomentose narrow bands on their 
posterior borders, silvery white hairs only on the sides, the fourth 
with a wider one, covered with silvery white pubescence extending 
in the middle as a short triangle not reaching the anterior border, 
on the fifth a broad median white-haired spot ; the dorsum is covered 
elsewhere with close black pubescence ; under side black, covered 
with grey tomentum and short silvery white pubescence, the last 
three segments blackish without the lighter tomentum and pubes- 
cence. Legs black, the fore tibiae on their basal half, the other 
tibiae on their basal two-thirds yellowish white ,the pubescence whitish 
on fore coxae, on femora black with white hairs intermixed on under 
side chiefly, on pale parts of tibiae white, on their dark parts and on 
all tarsi black. Wings hyaline, very slight traces of brown shading 
on the cross-veins, stigma and veins brown, no appendix. 

#7. Male is browner, the abdomen conical brownish black, 
not so thickly covered with black pubescence, scutellum brownish 
like the thorax with rather thick brownish pubescence. Legs with 
wholly black pubescence, the fore tibiae only yellowish white on 
their basal third. Face brown with brownish black pubescence, 
the palpi brownish with black hairs. Beard blackish. Antennae 
brownish black, the third joint long, slender, with a fairly prominent 
tooth at base. Eyes large, convex, with large facets on their upper 
surface reaching to the middle of the frontal triangle, the small 
ones occupying the lower part and continued as a narrow border 
to the vertex. 


Tabanus persis, , n. sp. 
(Pl, xive fig. 24.) 


Type male and female from Seistan, Persia, on the Afghanistan 
frontier, in Mr. Howlett’s coll., and others from same locality. The 
females are labelled by the collector, A. S. Leese, as taken off camels 
or horses and common, the males as uncommon and taken off 
horse, and off gauze door. 

This species, Atylotus-like in the shape of head though no hairs 
are to be discovered on the eyes with a lens, does not appear to 


252 Records of the Indian Museum. {[Vor. IV, 


have been described before; it has a general resemblance to Atylotus 
rusticus, I,., but is distinguished from it by the shape of frontal 
callosities and by its black antennae. 

A small species with greyish black abdomen reddish at sides 
of first two segments, clear wings, yellow legs, broad forehead with 
two transverse callosities, and black antennae. Length o type 
I2mm., ? type 13 mm., other females 13—15 mm. 

Face covered with whitish tomentum and thick silvery white 
pubescence, with a yellowish brown band above between the 
antennae and eyes. Palpi stout, short, ending in a point, white with 
white pubescence. Beard silvery white. Antennae black, a character 
which distinguishes it from other species of the same group, the 
first two joints with some greyish brown tomentum and black hairs, 
the third joint broader at base but with hardly a perceptible tooth, 
only an angle, with some grey tomentum on the basal part of joint. 
Subcallus and forehead covered with greyish tomentum, the latter 
hardly three times as long as it is broad, and a third narrower 
anteriorly, the frontal callus large, transverse, almost reaching the 
eyes, convex, black, shining, with its posterior border towards 
vertex irregular, the median callus is the same colour but smaller, 
not reaching the eyes, irregular in shape, in the type broken up, 
but in others heart-shaped, the pubescence of forehead short, yellow, 
scanty, thicker towards the vertex. Eyes with apparently one 
band starting from the frontal callus. Thorax blackish covered 
with grey tomentum, stripes indistinct. Scutellum the same. 
Abdomen narrow, blackish with grey tomentum, the first three 
segments reddish yellow at the sides, so that the blackish colour 
appears only as a median stripe, on each side round grey tomentose 
spots appear, and also as narrow median ones on the black colour, the 
segmentations and sides of abdomen reddish yellow, pubescence 
white at sides and on segmentations, black at apex, but always 
scanty ; under side paler, yellowish, covered with grey tomentum. 
Legs yellow, fore tibiae at apex and tarsi blackish, middle tarsi with 
the last four joints brown, the posterior tarsi almost wholly brown, 
femora with some whitish pubescence. Wangs clear, veins brown, 
no appendix or only the root of one, stigma yellow, very narrow 
and small. 

@. Similar, the thorax and abdomen more pubescent, the former 
with yellowish pubescence, the scutellum with long white hairs on 
posterior border, the abdomen with chiefly whitish yellow pubescence. 

Kyes large, bare, the large facets yellowish, very distinct, the 
smaller ones black, continuing to vertex as a narrow border, no 
stripe is visible. One male measures 16 mm., the reddish yellow 
colour of abdomen is more extended in the males. 


Tabanus polygonus, ~ 2? , Walker. 


Tast Dipt.,v, Suppl. i, p. 237, 357 (1854); Ricardo, Ann. Mag. 
Nat. Hist. (7), xvi, p. 201 (1905). 


. 


1git.] G. RICARDO: Revision of the species of Tabanus. 253 


Male (type) from Bagdad (W. K. Loftus), 50, 105; and another 
male from the same locality. 

Female (type) from same locality. 

The males would come in Brauer’s table near aufumnalis, 
spectabilis and rectus. The female, which is much smaller, may 
possibly not belong to the same species and has a slight resemblance 
in appearance to Atvlotus rusticus, L. 

A redescription may prove of use :— 

Reddish brown species, with grey median triangular spots 
on the abdomen. Eyes bare; no long hairs on the hind part of 
the head. 

Male. Head large; facets of eyes unequal, sharply divided, 
the large facets yellowish, extending over more than half the eyes, 
surrounded by a narrow zone of small facets above and a broad 
zone on the basal half. Frontal triangle grey, reddish at apex. 
Antennae red, with the apex darker ; the third joint with a small 
tooth, long and slender, the annulated portion not so long as the 
basal joint ; the ocelligerous tubercle small, brown. Face whitish, 
with white hairs ; beard white ; palpi white, the second joint oval, 
with white pubescence. Thorax blackish, with four grey stripes 
and greyish pubescence; the shoulders red; the sides reddish, 
with black hairs above and whitish ones below ; scutelium black, 
with grey tomentum. Abdomen long and narrow, reddish, with 
a row of median grey triangular spots, which on the second and 
last segments are surrounded by black margins ; the hind borders 
of the segments narrowly white, the pubescence black ; the under 
side brighter red, with some irregular black markings. Legs red; 
the coxae and femora with longish white hairs ; the tibiae with short 
black pubescence on the sides and some whitish yellow hairs on their 
flat surfaces ; tarsi with black pubescence, pulvilli yellow. Wings 
longer than the abdomen, hyaline, the stigma and veins yellow -; 
no appendix. 

Length 18 mm., width oi head 5 mm. 

Female. Eyes not large, the facets equal, with two bands 
(in the male I can see no trace of bands). Frontal callus reddish 
brown, almost square, a little wider at its base, nearly touching the 
eyes ; frontal stripe grey, forehead parallel, about five times as long 
as it is broad, with black markings (probably denuded), the middle 
callus being a black stripe not joining the frontal callus ; the vertex 
blackish ; no ocelligerous tubercle. Antennae at base red (the rest 
wanting). Frontal triangle grey. Face grey, with white hairs; 
beard white ; palpi pale yellow, the second joint long and slender, 
at the base stouter, gradually tapering to a point, nearly straight, 
with a few black hairs. On the abdomen the grey triangular spots 
are replaced by a broad stripe of greyish tomentum, the black pubes- 
cence is longer, the abdomen not so pointed as in the males. Wings 
with the first posterior cell distinctly narrowed at the apex (this 
is not the case in the male). 

Length 14 mm., width of head 4mm. _ Ricardo, Ann. Mag. Nat. 
Hist. (7), xvi, p. 201. 


254 Records of the Indian Museum. [VoL. IV, 


Tabanus pulverifer, 9 ~ , Walker. 


List Dipt., v, Suppl. 1,.p:236,,-356 (4654),;. Ricardo, Ann: 
Mag. Nat. Hist. (7), xvi, p. 201 (1905). 

Male (type) from Turkey in Asia (Loftus). 

» Three females from the same locality (co-types) are nearly 
related to Tabanus cordiger, Mg., but belong to a distinct species ; 
they differ in the following particulars :— 

Antennae wholly red, black at the apex. Forehead one-third 
narrower anteriorly, about four times as long as it is broad, frontal 
callus reddish brown. Coxae and femora reddish yellow, as are the 
tibiae, only the anterior tibiae with a brown apex ; tarsi reddish 
brown, all with white pubescence. The under side of abdomen has 
no median black stripe. The species is much lighter in colour and 
slighter in form than cordiger ; the four specimens vary in size from 
I2—I4 mm. In the female the band under the eyes is narrow and 
vellowish brown. In the male the black hairs on the hind part of 
the head are not present ; the under side of the abdomen is reddish, 
greyer at the apex. Ricardo, Ann. Mag. Nat. Hist. (7), xvi, p. 201. 


Tabanus quadrifarius, ? , Loew. 


Zeitschr. f. d. ges. Naturwis., N. F., ix, xliii, p. 414, 3 (1874). 
[ Atylotus. ] 

From Schahrud, Turkestan. 

Near Tabanus bifarius, loew, but ornamented with four black 
stripes on abdomen, and with the femora except the grey base easily 
distinctly luteous, fore tibiae except the apices and the posterior 
wholly, the posterior tarsi at base wholly luteous. Eyes with one 
band. Forehead with two black spots. Length 5#—6 lines. 
Wings 54—54 lines. Loew, Zeitschr., etc., p. 414. 

There are examples of Tabanus bifarius, Loew, in the Brit. 
Mus. coll. determined by Dr. Kertesz, from Greece and Hungary; 
these (females) have the frontal callus square, yellow, not reaching 
the eyes, the middle callus oblong, larger than the frontal callus, 
black. The femora wholly blackish, the tibiae yellow, black at 
apices, all tarsi blackish. 

Presumably Tabanus quadrifarius has likewise two calli on 
forehead and probably belongs to the subgenus Atylotus ; there is 
one female from Bagdad in Brit. Mus. coll. which may possibly 
be this species. 


Tabanus sabuletorum, 9? , Loew. 


Zeitschr. f. d. ges. Naturwis., N. F., ix, xlili, p. 414, 4 (1874). 
[? Atylotus.| 

From Schahrud, ‘Turkestan. 

Dull whitish, with the thoracic stripes and the scutellum 
cinereous. Eyes naked, banded. Forehead very wide, the lower 
callus large, the upper one in undenuded specimens irregularly 
shaped, black. First joint of antennae whitish, the second red, 


tgtt.} G. Ricardo: Revision of the species of Tabanus. 255 


the third black, towards the base widely red, above concave but 
hardly at all excised. Palpi short, thick, white, and white haired. 
Abdomen cinereous, white spotted. 

Legs black, femora greyish, knees and tibiae except the apices 
pale yellowish. Wings white, hyaline, veins however pale brown, 
auxiliary wholly and others towards base of wing dull yellowish 
with appendix. Length 5} lines, wings 44 lines. Loew, Zeitschr.., 
Cle. Pp. 4l4+ 

This species is not known to me. 


Tabanus spectabilis, ° , Loew. 


Verh. zool.-bot. Ges. Wien, viii, p. 605, 34 (1858). 

Tabanus ispahamcus, @, Rondani, Ann. Mus. Civ. Genova, 
i, p. 300, 8 (1873) [aspahanicus]. 

Rondani’s type was not labelled, but owing to its striking 
appearance, I had no difficulty in recognising the one male specimen 
from N. Persia described by Rondani, and have no doubt it is the 
male of Tabanus spectabilis, recorded from Europe and Brussa 
in Asia Minor. The type @ came from Servia. There are two 
females of this species in the Brit. Mus. coll. from Bulgaria. 


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- = 


‘ o 
eo MAMAS OARS 
: i sé 
a iew 
iy ’ | ' nh ee ; 
ie) 
im 4 ~~ = sae i Ti iss 
2 a ae aI : 
, ) 
ae ' ? } 
' 
, 
= - 
= 
- 
7 
: 
7 : a i 
i : 
- . - = 2 
= « 


Bellardia sinicus .. 
Group I ee 


xJ 
Introduction 


List of oriental speciesof Tabanus 
List of species of Tabanus from 


Persia, etc. 

Tabanus abazus 
abbreviatus 
abscondens 
adjacens .. 
agricola .. 
albilateralis 
albimedius 
albocostatus 
albofasciatus 
alexandrinus 
amaenus .. 
angusticornis 
annamitus 
apricus 
ardens : 
atrohirtus 
auriflamma 
auristriatus 
aurotestaceus 
autuninalis 
basalis .. 
bicallosus .. 
bicinctus .. 
bifarius 
birmanicus 
borniensis 
bovinus 
bromius 
brunneus . 
brunnipennis 
bubali 
canipalpis 
caerulescens 
ceylonicus 
chinensis .. 
chrysurus 
consanguineus 
cordiger 


INDEX. 


a 


Page 


170 Tabanus costalis 


128 
130 
133 
135 
136 | 
143 
149 
1607 
188 
200 
227 
1II 
231 


246 
246 
153 
IOI 
248 | 
135 
230 
156 
190 
146 
247 


crassus 
cyanops 
cyprianus 
decorus 
discrepaus 
dissimilis .. 
ditaeniatus 
dives ; 
diversifrons 
eggeri 
erythrocephalus 
explicatus 
factiosus .. 
felderi 
flavicinctus 
flavissimus 
flaviventris 
flavothorax 
flexilis 
formosiensis 
fulvicornis 
fulvimedius 
fulvissimus 
fulvus 
fumifer 
fumipennis 
fuscicauda 
fuscicornis 
fuscomaculatus 
geniculatus 
gigas 

glaber 
graecus 
hilaris 
hirticeps .. 
hirtipalpis 
hirtistriatus 
hirtus 
hoang 
humilis. 
hybridus , 
ignobilis .. 
immanis 
incultus 
indianus .. 
infuscatus 
inobservatus 
intermedius 
ixion 
japonicus 
javanus 
joidus Fi 
jucundus .. 


258 Index. 


Page 

Tabanus justorius .. . 192 | Tabanus puteus 
khasiensis x 193 quadrifarius 
lama ae as 241 tarus 
laotianus .. x 226 | rubicundus 
leucocnematus ae 131 | rubidus 
leucohirtus Se 208 | rufiventris 
leucopogon 56 184 | sabuletorum 
leucopterus oF 1 36 | sanguineus 
leucosparsus 38 187 servillei 
lunatus.. afd 247 sexcinctus 
malayensis ae 178 siamensis 
mandarinus ne 170 signifer 
melanognathus 56 226 significans 
miyajima Be 250 | simplissimus 
monotaeniatus a 159 | solstitialis 
montanus oie MA a speciosus .. 
miihlfeldi ee e247, spectabilis 
negativus - 137 | speculum .. 
nemocaliosus ne 137 stantoni 
nephodes .. ake 145 striatus 
nexus =e am 1ST, | subcallosus 
nicobarensis SP I9I subcinerascens 
nigrita.. i 247 subhirtus .. 
nigropictus a 2LO tataricus .. 
nigrotectus si 202 | tenebrosus, Coq. 
niveipalpis Hoe teh tenebrosus, WIk. 
non-optatus 56 140 | tinctothorax 
obconicus a 21S triangularis 
optatus.. ea 139 tricolor 
orientalis .. = 198 | tristis 
orientis .. ae 195 tuberculatus 
oxyceratus a 195 umbrinus 
pallidepectoratus 3: 209 uniformis 
palpalis .. A 212 unifasciatus 
pauper... is 207 univentris 
perakiensis a 204 | vanderwulpi 
persis nie a 2S varicolor .. 
polygonus sis 252 | virgo 
polyzonatus se 247 wy villei 
pratti ae ie 143 yao , 
pulchellus et 27, || yokoamensis 
pulchriventris .. 241 : Table of groups 
pulverifer .. 254 , Table of species 


pusillus .. a 242 | Therioplectes subgenus 


11) 


a PP aa 
| 


EXPLANATION OF PLATE XIII. 


Fic. 1.—Tabanus varus. 


9) 


9) 


+, 


>) 


+) 


+) 


») 


2 
3.— 
— 
5.— 
b= 
ln 
oS 
9 — 
LO. —— 
jG G—— 
L2Z2—— 


Dp) 


+) 


+) 


hirtipalprs. 


+”) 
btcallosus. 
hacinctus. 
nemocallosus. 
strtatus. 
albimeduus. 
brunntpennts. 
jucundus. 
puteus 
rubicundus. 


Ree. Ind. Mus., Vol. IV, Igtt. Plate X1Re 


G. Edwards, Bemrose Lt? Derby. 


i" 


¢ 
y 
¥ 


| Ae 


bert Ni 

\ i 

eo + 
BF yin 


EXPLANATION OF PLATE XIV. 


Fic. 13.—Tabanus stantont. 


yo 4s ge fumefer. 

joel Sue 0-52 dissimilts. 
16 =—© |. leucopogon, 

eae? er discrepans. 

ee Oe ee, ortentis. 

3 LO 3; palpalis. 

so 20 &, 

ae) ne subcinerascens. 
5 223— 35 ceylonicus. 
323. ee glaber. 


5 24.—- 4, persis. 


Rec. Ind. Mus., Vol. IV, rorr. Plate XIV 


G. Edwards, 


Bemrose,L*4 Derby 


—. 


Part V.—Revision of the Oriental Leptide. Revised and annotated Catalogue of 
Oriental Bombylide, with descriptions of new species. 


Vol. ITI, 1909. 


Part I.—The Races of Indian Rats. 


Part II.—Notes on Freshwater Sponges, X. Report on a collection of aquatic 
animals made in Tibet by Capt. F. H. Stewart in 1907, II. Note on some 
amphibious Cockroaches. Description de quelques nouvelles Cécidomyies des 
Indes. Description of new land and marine shells from Ceylon and S. India. 
Description of two new species of Cavanx from the Bay of Bengal. Remarks on 
some little known Indian Ophidia. Remarks on some forms of Dipsadomorphus. 
A pelagic Sea-Anemone without tentacles. Rhynchota Malayana, II. 


Part IIf.—Notes on the Neuroptera in the collection of the Indian Museum. New 
Indian Leptide and Bombylide, with a note on Comastes, Os.Sac., v. Heterostylum, 
Macq. Notes on the Trichoptera in the collection of the Indian Museum. 
Diagnoses of new species and varieties of Freshwater Crabs, I—3. Report on a 
small collection of Lizards trom Travancore. Descriptions of three new Cicinde- 
linze from Borneo. The relation between fertility and normality in Rats. Des- 
cription of a Barnacle of the genus Scalpellum from Malaysia. The Hemipterous 
family Poly:tenide. Notes on Freshwater Sponges, XI. Descriptions of two 
new shells from 5.India. Preliminary note on a new genus of Phylactolematous 
Polyzoa. Miscellanea. 


Part IV.—Description of a minute Hymenopterous insect from Calcutta. The Insect 


Fauna of Tirhut, No. 1. Descriptions of new species of Botta and Nemachilus. 


New Oriental Sepsine. A new species of Fredervicella from Indian lakes. Diagnoses 
of new species and varieties of freshwater crabs, No. 4. On some new or little- 
known Mygalomorph spiders from the Oriental region and Australasia. 


Vol. IV, 1910. 


No. I.—Second report on the collection of Culicidee in the Indian Museum, with des- 
criptions of new genera and species. 


Nos, II and III.—The Indian species of Papataci Fly (Phlebotomus), Taxonomic 
values in Culicide. 


No. IV.—Revision of the Oriental blood-sucking Muscide (Stomoxine, Phtlema- 
tomyia, Aust., and Pristirhynchomyia, gen. nov.). 


No. V.—A new arrangement of the Indian Anopheline. 


Vol. V, 1910. 


Part I.-—The Hydroids of the Indian Museum, I. Notes on Freshwater Sponges, XII. 
Descriptions of new Shells in the collection of the Indian Museum from Burma, 
Siam and the Bay of Bengal. Materials for a revision of the Phylactolematous 
Polyzoa of India. Studies on the aquatic Oligocheta of the Punjab. An undes- 
cribed Burmese Frog allied to Rana tigrina. Miscellanea. 


Part II.—Description d’Ophiures nouvelles provenant des derniéres campagnes de 
‘*]’Investigator ’’ dans 1’Océan Indien. Description d’Holothuries nouvelles 
appartenant au Musée Indien. The races of Indian rats, II. Description of a new 
species of Scalpellum from the Andaman sea. Descriptions of five uew species 
of marine shells from the Bay of Bengal. Notes on fish from India and Persia, 
with descriptions of new species. 


Part IiI.—A new genus of Psychodid Diptera from the Himalayas and Travancore. 
The Indian barnacles of the subgenus Smilium, with remarks on the classification 
of the genus Scalpcllum. On a sub-species of Scutigerella unguiculata, Hansen, 
found in Calcutta. The distribution of the oriental Scolopendride. Notes on 
Decapoda in the Indian Museum, I. Description of a new species of Nemachilus 
from Northern India. Notes on the larve of Toxorhynchites immtsericors, Wk. 
Description ¢f a South Indian frog allied to Rana corrugata of Ceylon. Contri- 
butions to the fauna of Yunnan, Introduction and Part I, Miscellanea. 


Part IV.—Notes .and descriptions of Indian Microlepidoptera. On some aquatic 
oligochaete worms commensal in Spongilla cartert. On Bothrioneurum iris, 
Beddard. Notes on nudibranchs from the Indian Museum. On the classification 
of the Potamonidae (Telphusidae). Catalogue of the pheasants, peafowl, 
jungle fowl and spur fowl in the Indian Museum. On certain species of Palaemon 
from Sonth India. Alluaudella himalayensts, a new species of degenerate (*) 
cockroach, with an account of the venation found in the genera Cardax and 
Alluauaella. Rhynchota Malayana, III. 


No. 


No. 


No. 


No. 


MEMOIRS 
of the 


INDIAN MUSEUM 


Vol. I. 


. 1.—An account of the Rats of Calcutta. By W. C. Hossack. Rs. 5-8. 
. 2.—An account of the Internal Anatomy of Bathynomus giganieus. By R. E, 


T,Lovp. Rs. 2. 


. 3 Aand B.—The Oligocheta of India, Nepal, Ceylon, Burma and the Andaman 


Islands, with an account of the anatomy of certain aquatic forms. By W. 
MICHAELSEN and J. STEPHENSON. Rs. 4-8. : 


4.—Investigator sicarius, a Gephyrean Worm hitherto undescribed, the type of a 
new order. By F. H. STEWART. Rs. 2. 


Vol. II. 


. 1.—Report on the Fishes taken by the Bengal Fisheries Steamer ‘‘ Golden Crown. ’’ 


Part I.—Batoidei. By N. ANNANDALE. Rs. 2. 


. 2,—An account of the Indian Cirripedia Pedunculata. Part I.—Family Lepadide 


(sensu stricto). By N. ANNANDALE. Rs. 2. 


3.—A description of the deep-sea fish caught by the R.I.M.S. Ship “ Investiga- 
tor’’ since the year 1900, with supposed evidence of mutation in Malthopsis, 
and Illustrations of the Zoology of the R.I.M.S. Ship ‘‘ Investigator,’’ Fishes, 
Plates XLIV—L, 19c9. By R. E. Lnoyvp. Rs. 4-8. 

4.—Etude sur les Chironomides des Indes Orientales, avec description de quelques 
nouvelles espéces d’Egypte. ParJ.J. KIEFFER. Rs. 2. 


Vol. IIT. 


_ 1.—Report on the Fishes taken by the Bengal Fisheries Steamer ‘‘ Golden 


Crown.’’ Part II.—Additional notes on the Batoidei. By N. ANNANDALE. Part 
III —Plectognathi and Pediculati. By N. ANNANDALE and J. T. JENKINS. 
Part I1V.—Pleuronectidae. By J. T. JENKINS. Rs. 3. 


2,—Studies in post-larval development and minute anatomy in the genera 
Scalpellum and Ibla. By F. H. StEw4RT. Rs. 4. 


Other Publications edited and sold by the Superintendent of the Indian 
Museum (also obtainable from Messrs. Friedlander & Sohn) issued 
by the Director of the Royal Indian Marine. 


Illustrations of the Zoology of the R.I.M.S. ‘‘ Investigator ’’ 1892. Fishes, Plates I to VII. 
Crustacea, Plates I to V, 1894. Fishes, Plates VII to XIII. Crustacea, Plates VI to VIII. Ech- 
inoderma, Plates I to III, 1895. Echinoderma, Plates IV and V. Fishes, Plates XIV to XVI. 
‘Crustacea, Plates IX to XV, 1896. Crustacea, Plates XVI to XXVII, 1897. Fishes, Plate XVII. 

= oon Plates XXVIII to XXXII. Mollusca, Plates I to VI, 1898. Fishes, Plates XVIII to 

XXIV. Crustacea, Plates XXXII to XXXV. Mollusca, Plates VII and VIII, 1899. Fishes, 

- Plates XXV and XXVI. Crustacea, Plates XXXVI to XLV, 1900. Fishes, Plates XXVII to 

_XXXV. Crustacea, Plates XLVI to XLVITI. Index, Part E I9Ol. Crustacea, Plates XLIX 

a % to LV. Mollusca, Plates IX to XIII, 1902. Crustacea, Plates LVI to L XVII. Crustacea, Plates 

_ LXVIII to LXXVI. Fishes, Plates XXXVI to XXXVIII, 1905. Crustacea (Malacostraca), Plates 

__LXXVII to LXXIX. Crustacea (Entomostraca), Plates I and II. Mollusca, Plates XIV to 

_ XVIII, 1907. Fishes, Plates XXXIX to XLIII. Crustacea (Entomostraca), Plates III to V. 

= Mollusca, Plates XIX ‘and XX, 1908.—Re. 1 per me Mollusca, Plates XXI to XXIII, 1909.— 
As. 8 per ‘plate. ; 


, y 


a SRE 6 


yo 


RECORDS 
of the 


INDIAN MUSEUM 


Vol. I, 1907. 


Part I.—Contributions to the Fauna of the Arabian Sea. Records of Hemiptera and 
Hymenoptera from the Himalayas. Further notes on Indian Freshwater Ento- 
mostraca. The Fauna of Brackish Ponds at Port Canning, Lower Bengal, I—IIT. 
A Sporozoon from the Heart of a Cow. Miscellanea. 


Pavt II7.—Revision of the Oriental Stratiomyide. Description of an Oligochete 
Worm allied to Chetogaster. The Fauna of Brackish Ponds at Port Canning, 
Lower Bengal, IV. Further Note on a Polyzoon from the Himalayas. Reports 
on a collection of Batrachia, Reptiles and Fish from Nepal and the Western 

_ Himalayas. The Fauna of Brackish Ponds at Port Canning, Lower Bengal, V. 
Notes on Oriental Diptera, I and II. Miscellanea. 


Pavt III.—Report on the Marine Polyzoa in the collection of the Indian Museum. 
The Fauna of Brackish Ponds at Port Canning, Lower Bengal, VI. A third note 
on Earwigs (Dermaptera) in the Indian Museum, with the description of a new 
species. Notes on Oriental Diptera, III. Description of a new snake from Nepal. 
Notes on a collection of marketable fish from Akyab, with a description of a new 
species of Lactavius. Description of two freshwater Oligochete Worms from the 
Punjab. Notes on Phosphorescence in Marine Animals. Notes on the rats of 
Dacca, Eastern Bengal. Notes on Freshwater Sponges, I—V. Miscellanea. 


Pari IV.—Nudiclava monocanthi, the type of a new genus of Hydroids parasitic 
on Fish. Preliminary descriptions of three new Nycteribiidee from India. Anno- 
tated Catalogue of Oriental Culicide. Notes on Oriental Diptera. Notes on 
Freshwater Sponges, VI, VII. Description of a new Cyprinid Fish of the genus . 
Danio from Upper Burma. Miscellanea. 


Vol. II, 1908-1909. 


Part I.—The retirement of Lieut.-Col. Alcock, with a list of his papers, etc., on Indian 
Zoology. The Fauna of Brackish Ponds at’ Port Canning, Lower Bengal, VII. 
Description of a New Dictyonine Sponge from the Indian Ocean. Notes on 
Freshwater Sponges, VIII. Remarkable cases of variation, I. Description of a 
new species of Lizard of the genus Salea, from Assam. ‘The Fauna of Brackish 
Ponds at Port Canning, Lower Bengal, VIII. Description of a new Cavernico- 
lous Phasgonurid from Lower Siam. Descriptions of new species of Marine and 
Freshwater Shells in the collection of the Indian Museum. Notes on Oriental 
Syrphide, I. Description of a new variety of Spongilla lovicata. Notes on 
Oriental Diptera, V. Miscellanea. 


Payit II,—Gordiens du Musée Indien. The Fauna of Brackish Ponds at Port Canning, 
Lower Bengal, IX. Description of a new species of Danio from Lower Burma. 
Rhynchota Malayana, I. Cimex votundatus, Signoret. Notes on Freshwater 
Sponges, IX. Fruit Bats of the genus Pteyopus inhabiting the Andaman and 
Nicobar Archipelagos. A new species of Sun-Bird obtained near Darjiling. 
Three Indian Phylactolemata. On two new species of Eagle-Rays (Myliobatide). 
Description of a new species of the genus Sesayma, Say., from the Andaman 
Islands. Descriptions of new species of Land, Marine, and Freshwater Shells from 
the Andaman Islands. 


Part III,—The Fauna of Brackish Ponds at Port Canning, Lower Bengal, X, XI. On 
some Oriental Solifugee with descriptions of new forms. The difference between 
the Takin (Gudorcas) from the Mishmi Hills and that from Tibet, with notes. 
on variation displayed by the former. On Caridina nilotica (Roux) and its 
varieties. Description of a new species of Chavaxes from the Bhutan Frontier. 
First Report on the Collection of Culicidee and Corethridee in the Indian Museum, 
with descriptions of new genera and species. Miscellanea. 


Pavt IV.—Report on a collection of aquatic animals made in Tibet by Capt. F. H. 
Stewart in 1907, I. Notes on Aculeate Hymenoptera in the Indian Museum, I. 
Indian Psychodide. Description of a new species of mouse from the Madura 
District, Madras. Some Cleride of the Indian Museum. The Fauna of 
Brackish Ponds at Port Canning, Lower Bengal, XII. Description of a new 
species of Saw-Fish captured off the Burma Coast. A new Sting Ray of the 
genus Tyygon from the Bay of Bengal. New Micro-lepidoptera from India and 
Burma. Notes on some Chrysomelid Beetles in the collection of the Indian 
Museum. Six, new Cicindelinee from the Oriental Region. Description of a 
new slug from Tibet. 


VIE. NEW ORIENTAL NEMOCEHRA. 
By E. BRUNETTI. 


In the present paper are described nearly fifty new species 
belonging to the first suborder of Diptera. Figures of some of 
them, especially wings of the somewhat closely allied species of 
Rhyphus and Dixa, will appear in my forthcoming volume on 
Nemocera (exclusive of Culicidae and Chironomidae) for the 
‘Fauna of British India”’ series. 


RHYPHIDAE., 


No less than six species of Rhyphus from the East have come 
before me, including the only one previously recorded (macult- 
pennis, Wulp); a European species, punctatus, F., about the 
identification of which there can be little doubt, and also a variety 
of the common European fenestralis, Scop. 

All these forms are rather easily separated by the following 
characters :— 


Table of spectes. 


A Antennae conspicuously vari-coloured. 
B Sub-apical clear spot in wing (at tip 
of 2nd longitudinal vein) elongated, 
and entirely clear Le .. maculipennis, Wulp 
BB ‘The above spot nearly circular, and 
enclosing a distinct dark round 
SpoeE 23% a: . pulchricornis, sp. nov. 
AA Antennae wholly black. 
C Thorax bluish ash-grey with chocolate- 
coloured stripes. 
D Wing more deeply and extensively 
marked; distinct blackish spot at 
tip, whole distal margin more or 
less light blackish grey .. .. fenestralts, Scop., var. 
nov. ¢ndicus. 
DD Wing much less deeply marked, and 
quite clear on distal portion, beyond 
the cross-veins; no apical spot 
whatever * oe .. punctatus, F. 
CC Thorax yellow, or brownish yellow, 
. with reddish brown stripes. 
E Costa darker, towards tip of wing 
showing distinctly a quite clear 


260 Records of the Indian Museum. [Vor lLyr 


square spot on costa, only descend- 

ing to the 3rd longitudinal vein, 

with a smaller contiguous spot 

below; a distinct, narrow brown 

streak beyond outer cross-vein ; 

distal part of wing below 3rd 

longitudinal, nearly clear, with pos- 

terior veins very lightly suffused .. dtstinctus, sp. nov. 
EE Costa much lighter, showing towards 

tip of wing only one, much less 

clear, oval spot, always descending 

below 3rd longitudinal vein, with- 

out any second spot adjacent; no 

brown streak outside outer cross- 

vein; distal part of wing below 

3rd_—s longitudinal, very slightly 

darker grey, and this towards the 

margin only, the posterior veins 

not individually suffused .. divisus, sp. nov. 


Rhyphus maculipennis, Wulp. 


A @ in the Indian Museum collection, dated 30-vii-10 (no 
locality given, but probably Assam), and one in my own collection 
from Peradeniya, Ceylon, xii-o7, are almost certainly this species, 
although in the former specimen both the hind legs are missing. 
The wing agrees perfectly with Van der Wulp’s plate. The an- 
tennae and abdomen in his type specimen were injured. Of the 
former, the first three (not two, as he says!) joints are yellow, 
joints 4 to 8 yellow with brownish marks, or wholly brownish, 9, 
Io quite black, 11, 12 yellowish white, 13 to 16 quite black, the 
16th with a short white style. The abdominal two basal segments 
are mainly dirty yellow, with black posterior borders, the re- 
mainder being brownish yellow with broadly black borders; the 
apical segments wholly blackish. Belly similar. The palpi are 
black. 


Rhyphus pulchricornis, mihi, sp. nov. 


@. Base of Darjiling Himalayas. Long. 3} mm. 


Head.—Frons, fully one-third width of head, reddish yellow, 
bare; vertex with a few hairs; ocellar triangle small, black. 
isolated; antennae long, 16-jointed, variegated, the joints mostly 
reddish yellow and black, with a short, white style and two or three 
bristles at apex. The antennal joints 1, 2, 3 are reddish yellow, 
4, 5, quite black, 6, 7, 8, reddish-yellow,, 9, 10; -black era 
brownish yellow, 13 to 16 black, the appearance of the antenna 
suggesting that it is liable to variation. Palpi bright reddish 
yellow, with a few bristles; proboscis very short, yellow. 


1 Quite possibly a variable character. 


IQII.] E. BruNnETTII1: New Oriental Nemocera. 261 


Thorax yellowish, with three wide, dark, soft reddish brown 
stripes. Sides yellow, with black streaks. Some bristles on the 
dorsum. Scutellum yellow, with a few bristles. 

Abdomen brownish yellow, blackish towards tip, posterior 
borders of basal segments widely black. Belly apparently con- 
colorous. 

Legs brownish yellow, minutely pubescent: tips of the 
femora, of the tibiae and of the tarsal joints blackish. 

Wings pale grey: anterior border a little yellowish on basal 
half; posterior border pale blackish, slightly deeper at tip. A dark 
irregular band from the costa, reaching to the discal cell, and 
another, of about equal width, placed between the first and the 
infuscated wing tip. The clear space immediately adjoining the 
infuscated wing tip encloses a pale blackish oblong spot placed 
lengthwise on the costa. Outer side of the discal cell with a 
black suffusion, a black round spot in the Ist basal cell; posterior 
cross-vein and the anal vein narrowly black suffused. Halteres 
pale yellowish brown. 

Described from a specimen in good condition in the Indian 
Museum collection from Siliguri (18—20-vii-07). 

N.B.—This species is very near Van der Wulp’s maculipenms, 
but I believe it quite distinct, differing in the wing marks, and in 
the wholly yellowish hind femora, these joints in Wulp’s species 
having a black ring in the middle. The palpi, too, in Wulp’s 
species are blackish and the thorax is described as yellow with 
black stripes. 


Rhyphus fenestralis, Scop., var. nov. indicus, mihi. 


o @. Himalayas and Assam. Long: 4-5 mm. 


A number of specimens in the Indian Museum, from the 
Himalayas, show a difference from the usual form of this common 
European species, the difference being constant in all the specimens 
examined. 

The apical spot in the wing is distinctly black, clearer cut, and 
of a different shape. In fenestralis (typical) it is brownish, and its 
inner sides form a right angle, and, at the edge of the wing, the 
spot extends distinctly below the 3rd longitudinal vein ; whereas 
in indicus, it is quite black, the edge much more clearly cut, and 
the distal half of the spot does not extend below the 3rd longitudi- 
nal vein, although above this vein it reaches the wing tip as usual, 
so that the inner form of the spot is not a right angle, but irregu- 
lar. Moreover, the abdomen is darker, especially in the @ in 
which sometimes the usual black bands extend over nearly all the 
abdomen. The abdomen in the o is lighter than in the 2 , and 
shows traces of a dorsal line of elongated spots. In size, the variety 
is slightly larger, and all the examples have the brown ring on the 
hind femora, but though always present, it varies in intensity, and 
is darkest in the o. 


262 Records of the Indian Museum. [VOE.rEVs 


From a careful comparison of the specimens with four 2 9 
from Austria, four ? @ taken by me in Wales, anda @ taken 
by Dr. Annandale in Mull, Scotland, 4—6-x-07 (all in the Indian 
Museum), I find no other difference from typical fenestvalis. Ihave 
no European o present to compare with, but in imdica both sexes 
appear equally common. 

This new form appears common in the Himalayas, the Indian 
Museum possessing it from Simla (7,000 ft.), 24-iv-o7 and 10-v-09 
[Annandale, common]; Matiana, Simla district (8,000 ft.), 28—30- 
iv-07 [Avnandale]; Darjiling (7,000 ft.), 5—9-viii-og [Paiva, com- 
mon); also from Ukhral, Manipur (6,400 ft.) [ Pettigrew}. 


Rhyphus punctatus, F. 


Though I have no specimen to compare them with, I have 
little doubt that three @ 9? in the Indian Museum represent this 
species, agreeing almost perfectly with Schiner’s description. ‘T'wo 
are from Kurseong, Darjiling (5,000 ft.), 10—26-ix-og [Lynch], 
and the third from the same place taken on 9g-ix-og by Dr. 
Annandale. 

Rhyphus distinctus, mihi, sp. nov. 

oe @. Darjiling district. Long. 3-4 mm. 

Head.—KHyes absolutely contiguous in ” from the conspicu- 
ously raised ocellar triangle, which occupies’ the whole vertex, 
almost to the base of the antennae. Under side of head blackish 
grey. Frons in @ one-fourth the width of the head, and, with 
vertex and face, whitish grey ; under side of head somewhat yellow- 
ish. Antennae wholly black with a little short grey pubescence, 
tips of both scapal joints sometimes narrowly brownish yellow; 
palpi black. Back of head with some soft long hairs, proboscis 
yellowish. 

Thorax brownish yellow ; with three moderately broad, some- 
what reddish brown stripes; the outer ones extending from just 
below the anterior margin, nearly to the posterior one. The middle 
stripe extends from the anterior margin nearly to the scutellum, 
tapering gradually ; often narrowly divided in front: there is also 
a small unicolorous transverse mark just below each shoulder. 

A dorso-central row of stiff hairs, gradually diminishing in 
length extends from the posterior margin forwards, thence curving 
towards and over the humeral swellings. 

There is a lateral row of 6 or 7 long stiff hairs above each 
wing and some post-alar ones. Scutellum brownish yellow, with 
a broad median brown band, and the extreme edges brown: 
metanotum shining dark brown. 

Abdomen dark reddish brown, with a moderate amount of 
soft short yellow hair; hind margins of segments more or less 
narrowly yellowish, sometimes the posterior corners of the seg- 
ments also. Basal segment sometimes lighter coloured. Belly 
yellowish, more or less dark marked, genitalia inconspicuous. 


19II.] E. BRUNETTI: New Oviental Nemocera. 263 


Legs light brownish yellow; tarsi dark; extreme tips of 
posterior femora black. ‘There is generally a more or less distinct 
broad irregular blackish band occupying about the middle third of 
the hind femora, and the hind tibiae are more or less blackish for 
some distance at both base and tips: legs minutely pubescent. 

Wings very pale grey, with brownish markings. The cross- 
veins are rather deeply but narrowly brown, whilst moderately 
dark brownish markings occur as follows: a square mark in the 
middle of the upper basal cell; a brown well-defined streak from 
the middle of the costa, narrowing gradually, passing between the 
above-mentioned square mark, and the inner transverse vein, 
crossing the base of the discal cell and continuing narrowly along 
the 5th longitudinal vein: a broader, also well-defined stripe from 
the costa (narrowing hindwards) passing clear of the outer trans- 
verse vein, terminating at the upper fork of the lower branch of 
the 4th longitudinal. 

The tip of the wing down to the 3rd longitudinal vein is 
brown, leaving in front of it a distinct square shaped, quite clear 
spot, touching the costa and contiguous to the previously des- 
cribed stripe. It may be described differently by saying the whole 
of the costa is brownish, deepening towards the tip, with a quite 
clear square spot placed just touching the tip of the 2nd longitu- 
dinal vein, extending from the costa to the 3rd longitudinal. The 
distal part of the wing from below the 3rd longitudinal down to 
a little beyond the lowest branch of the 4th, is light brownish grey 
leaving a clearer streak across the rst, 2nd and 3rd posterior cells, 
just before their centres and almost below, but a little previous 
to, the clear square costal spot. Nearly below this latter spot, but 
a little beyond it, yet just touching it in the Ist posterior cell is 
a nearly upright, oblong quite clear spot, on the outer side of which 
the brown colour is distinctly deeper for a very small space. The 
6th longitudinal vein is very narrowly and not deeply suffused. 
Halteres yellowish. 

Described from a number of both sexes from Darjiling 
(7,000 ft.), taken by Mr. Paiva, 5—8-viii-og, and a ¢ taken by 
Dr. Annandale at Kurseong, 2,000 feet below Darjiling, 4-ix-o9. 


Rhyphus divisus, mihi, sp. nov. 


o @. Himalayas. Long. 34—5 mm. 


This species considerably resembles distincius but differs in 
some minor characters, and very essentially in the wing markings. 
The row of stiff hairs behind the eyes and across the vertex is 
stronger. The basal abdominal segments in the @ are more 
yellowish on their posterior borders, and the middle line on the 
thoracic dorsum is very distinctly divided. The blackish band 
on the hind femora is confined to the @, whereas in distinctus it 1s 
sometimes present, though indistinctly, in the @. The wing 
markings are entirely different. 


264 Records of the Indian Museum. [Vor sve 


On a pale grey ground colour, the cross-veins are deeply but 
narrowly infuscated as is also the 5th longitudinal vein; the 
proximal two-thirds of the marginal cell is rather deeply blackish, 
with a slightly paler spot in a line with the outer cross-vein; the 
distal part of the wing is light blackish grey, much deeper at tip 
just above the 3rd longitudinal vein and a little below it, leaving 
a nearly clear oval spot touching the costa, at the tip of the 2nd 
longitudinal and extending downwards into the Ist posterior cell, 
a second, much fainter and more irregular clear spot occurs just 
beyond the outer cross-vein. The praefurca appears slightly 
obliterated about its middle. Halteres yellow. 

Described from a good series of both sexes in the Indian 
Museum from the following localities :— 

Darjiling (7,000 ft.), 5—12-vili-og [Jenkins and Patva, con- 
mon]; Kurseong, 10—26-ix-og [Lynch]; Gangtok, Sikhim State, 
g-ix-09; also two pairs taken im cop., Darjiling, 5-vili-og, and 
Gangtok, 8-ix-og. An apparently immature specimen from Phagu, 
Simla hills, 12-v-09 [Annandale]. 

A @ andtwo 2 @ taken at Darjiling in company with the 
others have the thoracic markings almost obliterated, and the clear 
parts of the wing more extended; but they apparently belong to 
the same species. 

N.B.—Of the three species (fenestralis, distinctus and divisus) 
taken by Mr. Paiva at Darjiling in August, 1909, he says that 
some were taken on windows and others along the roads of the 
town, where they were hovering in small swarms under the shade 
of the trees, but it would be impossible to say now whether all 
the species occurred in both habitats or not, as at the time of 
collecting, identification was impracticable. 


DEX TIDAL: 


This family, likethe Rhyphidae, contains but a single genus—, 
and this one, Dixa, has not previously been recorded from the 
East. Five species are here offered as new, as I am unable to 
identify any of them with such descriptions of Palaearctic species 
as are accessible to me. 


DIXA, Meig. 
Table of species. 


A Wing with more markings than a 
narrow transverse streak across the 
middle. 
B Wing tip not infuscated. Thoracic 
stripes dark brown, abdomen brown. 
C Wing nearly clear; only a few infus- 
cated spots, mostly in basal and 
anal cells: no deeper black spots .. montana, sp. nov. 
CC - Wing with several blackish streaks ; 
and four rather deep black marks 


IgII.| E. BRUNETTI: New Oriental Nemocera. 205 


contiguous to and below the Ist 

longitudinal vein . maculipennis ,sp. nov. 
BB Wing tip distinctly infuscated for. a 

little distance. Thoracic stripes 

brownish yellow; abdomen brownish 

yellow .. . ochrilineata, sp. nov. 
AA Wing with only a narrow ‘transv erse 

streak across the middle. Thoracic 

stripes dark brown. Abdomen 

blackish. 

D The transverse streak on the wing ex- 
tending over the posterior cross- 


Veil «. % .. bistriata, sp. nov. 
DD The transverse streak not continued 
over the cross-vein - .. bifasciata, sp. nov. 


N.B.—D. bifasciata is possibly the @ of bistriata. 


Dixa montana, mihi, sp. nov. 
@ ¢. Western Himalayas. Long. 2-3 mm. 


Head.—Frons and back of head, lighter or darker, grey. 
Proboscis robust at base, pointed, moderately long, pubescent, 
brownish yellow; paipi dark brown. Antennae: Ist joint large, 
globular, yellowish brown, flagellum of apparently 12 joints, 
brownish yellow, closely pubescent, filamentous towards the tip, 
making it difficult to discover the exact number of joints. 

Thorax.—Dorsum grey, sometimes a little yellowish, some- 
times almost whitish. Three dark brown stripes, the median one, 
which is very narrowly divided more or less by a longitudinal pale 
stripe, runs from the anterior margin to beyond the middle, pos- 
terior to which it continues, much narrowed. The two outer 
stripes which are separated from the median one by a narrow 
space, begin some distance behind the shoulder and continue to 
the posterior margin of the dorsum; the space between them, 
posterior to the broad part of the median dark stripe, being light 
grey, traversed longitudinally by the attenuated continuation of 
the median stripe. A more or less indistinct transverse streak, or 
darker space, on the shoulders, brown or grey of a lighter or 
darker shade, sometimes with traces of a thin line connecting 
them. Scutellum yellow, metanotum brownish. Sides of thorax 
brownish or brownish grey, irregularly tinted. 

Abdomen dark brown, with very sparse pale has. Geni- 
taliain ” very small, yellowish, mainly concealed. The tips of 
(presumably) a pair of small claspers are visible. Ovipositor in ¢ 
small, with a reddish brown tip. 

Legs pale brownish yellow. ‘Tips of femora and tibiae, and 
the tarsi towards the tips, blackish; the hind tibiae being very 
distinctly though not greatly incrassated. 

Wings pale grey; venation in accordance with the generic 
description. Three moderately small pale brown spots. The tst 


266 Records of the Indian Museum. [VOL aie 


embraces the end of the praefurca, the base of the fork of the 
2nd vein, and the anterior cross-vein: the 2nd spot is in the 
middle of the basal cell: the 3rd is placed across the middle of the 
5th vein, thus falling in both the 2nd basal and the anal cells. A 
very pale grey, irregular, narrow but just distinctly perceptible 
streak joins the fork of the 2nd vein to the fork of the 4th. In 
addition there are some indistinct pale grey spots placed appar- 
ently irregularly in the basal half of the wing. Halteres yellowish. 

Described from three ~ o@ and two 2 & in the Indian Museum 
collection, all captured by Dr. Annandale in the Simla district, 
with the following data: Simla (7,000 ft.), I0-v-og (type o); 
Barogh (5,000 ft.), 10-v-10; Phagu (9,000 {t.), II-v-og (type 2 ). 

Types in Indian Museum. 

Nores.—This species must bear some resemblance to the 
common European D. maculata, Mg. Of the descriptions available 
to me that of this species is the only one in which the slight but 
very distinct incrassation of the hind tibiae is mentioned. Yet 
this character, although distinctly present in all five Oriental 
species treated of here, and apparently common to both sexes, 
has not, so far as I am aware, been described as generic, unless 
indeed, Macquart is referring to this peculiarity in his words 
‘““jambes terminées par deux pointes, peu distinctes.’’ 

In the European species, maculata, the thorax is described as 
pale yellow, and there are stripes on the sides of the thorax, the 
base of the wing is yellowish, and lesser differences also are appa- 
rent between Meigen’s species and the present one, which may be 
regarded as distinct. 


Dixa maculipennis, mihi, sp. nov. 


@ @. Eastern and Western Himalayas. Long. 2-2} mm. 


Head.—Back of head, vertex and frons light grey. Proboscis 
yellowish, distinctly dark brown at base; palpi dark brown. 
Antennae with two very distinct basal. (scapal) joints, the Ist 
circular, very short, the 2nd subglobular, both orange-yellow : 
flagellum of at least 14 joints, possibly one or two more, if such 
are shorter, as they are not at all easily counted. 

Thorax rather bright yellowish. Three dark brown thoracic 
stripes as in the previous species ; and on each shoulder a distinct 
but not sharply defined brownish, curved, transverse streak, 
connecting the tip of the median stripe with the tip of the outer 
stripe. Prothorax forming a sort of imperfect collar, swollen on 
each side into an elongate lobe, lying close to the thorax. Scutellum 
dark brown, with a broad yellow median indistinct stripe; meta- 
notum dark brown. A small brown scutellar bridge with a 
narrow yellow upper margin, joins the scutellum on each side to 
the wing base. 

Abdomen brownish, a small blackish mark towards each side 
on the posterior margins, the segments becoming blackish towards 


19II.| E. BRunETI1: New Oriental Nemocera. 267 


the tip of the abdomen. Genitalia in ~ black, a pair of claspers, 
large, conical, fleshy, the Ist joint of which is vellowish; the 2nd 
joint is also conical, smaller, apparently pointed at the tip. (The 
claspers are closed together so that exact vision is impossible.) 
Ovipositor blackish. 

Legs pale yellowish; tips of femora and tibiae, and the tarsi 
towards the tips, narrowly black: hind tibiae slightly but distinctly 
incrassated at tips. 

Wings.—Venation normal, Pale grey, costal cell unmarked, 
the brown spots beginning on the Ist longitudinal vein. Four 
distinct brown spots in a row, with fairly clearly cut sides, are 
placed in juxtaposition to this vein; the Ist small, rounded, near 
the base ; the 2nd, larger, squarish, below the tip of the auxiliary 
vein, both these two spots limited posteriorly by the 4th vein. 
The 3rd spot is a streak, beginning beyond the origin of the 2nd 
vein, and ending on the 4th vein, proximad of the anterior cross- 
vein. The 4th spot is over the fork of the 2nd vein. Much 
lighter pale brownish grey smail spots are placed, possibly irregu- 
larly over the rest of the wing. In the single specimen present 
they are situated as follows: Two in the 2nd basal cell, two in 
the anal cell, one in the axillary cell; one each in the marginal, 
submarginal, and the Ist posterior cells, all united more or less 
into a streak, placed towards the tip of the wing. ‘wo each in the 
basal (one basal, one central) and 2nd submarginal cells, also in 
_the Ist posterior cell ; one each at the base of the 3rd, 4th, and 
5th posterior cells. Halteres yellowish. 

Described from one o@ and one @,the o from Darjiling, 
29-v-10, taken by me, the 9 from Matiana (8,000 ft.), Simla 
district, 28—30-iv-07 [| Annandale]. 

Types in Indian Museum. 

Notre —Kasily known by its marmorated wings. 


Dixa ochrilineata, mihi, sp. nov. 


-@. Darjiling district. Long. 2 mm. 


Head light yellow. Eyes separated by a frons one-third the 
width of the head. Proboscis yellowish, with rather well-devel- 
oped terminal dark brown lips; palpi dark brown. Antennae: Ist 
scapal joint very short and indistinct, 2nd large, globular, yellow : 
flagellum of 14 to 16 joints (2), dark brown, extreme base of Ist, 
yellow. 

Thorax distinctly gibbous, prominent in front, yellow. ‘Three 
brownish yellow dorsal stripes, somewhat normally placed, the 
median one attaining the anterior margin, the outer ones dis- 
tinctly curved towards and over the sides anteriorly. Some soft 
long hairs in the neighbourhood of the stripes and in front of the 
wings. 

Scutellum and metanotum yellow, the former a little brown 
at the sides, the latter in the middle. Sides of thorax with a dark 


268 Records of the Indian Museum. [Vos Ve 


brown lateral stripe on a level with and passing across the bases of 
the coxae. 

Abdomen brownish yellow, a little darker towards the sides. 
Ovipositor small, inconspicuous, pale yellow. 

Legs.—Coxae and femora pale brownish yellow, femora a 
little lighter near the tips, the tips themselves blackish brown 
ringed. Tibiae and tarsi brownish. 

Wings.—Venation normal. Very pale grey, costal cell wholly 
unmarked. Tip of wing very pale blackish from beyond the fork 
of the 2nd vein, the shade extending posteriorly as far as the 2nd 
posterior cell, filling it. A dark brown streak from the Ist longi- 
tudinal vein, passing over the origin of the 3rd vein, the anterior 
cross-vein, then in diminished intensity to the hind margin of the 
wing by way of the posterior cross-vein and the last section of the 
5th longitudinal vein. A very pale blackish spot in the Ist basal 
cell, near its tip, an elongate one in the basal part of the 2nd 
basal cell, turning down at its proximal end into the anal cell. 
Wings a little yellowish at the base. Halteres pale yellow. 

Described from @ from Kurseong, 9-ix-og [Annandale|. 

Type in Indian Museum. 


Dixa bistriata, mihi, sp. nov. 


gy. Darjiling. Long. 24 mm. 

Head.—Proboscis and palpi brownish yellow, the former brown 
at the tip. Antennae: Ist scapal joint very wide and short, 
saucer-like, 2nd normal, subglobular ; flagellum of not less than 
I2 joints, closely pubescent: the whole antennae dark brown, 
except the pale yellow extreme base of the Ist flagellar joint. 

Thorax pale yellowish. Three dark chocolate-brown stripes 
of normal pattern, the median one extended somewhat linearly in 
front just below the shoulders. The shoulders themselves with a 
very pale blackish streak. 

Scutellum and metanotum dark brown, a narrow yellowish 
stripe in the middle of the former. Sides of thorax yellowish, 
with brown marks. 

Abdomen blackish, with a little pale yellow pubescence. 
Genitalia of moderate size, consisting of a pair of dark brown 
claspers, of which the second joint is cylindrical, much thinner, 
yellow ; apparently a small dorsal plate is present in addition to 
the moderately large ventral one. 

Legs pale brownish yellow, tips of femora and tibiae nar- 
rowly black ringed ; tarsi darker. 

Wings practically clear, a blackish narrow irregular streak 
in the middle from the 1st longitudinal vein, over the cross-veins, 
continued in an attenuated form along the posterior cross-vein and 
the terminal section of the 5th longitudinal vein. A very pale 
blackish streak in the basal part of the anal cell. Halteres yellow. 

Described ftom a single @ taken by myself at Darjiling, 29-v-10. 

Type in Indian Museum. 


eked Saath 


19Q1I.] E. BrunETrII: New Oriental Nemocera. 269 


Dixa bifasciata, mihi, sp. nov. 


?. Western Himalayas. Long. nearly 3 mm. 


Head. —The broad flat vertex light grey dusted, through which 
the dark ground colour can be seen. Proboscis vellowish brown, 
palpi long, concolorous. Antennae with its two scapal joints very 
distinct, dark brown; flagellum brown, except the pale yellow 
extreme base of the first joint. 

Thorax pale yellowish. Three dark brown stripes of normal 
pattern, the outer ones continued much further forward than in 
the last species. Dorsum between the stripes greyish. Scutellum 
yellowish, metanotum dark brown. Sides of thorax yellowish, 
brown marked. 

Abdomen blackish, with whitish pubescence ; belly similar. 

Legs pale yellowish, knees almost imperceptibly black, tarsi 
darker. Hind tibiae with the slight incrassation at the tip as usual, 
but yellowish. 

Wings very pale grey, a rather dark brown narrow short 
streak in the centre over the cross-veins, not extending to the 
posterior cross-vein. A pale blackish streak filling the basal half 
of the anal cell. Halteres pale yellow. 

Described from a single @ taken by Dr. Annandale at Phagu 
(9,000 ft.) in the Simla district. 12-v-09. 

Type in Indian Museum. 

Norers.—This species is very near bistriata, and may possibly 
be identical with it. The differences lie in the respective lengths of 
the Ist scapal joint and in the lengths of the thoracic stripes; the 
femora and tips tips, black ringed in one species and practically 
all yellow in the other. The wing marks in the two species may 
be differentiated as follows: In bifasctata the central streak stops 
before or at the posterior cross-vein, this vein itself not being at 
all suffused, whilst in b7strviata the suffusion is continued narrowly 
to the posterior margin. The basal pale streak in the anal cell 
is much more distinct in bifasciata than in bistriata. When placed 
side by side the two forms appear distinct, although the descrip- 
tions read so similar. 


BIBIONIDAE. 
PLECIOMYIA, mihi, gen. nov. 


Lt YPE Penthetria melanaspis, Wied., Auss. Zweifl., 1, 72 (1828) 
Plecia melanaspis, auct. 

Allied to Plecta, W., from which it differs by the 4th longitu- 
dinal vein rather broadly forking immediately at the point of con- 
tact with the anterior cross-vein. 

The 3rd longitudinal vein forks almost immediately beyond 
the anterior cross-vein, the upper branch lying almost parallel to 
the lower one. 

A third but lesser character is the presence of twelve joints 
to the antenna, two short normally shaped basal ones forming 


270 Records of the Indian Museum. [Vor: BV, 


the scape, a comparatively long first flagellar joint, followed by 
eight others of a flattened bead shape, with a moderately long, 
conical terminal joint. This is not entirely a conclusive character, 
as at least one other allied Oriental species (Plecia fulvicollis, F.) 
has twelve-jointed antennae, instead of ten- or eleven-jointed as 
in the other allied species belonging to this region. 


Pleciomyia melanaspis, Wied. 


Even as early as 1828 Wiedemann recognised a generic differ- 
ence between this species and the species of Plecia, by placing it 
in Penthetria. I have not seen any other species that can be 
referred to my new genus. 

P. melanaspts is quite common apparently all along the IHima- 
layas, the Indian Museum series representing Naini Tal, v-1893; 
Bhim Tal, 1g —22-ix-06; Theog (Simla district), 27-iv-07 ; Kurseong, 
g-ix-og ; Siliguri, 18—2o-vii-0o7 (both Darjiling district, hills and 
plains respectively). From the latter locality there is a @ and 
@ im cop. Also from Soondrijal and Katmandu (both Nepal), 
Sikhim and Mungphu in the EK. Himalayas, and Sibsagar, Shillong, 
Margherita in Assam. 

It is known from Siberia, China and Japan, and I found it 
common both at Darjiling, 2I-ix to 2-x-o8, and also at Hankow, 
China, 22—-26-1v-06, but only sparingly at Mussoorie, 20-v-05. 


PLECIA, Wied. 
Table of species. 


A The 3rd longitudinal vein forks at 
some distance beyond the anterior 
cross-vein (nearly at half the length 
of its lower branch) and always dis- 
tinctly beyond the fork of the 4th 
vein. 
B Whole thorax reddish yellow, including 
dorsum, sides, scutellum and meta- 
notum. Antennae 12-jointed .. fulvicollis, F. 
BB Dorsum of thorax and scutellum red- 
dish yellow; the sides and metano- 
tum black. Antennae 10-jointed .. tergorata, Rond. 
AA The 3rd longitudinal vein forks shortly 
beyond the anterior cross-vein (dis- 
tinctly before one-third of the length 
of its lower branch), and approxti- 
mately opposite the fork of the 4th 
vein; the upper branch lying almost 
parallel to the lower one, not almost 
erect as in Division A. 
C Not wholly black species. Dorsum of 
thorax reddish yellow. Antennae 
11-jointed HE se .. indica, sp. nov. 


IQIl.| E. BruneEtt1: New Oriental Nemocera. 271 


CC Wholly black species. 
D Legs wholly black. Long. 8—12 mm. 


Antennae I1-jointed zh .. atra, sp. nov. 
DD Legs mainly brownish. Long. 6-7 mm. 
Antennae 1r0-jointed se .. obscura, sp. nov. 


Plecia fulvicollis, F. 
P, subvarians, Wik. 
P. thoracica, Guer. 

This species appears as common in the plains of the eastern 
tropics as melanaspis, Wied., is in the higher and more northern 
parts. The Indian Museum has it from many parts of India, 
Lower Burma, Assam, Java and Ceylon. 

I have taken it myself at Jubbulpore, Meerut (both India), 
and in Java. It is known from many of the East Indian and 
Philippine Islands. 

Through the kindness of Mr. EK. K. Austen, who has examined 
the type 7 and @ of subvarians, Wik., in the British Museum, I 
am able definitely to allot this name synonymic rank, and from 
notes on thoracica, Guer., supplied by the same gentleman I have 
no hesitation, knowing the variability of /fulvicollis from the 
large number of specimens examined from many parts of the Kast, 
in regarding Guerin’s species as also identical. 

Moreover, Van der Wulp’s reference to thoracica is incorrect; 
it should be Belanger’s Voyage aux Indes Orientales (1833). 

Like many species in this family, it varies considerably in 
size, from 54 to 7 mm., one specimen in the Indian Museum 
being barely 4 mm. long. 


Plecia tergorata, Round. 


In the Indian Museum from Bhim Tal (4,500 ft.), 1g—27-ix-00 ; 
Sukna (500 ft.), I-vii-o8 [both Annandale|; Darjiling, 6—g-viii-0g9 
[Paiva]; Shan Hills, Upper Burma [J. Coggin Brown]. A pair 
in cop. from Bhim Tal, 27-1x-07. 

It is also known from Burma, Borneo and Java. 

N.B.—In spite of Rondani’s remark that the wing is wholly 
black, without trace of yellow at the base, three or four of the 
above-mentioned specimens have the wings wholly yellowish 
brown, yet there can be little doubt of their identity with this 
species. In both Pleciomyia melanaspis and Plecia fulvicollis the 
wings are sometimes more brown than black. The scapal joints 
and first flagellar joint of the antennae are relatively longer than 
in either melanaspis or fulvicollts. 


Plecia indica, mihi, sp. nov. 


@. Himalayas. Long. 6—9 mm. 


Entire body black, with the exception of the dorsum and 
upper part of the thorax, above the ridge line about the insertion 


272 Records of the Indian Museum. [Vor,.21Ve 


of the wings. ‘This part is wholly bright ferruginous red, and very 
minutely pubescent. 

Wings blackish, darker on anterior border; stigma and 
halteres black. Antennae as in atra, but the first flagellar joint 
hardly longer than those immediately following. 

Described from two @#@ and several 2 2 in the Indian 
Museum from Darjiling, 2-x-08 [Brunett:]; Theog (Simla hills), 
27-iv-0o7, and Kumaon, ix-1906 [both Annandalej; Soondrijal 
(Nepal), Kangra Valley (4,500 ft.) and Kimoli, 24-x-07. I also 
took it at Darjiling, 1o—16-x-05. 


Plecia atra, mihi, sp. nov. 


9. Nepal and Western Himalayas. Long. 8—12 mm. 


Entive body deep velvet-black, especially on the dorsum of the 
thorax. Antennae of eleven joints. the scapal two, subcylindrical, 
the 2nd being wider at the tip; the 1st flagellar joint is cup-shaped 
at its base; the following seven joints of equal size, of flattened bead 
shape, the last one, small, conical; the whole antenna straight 
and slightly pubescent. 

Abdomen trough, minutely pubescent. Legs bare, pulvilli 
greyish white. 

Wings blackish, anterior part much darker, stigma black ; 
halteres greyish black. The 4th longitudinal vein not forked 
until some distance from the anterior cross-vein. Upper branch 
of the 3rd longitudinal vein long, almost parallel to the lower one, 
originating close to the anterior cross-vein. 

Described from four @ 2 in the Indian Museum from Kumaon 
(Bhim Tal, 4,500 {t.), Ig—22-ix-06 [Annandale], and from Soondri- 
jal, Nepal. 

This is the only wholly black eastern species except my 
obscura, which is more ditty black in colour with a tendency to 
brown in the legs, besides being smaller in’size. 


Plecia obscura, mihi, sp. nov. 


o @. Western Himalayas. Long. 6-7 mm. 


Wholly dirty black, minutely pubescent. Thorax, on dorsum 
and sides sometimes brownish (in one specimen). Femora dark 
mahogany-brown, remainder of legs blackish brown. Antennae 
rather stout, black, scapal joints equal in length, short; Ist 
flagellar joint longer than each scapal joint, slightly pinched in 
the middle; remaining seven joints subequal, normal (no obvious 
minute apical joint in one example, antennae incomplete in the 
other two). 

Vertex in ~ wholly occupied by the very large cup-shaped 
ocelli, which are placed, so to speak, on their sides with their 
bases united. 

Genitalia in 7 : a pair of strong hairy claspers, two-jointed, 
the basal joint the longer and stronger, the 2nd joint ending 


TGR] E. Brunetti: New Oriental Nemocera. 273 


apparently in a single claw. In the @ the genital organ is narrow, 
cylindrical, short, ending in a pair of rather slender palp-like 
appendages. 

Wings blackish brown, darker on anterior part: venation as 
in indica, 

Described from two ~ & and one @ in my collection captured 
by me at Mussoorie, 24-v-05. Not in very good condition but the 
specific characters quite distinct enough to recognise as represent- 
ing a good species. 


BIBIO, Geoff. 
Table of spectes. 


A Thorax partly or wholly reddish yellow 
or brownish yellow (at least the 
dorsum wholly ved in all the species 
except discalis, in which it is black). 

B- Dorsum of thorax yellowish. 


C Abdomen wholly reddish yellow .- hortulanoides, 2, sp. 
nov. 
CC Abdomen wholly black at .. bicolor, Wik. 
BB Dorsum of thorax black... .. discajts, sp. nov. 
AA Thorax wholly black. 
D Abdomen wholly reddish yellow .. abdominalis, sp. nov. 


DD Abdomen wholly black. 

E Larger species; at least Io mm. long. 
Basal section of 3rd longitudinal 
vein usually much longer than the 
anterior cross-vein. 


F Species 14 mm. long ag: .. hortulanoides, ¢ , sp. 
nov. 
FF Species 10 to 12 mm. long .. .. obscuripennis, Meij. 


EE Smaller species; at most 7 mm. iong. 
Basal section of 3rd longitudinal 
vein barely, if any longer than the 
anterior cross-vein. 
G Femora wholly bright reddish or 
yellowish. 
H Femora yellowish, tibiae yellow .. Johannis, 1. 
HH Femora reddish, tibiae and tarsi black rufifemur, sp. nov. 
GG Femora wholly black or dark brown. 
I Hind tibiae reddish yellow. Veins on 
posterior part of wing distinct to 
hind margin, but paler than those 
in anterior half .. - .. fuscittbia, sp. nov. 
II Hind tibiae dark brown or black. 
Veins uniformly distinct, or those 
on posterior half of wing paler, and 
either distinctly continued to the 
hind margin or abbreviated. 


274 Records of the Indian Museum. [Vo... IV; 


J Veins on posterior half of wing, viewed 
from certain directions, as distinct 
as those on anterior half. Hind 
tibiae black b. s. .. approximatus, sp. 
nov. 
JJ Veins on posterior half of wing less 
distinct than those on anterior half. 
K Hind tibiae dark brown, lower branch 
of 4th vein and upper branch of 5th 


vein not reaching border of wing .. defectus, sp. nov. 
KK Hind tibiae black. All the veins 
attain the wing margin .. .. proximus, sp. nov. 


N.B.—The above table is constructed for the sake of con- 
venience only, and does not illustrate the affinities of the species. 
The order in which the descriptions are arranged is intended to 
represent their affinities. Schiner and others have adopted the 
relative lengths of the basal section of the 3rd longitudinal vein 
and the anterior cross-vein, as the preliminary distinction in 
separating the species, but in view of its decided variability in 
obscurtpennis and the closely allied European marci, L., it seems 
safer to distinguish the species comprised in the present paper by 
their conspicuous differences of colour. 


Bibio hortulanoides, mihi, sp. nov. 


o 2. Darling district. Vones.c 14 mime 99 12st 

Head.—In o& eyes closely contiguous from the vertex to 
immediately above the antennae, leaving a very small frontal 
triangle, the eyes with dense dark brown hair. Proboscis, palpi 
and antennae black with thick long dark brown hair, which is also 
long and thick behind the vertex. Vertical triangle conspicuously 
elevated bearing the three ocelli. 

In the ¢ the frons is one-third the width of the head, shining 
black, with some black short hair, ocellar tubercle as in 7. The 
other parts as in the ~ but the hair is more blackish than brown, 
whilst the pubescence on the proboscis, palpi, under side of head 
and behind the eyes is yellowish. 

Thovax.—In ~, dorsum, scutellum and sides shining black, 
densely covered with blackish brown hair. Inthe 2 the dorsum 
is bright brownish yellow (with microscopic concolorous pubes- 
cence), the colour very sharply delimited. The remainder of the 
thorax black, moderately shining, with short black hairs. Scu- 
tellum and scutellar ridge black ; metanotum black. 

Abdomen.—In o, shining black, wholly covered on all sides 
with thick blackish brown hair. ‘The genital organs shining black, 
obtuse, bilobed. In the 9, bright brownish yellow with short 
concoiorous pubescence; belly similar. Genital organs incon- 
spicuous. 


IQII.] E. BRUNETTI: New Oriental Nemocera. 2 
a 75 


Legs shining black, pubescent, spines on fore tibiae with a 
dull carmine tinge, pulvilli yellowish white. 

Wings brownish in @ as in obscuripennis, Meij.; in @ very 
pale grey, conspicuously lighter than in the ~, anterior border a 
little blackish in the ~, but wholly deep black in the 9. Stigma 
large, black. Halteres black. 

Described froma @ and @ in bad condition in the Indian 
Museum (labelled simply ‘“‘Ind.’’), which were returned by Bigot 
marked ‘‘ hortulanus 7 2 ?”’; also a perfect 2? obviously of the 
same species taken at Kurseong, I5-v-I0. 

Types in the Indian Museum. 

Nore.—This is either a large and well-marked variety of 
hortulanus, 1,., or distinct. ‘The differences apparent are, the much 
greater size of the ~, hortulanus being generally about 8 to II 
mim. in length; the brown, not whitish wings in the o, the yel- 
lowish hair in the 2 on the under side of the head and behind 
the eyes, the hair in this position in hortulanus ( 2 ) being black. 


Bibio obscuripennis, Meij. 


This species was abundant at Darjiling during a few days 
before and after October 16th 1905, when I captured several pairs 
in cop. in addition to a good number of males and a lesser number 
of females. My identification of the species was confirmed by Herr 
Meijere. 

Its chief distinction from marci, l., to which it bears a remark- 
ably close resemblance, is the brown wings in the ~. In marci 
they are nearly clear, with a whitish tinge. The basal section of 
the 3rd longitudinal vein in both species varies distinctly in its 
length relative to that of the anterior cross-vein, in some speci- 
mens being one and a quarter times as long, in others as much as 
nearly double, and it cannot be relied on exclusively as a specific 
character. 

The species is represented in the Indian Museum by specimens 
from the following localities : Chitlong (Nepal) ; Darjiling, 16-x-05 
[Brunetti]; 27-v-10 (6,000 ft.) [D’Abreu]; Kurseong, 23-iv-10 
[D’ Abreu]; Naini Tal, 28-ix-o7; Matiana (Simla hills, 8,000 ft.), 
28, 30-iv-07 [Annandale]; Chinese Frontier, N. W. Burma (4— 
5,000 ft.), xi-ro [C. W. Beebe]. 


Bibio proximus, mihi, sp. nov. 


@. Darjiling. Long. 5 mm. 

Head wholly black, clothed with black hairs, including dense 
pubescence on the eyes. 

Thorax wholly black, dull, dorsum rather more shining ante- 
riorly , pubescence black. 

Abdomen black with black pubescence. 

Legs black: hind femora barely incrassated posteriorly, dis- 
tinctly less so than in marci; hind tibiae more incrassated towards 


276 Records of the Indian Museum. [Vo.. IV, 


tips than in that species, giving a distinct clubbed appearance: 
hind metatarsus distinctly incrassated, the second joint of the tarsi 
two-thirds as long as the metatarsus. (In marci, the metatarsus 
is not at all incrassated, and is twice the length of the next joint.) 
Spines on fore tibiae carmine tipped. 

Wings very pale grey, only the anterior veins dark, those on 
hind part of wing normally pale: rst section of 3rd vein barely as 
long as anterior cross-vein (in marci it is always distinctly longer, 
often twice as long). Halteres black. 

Described from a single @ from Darjiling taken November, 
1910, by Mr. D’ Abreu. 

Type in Indian Museum. 

Notes.—This species is intermediate between marci, L., and 
obscuripennts, Meij. From the former it is distinguished by the 
black, not dark brown pubescence; the shortness of the basal part 
of the 3rd longitudinal vein, and the incrassation of the hind 
metatarsus. Minor differences are the lesser incrassation of the 
hind femora, the greater incrassation of the hind tibiae and the 
pale grey wings, as contrasted with the quite whitish wings of 
marct. 

When placed by the side of a specimen of marci, these differ- 
ences appear sufficiently distinctive. 

From obscuripennis, Meij., it differs by its smaller size and 
clearer wings and in the structure of the hind metatarsus, which in 
Herr de Meijere’s species is not distinctly incrassated and is twice 
the length of the following joint. 


Bibio abdominalis, mihi, sp. nov. 


o @. Western Himalayas. Long. 7-8 mm. 


Head wholly black; vertex very small, shining black, with 
small distinct ocellar triangle. Eyes (which are densely pubescent 
in the o@ with very dark brown or nearly black hair), contiguous 
from vertex to antennae, the frontal triangle very small. Antennae 
and palpi wholly black, moderately pubescent. 

Thorax shining black, closely pubescent; a fan-like bunch 
of hair in front of and another one below the root of the wing. 
Scutellum and metanotum shining black, the former with long soft 
black hairs on posterior margin, with a tendency to curl forwards ; 
metanotum bare. 

Abdomen shining black in @, with close long black pubes- 
cence ; belly similar. Genital organs confined in a hard squarish 
segment-like case. In the @ the abdomen is normally wholly 
reddish yellow, with minute sparse black pubescence; belly similar. 
Genital organs apparently comprised in a flat elevation on the 
under side of the last abdominal segment.! 


1 In the type ¢ and 2, which still remain united, both abdomens are con- 
siderably stretched out, as though the insects in life had endeavoured to separate 
themselves, yet no extension of any part of the male genitalia can be seen. There 


IgrtI.] E. BRuNETII: New Oriental Nemocera. 274 


Legs shining black, anterior femora with dense black hair, 
hind pair with the hair much sparser : tibiae and tarsi moderately 
pubescent. 

Wings dark grey in 2, a little blackish in ? , in both sexes 
somewhat iridescent in certain lights, costal cell and stigma brown- 
ish, veins dark brown. The basal section of the 3rd longitudinal 
vein is almost exactly equal to the anterior cross-vein. The 4th 
longitudinal vein forks immediately before the posterior cross-vein. 
Halteres blackish. 

Described from a type @ and @ taken im cop. by Mr. A. D. 
Imms at Badrinath (10,200 ft.), Garhwal district, 27-v-10, also 
from one additional @ and three additional 2 @ of the same 
date and locality, and a 2 taken by Dr. Annandale at Phagu 
(Simla district, 9,000 ft.), II-v-09. 

Types in the Indian Museum (also most of the other speci 
mens). 

NotTes.—One of the three females referred to above has the 
abdomen blackish except towards’ the side, where the colour is 
sufficiently distinct to identify the specimen with certainty with 
this species. 

This species is very like B. marci, L,., the common Palaearctic 
and North American species, but in that species the @ abdomen 
is wholly black as in the ~, so that in spite of the resemblance 
between the males, the specific difference between the two forms 
is indisputable. 


Bibio approximatus, mihi, sp. nov. 


? = venosus, Meig., var. 

9. Darjiling district. Long. 5-6 mm. 

Body wholly black, head much flattened, slightly pubescent. 
Dorsum of thorax with sparse short black hairs; a very small 
brownish yellow spot just below posterior corners of dorsum. 
Scutellum and metanotum black. Legs with terminal spines on 
fore tibiae reddish brown, apical spurs on posterior tibiae, brown- 
ish yellow. Hind metatarsi not at all thickened, nearly twice as 
long as the 2nd tarsal joint. Abdomen with very pale yellowish 
hairs. Ovipositor normal. 

Wings pale yellowish grey, hardly darker on anterior part ; 
veins pale brownish yellow, but clearly defined; those on hinder 
part of wing when viewed from certain directions appear almost 
as dark as the anterior veins. Stigma large, well defined, oval, 
black, a slight narrow blackish suffusion over base of 5th longi- 
tudinal vein. Basal portion of 3rd longitudinal vein almost exactly 
equal to the length of the anterior cross-vein. Halteres black, 
but basal half of stem yellowish. 


is a small thick rounded organ between the two abdomen tips of the same red 
colour as the ? abdomen, but it is not obvious to which abdomen it really 
belongs. 


278 Records of the Indian Museum. [Vor. IV, 


Described from one @ from Kurseong, November, rgio, taken 
by Mr. D’ Abreu. 

Type in Indian Museum. 

Note.—This species must bear a considerable resemblance at 
first sight to Bibio venosus, Meig., owing to the rather clear wings, 
the distinctness of the veins, and the similarity in size. Though 
the abdomen is given as only a little over 5 mm. in length, it is 
probably more in living specimens, as the single example examined 
appears to be shortened by shrinkage; otherwise the wings 
are abnormally long, their full expanse being 20 mm. The 
relative length of the hind metatarsus to the following joint is 
practically identical in both species, and it is quite possible the 
specimen before me really is a variety of venosus, Meig., bearing 
about the same relation to it as obscuripennis does to marci, the 
only suggested differences in my new species from venosus being 
the grey instead of clear wings, and the pale yellow instead of 
whitish colour of the abdominal pubescence. 


Bibio discalis, mihi, sp. nov. 


9. Western Himalayas. Long. 54 mm. 

Head.—F rons over a third the width of the head, dull black 
with sparse short black hair. Ocellar triangle normal. Pale hairs 
behind the vertex. Antennae black, scape yellowish. Tip of pro- 
boscis yellowish and the 1st joint of the palpi also, the remainder 
black, all the organs with black pubescence. 

Thorax.—Dorsum dull black, sharply delimited from the 
wholly yellowish remainder of the thorax. Prothorax consider- 
ably enlarged, conspicuously over the anterior corners of the 
dorsum. Scutellum yellowish, metanotum black. ‘The whole 
thorax, with the scutellum with pale yellow hairs. 

Abdomen black; pale yellow hairs on both dorsal and ven- 
tral sides, extreme tip and belly yellowish. 

Legs yellowish, knees narrowly black ; fore tibiae and tarsi 
(in one example) with a slight pale reddish brown tinge and the 
spurs carmine tipped ; tips of tarsi brown. 

Wings pale grey, anterior border no darker, stigma light 
brown, elongate. Halteres yellowish. 

Described from two @ 2 from Phagu, Simla district (9,000 ft.), 
II-v-og, taken by Dr. Annandale. 

Type in Indian Museum. 

Notre.—This species has considerable resemblance to B. johan- 
mis 2 , but the distinctly yellowish thorax with well-defined black 
dorsum at once distinguish it from the European species, in which 
the whole thorax is black or blackish brown. 


Bibio johannis, L. 


Two males, taken by Dr. Annandale at Matiana, 28—30-iv-07, 
and another from Theog (9,000 ft.), 2-v-07 (both places, Simla 
district), certainly represent this well-known European species. 


IgII.] E. BRUNETTI: New Oriental Nemocera. 279 


Bibio rufifemur, mihi, sp. nov. 

9. Darjiling. Long. 6-7 mm. 

Head black ; antennae with a little blackish pubescence ; some 
short grey hairs behind head. Ocelli close together on a small 
triangle on extreme vertex ; frons wholly bare. 

Thorax black, a little brownish grey hair on pleurae. Scutel- 
lum and metanotum shining black, former with a few short black 
hairs on hind margin. 

Abdomen black, roughened, with black and brown hairs. 

Legs.—Coxae shining black, with microscopic yellow pubes- 
cence, and a few longer hairs; tips reddish on outside. Femora 
bright orange-red, with sparse, very short, yellow hairs, tips black. 
Remainder of legs black, with short black or brown hairs, tips of 
tarsi joints narrowly yellowish brown. 

Wings brownish grey, darker anteriorly ; costal cell rather 
dark brown ; stigma distinctly dark brown, rather small, elongated 
egg-shaped, not touching 3rd longitudinal vein. Veins on front 
half of wing brown, 4th, 5th and 6th veins brownish yellow: 4th 
vein forks rather widely at junction with posterior cross-vein and 
just beyond tip of 2nd vein. Halteres black. 

Described from two 2 @ in my collection, taken by me at 
Darjiling, 20-x-05. 


Bibio fuscitibia, mihi, sp. nov. 
@. Western Himalayas. Long. 5-6 mm. 


Head black, with rather long and thick black hairs, mixed 
with some dark brown ones; antennae and palpi black, mouth 
greyish white. 

Thorax and scutellum shining black, black haired. 

Abdomen black, moderately shining, with blackish and dark 
brown hairs. Belly similar. 

Legs normal ; femora black with brown hairs ; fore tibiae dark 
brown, ending in a light brown spine, posterior tibiae reddish 
brown, middle pair darker, black at tips, shortly setose; hind pair 
with softer hairs, darker towards tips; tarsi brown, darker at tips ; 
hind metatarsus thickened, one-and-a-half times as long as next 
joint. 

Wings nearly clear, stigma dark brown, moderately long, and 
distinct: 4th longitudinal vein forking exactly at junction with 
outer cross-vein, the branches not widely separated. Halteres 
black. 

Described from a @ in the Indian Museum collection from 
Phakia, Kumaon district (10,700 ft.), taken by Mr. A. D. Imms, 
3-vi-09. 

N.6.—This species bears a considerable resemblance to the 
European B. laniger, Mg., but that species has greyish brown, 
thicker, woolly hair, and a comparison of specimens of the two 
species shows them to be quite distinct. 


280 Records of the Indian Museum. [VoreiVe 


Bibio defectus, mihi, sp. nov. 


@. Darjiling. Long. 3} mm. 

Head and appendages wholly black, very shortly pubescent. 

Thorax shining black, with dark brown hairs; scutellum and 
sides black. 

Abdomen black with brown hairs, greyish hairs at the tip 
and at the sides near the base. 

Legs dark brown, hind femora nearly black, somewhat long, 
distinctly though not greatly clubbed, hind tibiae a little lighter, 
with darker streaks, also clubbed. 

Wings pale brownish grey; stigma moderately large and 
distinct ; 4th longitudinal vein forking distinctly proximad of the 
posterior cross-vein, the branches moderately diverging ; lower 
branch of 4th and upper branch of 5th veins shortened, not reach- 
ing the wing-margin. Halteres black. 

Described from a unique @ from Kurseong, taken 13-viii-09 
[Paiva]; preserved in the Indian Museum. 

NotE.—This species comes in the group containing the Euro- 
pean species clavipes, varipes, etc. 


Dilophus graciosus, Big. 
Redescription. 


oa” @. N.India; China. Long. 7 3mm., 2 34-33? mm. 


@. Hyes rather bright reddish brown, some long whitish 
hairs below the head. Thorax shining, abdomen somewhat 
roughened ; posterior margin of scutellum, and a scutellar ridge 
below it, connecting it with the metanotum, bright brownish yel- 
low. ‘The whole body with short sparse pale hairs. Legs reddish 
brown, with yellowish hairs (longest on the femora) ; coxae and 
tarsi tips darker. Wings (damaged) nearly clear, stigma dark 
brown, distinct but ill-defined ; halteres black, rather large, stems 
yellow. 

2. Head black, shining, eyes long, antennae jet black. 
Thorax bright ferruginous, with a thin black dorsal line, which 
widens anteriorly and extends along the anterior borders of the 
thorax nearly to the shoulders. 

Abdomen dull ferruginous, dorsum of segments blackish, the 
first two or three segments nearly wholly blackish on upper side. 
Belly ferruginous. Legs black, all the coxae, the fore femora 
wholly, middle femora wholly, except the tips, and the basal half 
of the hind femora, bright ferruginous. Wings uniformly yellow- 
ish grey, stigma large, clear cut, black. 

Redescribed from specimens of both sexes in the Indian Museum 
including the type ~ and @, which were taken in cop. by Major 
Sage in September 1890 at Dharamsala, Western Himalayas. The 
other specimens are from Amangarh, Bijnor District, United 


EQLIs| E. BRuNETII: New Oriental Nemocera. 281 


Provinces, 24-ii-I0 ; Kurseong, 24-vi-ro ; Theog (8,000 ft.), 2-v-07 ; 
Phagu (9,000 ft.), 4I1-v-og [Annandale]; Kumaon district 
(5,700 it.), July 1909 [Jmms] ; Darjiling, 21-iv-10 [Beebe]. 

A ¢ in the same collection from Yunnan, South China, 
almost certainly represents the same species. 

Type o and @ in Indian Museum. 


Scatopse brunnescens, mihi, sp. nov. 


@. Ceylon. Long. 2 mm. 

Whole body and legs dark shining brown, slightly yellowish, 
tinged here and there with short whitish pubescence. Dorsum of 
abdomen nearly black, belly dark brownish yellow. Antennae 
with 2nd scapal joint a little longer than the Ist, followed by six 
flagellar short, wide, rounded joints, with a long (equally broad at 
its base), conical terminal joint; all the antenna with close greyish 
pubescence. 

Wings with the two anterior distinct veins joined towards the 
base by a short cross-vein, the Ist of these veins ending before 
the middle of the costa, the 2nd ending some little distance before 
the tip of the wing, at the spot where the costal vein ends. Of 
the faint veins, the Ist is perfectly straight, ending some little 
distance below the tip of the wing, forked beyond its middle, the 
branch vein being anterior to the vein itself, the prong of the 
fork ending exactly at the wing tip. The next vein is slightly 
curved, attaining the wing-border faintly : the 3rd and 4th veins 
are short and distinct, both curved rather suddenly downwards 
towards their tips to the hind margin. 

Described from one specimen from Peradeniya 
[Gravely |. 

Type in Indian Museum. 


Q-Vili- 10 


) 


Scatopse nigronitida, mihi, sp. nov. 


o @. Western Himalayas. Long. 2-24 mm. 


Wholly shining black with minute black pubescence. Femora 
with a tinge of dark brown in certain lights. Wings clear; 4th vein 
originating immediately proximad of tip of Ist, the branches 
diverging gradually up to half their length, afterwards diverging 
widely. Upper branch of 5th nearly straight ; lower branch, ort 
ginating near base, almost at right angles, nearly straight. Hal- 
teres thick, brown. 

Described from two pairs taken in cop., an additional ~ and 
2 2 2 in the Indian Museum from Dharampur (5,000 ft., Simla 
hills), 14-v-08 [Annandale]. 


282 Records of the Indian Museum. [VOL.cLvs, 


SIMULIIDAE. 
SIMULIUM. 
Table of spectes. 


A Thorax distinctly reddish brown, with 
short yellow hair ; rufithorax, sp. nov. 
AA Thorax black; at most with a dull 
reddish brown tinge occasionally. 
B Thorax with grey shoulder spots, and 
a wide greyish band on posterior 
margin gVYUSESCENS, SP. NOV. 
BB Thorax without either prey, Shoulder 
spots or posterior marginal band. 
Abdomen destitute of any sign of 
yellow hair. 
D Hind metatarsus much incrassated, 
nearly as long and as large as the 
Gl iaeee metatarsalis, sp. nov. 
DD Hind metatarsus not SO conspicuously 
incrassated: distinctly less in size 
than the tibia.! 
EK Antennae wholly black. Long.2}mm_ gvisei/rons, sp. nov. 
EE Antennae reddish yellow at base. 
Long. 13mm. .. “is .. rufibasis, sp. nov. 
CC Abdomen with short bright yellow 
hair, or with rather shagey brownish 
yellow hair. 
Abdomen with normal, short, very 
bright yellow hair. 
G Abdomen with at least the first two 
segments distinctly yellowish or 
brownish yellow, often several seg- 


ments so coloured ae .. indicum, Becher. 
GG Abdomen all black ae . aureohirtum, sp. nov. 
FF Abdomen with distinctly longer shaggy 

brownish yellow hair a . senilis, sp. nov. 


Simulium rufithorax, mihi, sp. nov. 


¢ 2. Bombay; E. Himalayas. Long. 2 mm. 


Head.—Eyes dull red, the upper facets in the ~ very large, 
the lower ones small, sharply demarcated from each other, the 
eyes being absolutely contiguous from the extreme vertex to the 
antennae. In the @ the facets uniformly small, the frons dark 
grey, nearly blackish, covered with golden yellow hairs. Proboscis 
and palpi blackish, the former with well-developed labella at the 


1 Probably a sexual character, but no better one offers itself, and the two 
forms may possibly be the 7 and 2 of the same species. 


TOLLE. E. BRUNETII: New Oriental Nemocera. 283 


tip, the latter with the last joint distinctly the largest. Antennae 
more or less reddish brown (in the type o, the scape and two 
basal flagellar joints reddish,’ in one ? wholly red, in the other 
only slightly so at the base, in all cases with whitish pubescence 
over the whole flagellum). A row of whitish hairs behind the eyes 
encircling the back of the head. 

Thorax wholly rather dull but obviously reddish brown on 
dorsum, covered with bright golden yellow hairs, sides of thorax 
wholly blackish grey. Scutellum reddish brown, metanotum dark 
grey. 

Abdomen blackish, the posterior margins of the segments very 
narrowly whitish, dorsum with a few pale scattered hairs. 

Legs.—Apparently normally the coxae and femora dull reddish 
brown or brownish yellow, the tibiae and tarsi blackish, but the 
brownish colour extends at least to the hind femora in one of the 
female specimens. Hind metatarsus much enlarged in both sexes, 
especially so in the » , in both sexes as long as or slightly longer 
than the rest of the tarsus and about two-thirds as long as the 
tibia. 

Wings clear, venation normal. Halteres brownish or blackish. 

Described from one @ from Kanara, Bombay, viii-o7, and two 
° @ from Karwar, Bombay, viii-o7. Two @ @ in the Indian 
Museum are from Kurseong, 10o—26-ix-o9 [Lynch], and 7-1x-09 
(type 2 ) | Annandale}. 

Type o in the Pusa collection, type 2 in the Indian Museum. 


Simulium grisescens, mihi, sp. nov. 
@. Darjiling district. Long. 14 mm. 


Head set very close on the thorax. Eyes with conspicuously 
large facets on upper side; a moderately narrow face below the 
antennae, light grey. Antennae black, with rather thick short 
grey pubescence, scape brownish yellow. Proboscis blackish, 
yellowish at tip; palp black. 

Thorax deep velvet-black, evidently originally covered with 
the short bright yellow hairs common to most of the Oriental 
species, shoulders broadly and squarely pale blue greyish dusted, 
the inner corner of the spot nearly rectangular; also a broad 
similarly coloured band on the posterior margin. This rather 
gives the appearance of the dorsum being of blue-grey ground- 
colour with a broad deep black transverse band occupying the 
major portion of the surface with a contiguous perpendicular 
broad stripe reaching to the anterior margin. Sides of thorax 
conspicuously blue-grey dusted; scutellum blackish, metanotum 
black. 

Abdomen black, apparently with a little brownish yellow 
pubescence; Ist segment with blue-grey shimmer, the fan-shaped 
side hairs brownish yellow. Belly black. 


! Under a strong light, the microscope reveals the whole antenna as dull red- 
dish brown, though the flagellum appears mainly black to the naked eye. 


284 Records of the Indian Museum. [Vora 


Legs.—Fore coxae pale yellowish, posterior coxae blackish 
grey. Fore femora principally brownish yellow, a little blackish 
towards the tip: posterior femora blackish, the base shortly 
yellowish. Fore tibiae and tarsi shining black, the proportions of 
the joints as in indicum, but the three first tarsal joints are 
widened, although none of them to such an extent as in that 
species: 4th and 5th joints very narrow. 

Middle tibiae black, narrowly yellowish at base, metatarsus 
yellowish, black at tip, rest of tarsus black, the joints narrowly 
yellowish at base. Hind tibiae considerably widened, black, very 
narrow at base, where it is a little yellowish; hind metatarsus 
much enlarged, basal half yellowish, the rest black: remainder of 
tarsus black, base of 2nd joint a little yellowish: hind metatarsus 
much longer than the rest of the tarsus and practically as long as 
the tibia. Both hind tibia and metatarsus with long hairs on the 
upper side. 

Wings quite clear, venation normal, halteres bright yellow, 
large and thick. 

Described from a unique type male in the Indian Museum 
from Kurseong, taken by Mr. Lynch, 10—26-ix-09. 

NotE.—This is a very distinct species, recognisable by the 
bluish grey dusted spots on the shoulder; the similarly coloured 
band on the posterior margin of the thoracic dorsum, and the first 
abdominal segment tinged with the same colour also. The third 
fore tarsal joint being widened is also a specific distinction, and 
noticeable when specimens of grisescens and indicum are placed 
side by side. 


Simulium metatarsalis, mihi, sp. nov. 


@. Darjiling district. Long. barely 2 mm. 


Head very closely applied to the thorax. Eyes contiguous 
from vertex to antennae, upper facets very large. Proboscis and 
palpi black. Antennae black, with close greyish pubescence. 

Thorax black; dorsum with close bright yellow hairs which 
appear reddish orange when viewed from certain directions. 
Scutellum black, with yellow hairs. Sides of thorax and metano- 
tum black. Lower part of thorax anteriorly, grey dusted. 

Abdomen wholly black with rather sparse black pubescence, no 
trace of gold hairs. Belly similar. The hairs towards the sides of 
the Ist segment light brownish. 

Legs.—Fore coxae yellowish, posterior pairs blackish. Fore 
femora and tibiae dull yellowish, with golden yellow hair nearly 
as far as the tip of the latter, which with the whole tarsus, is 
black, the tarsus narrow, of equal width throughout. In spite of 
the gold hair the fore tibiae viewed from above appear whitish 
grey. Middle femora and tibiae slender, basal half of each 
yellowish, with some yellow hair, apical half black, as are the 
slender middle tarsi. Hind femora mainly black, slightly pale 
yellowish at base, tibiae much incrassated, basal half yellowish, 


IOEK.| E. BRUNETTI: New Oriental Nemocera. 285 


with gold-yellow hair, metatarsus much incrassated also. Nearly 
as large as the tibia. Relatively larger than in any other Oriental 
species, the basal half yellowish, rest of tarsus black. The hind 
tibiae and metatarsi with numerous long hairs on upper side. 

Wings colourless, venation normal. MHalteres brownish. 

Described from a perfect unique @ from Kurseong, taken 
28-iii-10, by Dr. Annandale. 

Type in the Indian Museum. 

NoteE.—From the general resemblance between the two 
species, it seems possible that griseifrons may be the @ of this 
species. 

Simulium griseifrons, mihi, sp. nov. 


@. Western Himalayas. Long. nearly 2} mm. 


Head.—From between one-third and one-fourth the width of 
the head; grey dusted, without any vestige of gold hair; face 
convex, blackish. Proboscis and labella reddish brown, palpi 
black. Antennae wholly black. Back of head blackish grey 
(seen from above, whitish grey), with some black hairs, which 
form an irregular fringe behind the eyes. 

Thorax (partly denuded) black, apparently covered with 
short golden yellow pubescence. Scutellum black, covered with 
gold pubescence and with a row of long soft black hairs on hind 
margin. Sides of thorax blackish. 

Abdomen black, the extreme edge of each segment pale. 

Legs.—Fore coxae dull brownish yellow, posterior coxae black. 
Femora brownish yellow, apical half black, the colour less pro- 
nounced and less extensive on the fore pair. ‘Tibiae, basal two- 
thirds yellowish, apical third black; the outer sides of the tibiae. 
seen from above, appear whitish. Tarsi black, the anterior ones 
with metatarstis as long as the rest of the tarsus, but only slightly 
wider. Hind metatarsus considerably incrassated, distinctly 
longer than rest of tarsus, and with the basal half yellowish 
white. 

Wings clear; venation in accordance with the generic charac- 
ters. Halteres pale yellow. 

Described from one @ from Kalighat, Kumaon (6,000 ft.), 
Western Himalayas, taken by Mr. A. D. Imms, 4-vi-Io. 

Type in Indian Museum. 

NotE.—Possibly the @ of metatarsalis. ‘The only distinctive 
character between them seems to be the difference in the size of 
the hind metatarsus, and this is probably a sexual one. 


Simulium rufibasis, mihi, sp. nov. 


9. Darjiling district. Long. 14 mm. 


Head.—Frons blackish grey without trace of gold hair, face 
light ash-grey. Proboscis and labella dull dark reddish brown; 
palpi black. Antennae black, basal two or three joints reddish 
yellow. 


286 Records of the Indian Museum. [VOLeEV, 


Thorax as in previous species, except that the scutellum is 
slightly reddish brown at the tip, and that (being denuded) no 
golden yellow hair is visible on its dorsum, though such is prob- 
ably present in perfect specimens. 

Abdomen wholly black, dull. 

Legs.—Fore coxae yellowish, posterior coxae black: all 
trochanters brownish yellow. Femora black, but fore pair a little 
yellowish at the base. Fore tibiae whitish yellow, except a little 
blackish at the tip, posterior tibiae yellowish on basal half, 
blackish on apical half. Tarsi exactly as in the previous species. 

Wings clear; venation normal. Halteres bright yellow. 

Described from a2 single type @ in the Indian Museum, taken 
by Dr. Annandale at Kurseong, 18-vi-10. 


Simulium indicum, Becher. 


This species was described originally (Journ. Asiat. Soc. 
Bengal, lili, 199, pl. xiv) in 1884 from female specimens in the 
Indian Museum from Assam. 

It appears to be fairly common and widely distributed along 
the Himalayas and into Assam, the Museum possessing specimens 
from Mussoorie (7,400 ft.), 19-x-09 [Badley Scott]; Simla (7,000 
ft.) , 14-iv-07 [Annandale]; Darjiling (7,000 ft.), 12-viii-o9 [ Jenkins]; 
Kurseong, 10—26-ix-0g [Lynch]; Sylhet, 18-iv-o5 [Hall]; Khasi 
Hills, ‘‘ Assam,” 2-iii-o5 [Hall]; and Jaunsa, Tons Valley [C. G. 
Rogers]. 

Of the ~ , which has apparently not previously been recorded, 
there are two specimens from the island of Bombay, 25-xi-09, one 
of which I propose as the new type of the species, the original 
type ¢ not being traceable. 

Two other @ @ in the same collection appear to represent a 
variety with wholly black abdomen; they come from Darjiling, 
1o-viil-0g [Paiva], and Ukhral, Manipur (6,400 ft.) | Petizgrew]. 

The extent of the yellow on the abdomen is very variable, in 
the series of specimens examined. 

In general appearance the species resemble aureohirtum, but 
differs in the following points :— 

The frons is shining greyish white, the humeral calli are dis- 
tinctly and rather broadly reddish brown, the colour varying in 
extent; the sides of the thorax are distinctly grey dusted (in one 
example there is a grey dusted band along the lateral margins of 
the dorsum); the antennae are always more or less reddish brown 
at the base. The two principal specific characters are, the yellow 
colour of, generally at least, the first two abdominal segments, 
and the fore metatarsi are distinctly incrassated as well as length- 
ened. The fore coxae are yellowish, the posterior ones blackish; 
the femora and tibiae are pale on the basal half, the remainder 
being black; the fore and hind tibiae are shining whitish yellow. 
Tarsi black, posterior metatarsi pale on basal half, more so on 
hinder pair. The fore metatarsus and hind metatarsus are both 


IQII. | E. BRUNETTI: New Oriental Nemocera. 287 


lengthened and considerably incrassated; the golden yellow hair 
on the hind legs reaches to and covers the metatarsus; it is ap- 
parently sparser on the anterior legs. 


Simulium aureohirtum, mihi, sp. nov. 


o @. Assam and Bombay. Long. 2-3 mm. 


Head.—The large upper facets of the eye in the @ relatively 
smaller than in the other species, the eyes closely contiguous from 
vertex to antennae. Antennae dull reddish brown, varying to 
nearly black; sometimes scape and one or two basal flagellar joints 
pale also, the remainder blackish, occasionally the whole antennae 
brownish yellow. Frons nearly one-third of the head in ¢ , black- 
ish grey with rather thick bright vellow hair; face dark grey with 
a few yellow hairs. Proboscis and palpi blackish or dark brown. 

Thorax blackish, occasionally with a little dull reddish brown 
tinge; with close bright yellow hair covering the whole of the 
dorsum. Shoulders sometimes reddish brown, the colour occa- 
sionally extending narrowly along the anterior margin of the thorax. 
The sides blackish grey. bare. Scutellum normally black, but 
sometimes reddish brown, always with close yellow hair. Metano- 
tum blackish. : 

Abdomen blackish, with gold-yellow hair: in some specimens 
the characteristic ridge on the first segment is pale on the hind 
margin; the usual fan-shaped row of yellow hairs towards each 
side on the first segment. Belly black. 

Legs.—Normally , fore coxae brownish yellow or pale yellowish, 
posterior coxae black or blackish grey. Femora principally reddish 
brown or brownish yellow, with a broader or narrower apical band, 
this band generally wide on the posterior legs; often the fore 
femora wholly pale; sometimes the remaining femora also almost 
wholly so. ‘Tibiae generally with basal half (or rather more) 
brownish yellow, the remainder black, sometimes whoily dark 
brown or brownish black; a slight whitish grey shimmer is visible 
inl some specimens viewed from above. ‘farsi blackish or dark 
brown, basal half of metatarsus normally pale, the extent of the 
colour varying considerably. Fore tarsi ~ @ not widened, meta- 
tarsus barely as long as rest of tarsus, whole tarsus distinctly longer 
than tibia. Middle tarsi in both sexes about as long as the tibia, 
metatarsus equal to the remainder of the tarsus. Hind tarsi in 
@ considerably thickened, three-fourths as long as tibia, and 
distinctly longer than the rest of the tarsus, whole tarsus a little 
longer than tibia. Hind tarsus in @ of the same proportion as in 
the o except being less thickened. 

The coxae, femora, outer side of tibiae, and the under side of 
the hind metatarsus (the latter in @ only) bear golden yellow hair 
more or less prominently in «, more sparsely in @. 

Wings clear, venation normal. Halteres brownish or brownish 
yellow. 


288 Records of the Indian Museum. [Vor EVs 


Described from two ~ » from Umling, Assam, iii-o7 (includ- 
ing type «), and several 2 @ from Kanara, Bombay, vili-07 (in- 
cluding type 2), alsotwo @ @ from Umling, 1i1-07, all in the Pusa 
collection. A ? with all the pubescence rubbed from the abdo- 
men is probably this species. It is in the Indian Museum, from 
Kurseong, 6-ix-09. 

Types in Pusa collection. 

Nores.—Although none of the specimens are in good condi- 
tion, the specific characters as illustrated by the full series seem to 
be sufficiently distinctive from indicum to warrant the claim of the 
species to specific rank. The differences I perceive are, (i) the 
abdomen is wholly black, instead of yellowish at the base ; (11) the 
blackish grey, not whitish grey sides to the thorax ; (iii) the barely 
widened fore tarsi in the 2, compared with the conspicuously 
widened two first joints in zndicum. ‘The amount of bright yellow 
hair also appears greater in this species, especially on the legs. 


Simulium senilis, mihi, sp. nov. 


7. Western Himalayas. Long. 3 mm. 


Head.—¥yes closely contiguous, the head applied very closely 
to the thorax. Proboscis, palpi, antennae and under side of head 
wholly black. 

Thorax black, with short bright yellow hair. Sides black, 
with a little greyish reflection. Scutellum black, with bright 
yellow hair, metanotum black. 

Abdomen black with moderately sparse but rather ragged 
yellowish or brownish yellow hairs which are distinctly longer 
than in any other Oriental species, and present an appearance of 
shagginess. ‘The first segment has the characteristic (generic) 
ridge a little pale, with, towards each side, very long shaggy 
brownish yellow hair which reaches nearly to the middle of the 
abdomen. 

Legs.—Fore coxae pale yellowish, posterior coxae blackish. 
All femora yellowish, a little darker at the tip, especially the 
hind pair. All tibiae dirty yellowish, a little darker towards the 
tip ; both femora and tibiae with golden yellow hair, which is less 
obvious on the latter. Anterior tarsi black, narrow, not at all 
enlarged, metatarsus equal in length to the remainder of the 
tarsus, the whole tarsus about as long as the tibia. Hind meta- 
tarsus enlarged, two-thirds as long as the tibia, yellowish, the tip 
black, remainder of tarsus black with the bases of one or two of 
the joints a little yellowish. 

Wings colourless, venation normal. 

Described from a single @ in good condition in the Indian 
Museum taken by Dr. Annandale, 8-v-07, at Phagu (8,700 ft.), 
Simla district. : 

NotEe.—The length and somewhat shaggy appearance of the 
hair on the abdomen effectually distinguishes this species from 
other eastern ones. 


IQEL.) E. BRuNETII: New Oriental Nemocera. 289 


PSY CHODIDAE. 
Table of subfamiltes. 


The 7th longitudinal vein absent.' The 
2nd longitudinal forks at, or only very 
little before, the middle of the wing. 
Prongs of upper branch always fork 
distad of middle of wing (in P. mala- 
bavicus, at the middle). Genitalia of 
the o with three pairs of append- 
ages; ovipositor in ¢ without horny 
scimitar-shaped pair of valves . PHLEBOTOMINAE. 

N.B.—One genus only, Phlebotomus Rond., 
is Ouentt, 

The 7th longitudinal vein conspicuously 
present. The 2nd longitudinal forks 
quite near base of wing ; always before 
the first fourth of the wing’s length. 
Prongs of upper branch always fork 
proximad of middle of wing. Genitalia 
in the o with two (Psychoda Latr., 
Pericoma Wlk.) or three (Brunetiza 
Ann., Parabrunettia gen nov.) pairs of 
appendages ; ovipositor in @ consisting 
of a conspicuous horny scimitar-shaped 
projecting appendage, forming a pair 
of valves .. sd ei .. PSYCHODINAE. 


Since the publication by me, comparatively recently (Rec. 
Ind. Mus., ii, 369), of several new species of Psychoda and 
Pericoma from India, a considerable quantity of fresh material has 
accumulated at the Indian Museum, but it is to be regretted that 
the present notes and description do not in any way bring our 
knowledge of the Oriental, or even the Indian species up to date, 
as there still remains a residuum of small obscure specimens 
which I have been unable to deal with successfully. They may 
represent few species or a number of either described or of undes- 
cribed ones. Much further study is required on the limits of those 
already known before additional ones should be described. At 
present I am able to sink my Pericoma appendiculata definitely as 
the 2 of spinicornis; and P. bella as a synonym of margini- 
notata. No further specimens of Psychoda dtstincta, albonotata 
and albonigra have come before me, but good or fair series of 
bengalensis. nigripennis, and vittata have stamped these as 
good species. The first three species of Psychoda in my table 


) In Phiebotomus papatasii an exceedingly short 7th longitudinal vein occurs 
at the base of the 6th running almost direct to the wing-margin. It is liable to 
be entirely overlooked but Grassi figures it, and it is found in P. perturbans, Meij. 


290 Records of the Indian Museum. [Vor. IV, 


(l. c., p. 370) are referred to a new genus, closely allied to the 
recently established Brunettia, Annandale, but of these species, no 
further specimens have been seen of squamtpennis and argenteo- 
punctata. 

Four genera are recognised here in the Psychodinae Psychoda 
and Pericoma, which never possess conspicuously scaled wings and 
conspicuous chaetae together in the same species;' and Brunettia 
and Parabrunetiia (the latter, new) in which both scaled wings and 
tolerably or very conspicuous chaetae on the flagellar joints of the 
antennae are simultaneously present. JI am still much in doubt, 
however, as to the limits of both these latter genera. 

Eaton’s genera are, to my thinking, untenable,” yet the 
group of species with thickly scaled, broad wings, generally with 
hairs closely covering the surface also, and with distinct chaetae 
on the flagellum, seems a well-defined one ; all of these possess the 
venation of Psychoda except superstes, which, being the first 
species described, must remain typical of Brunettia. 

At the present state of our knowledge it appears to me that 
it will eventuaily be found that the three genera Psychoda, 
Pericoma and Parabrunettia all possess some species with and 
some species without: (1) areas of conspicuous extent on the 
wings covered with scales ; (2) hairs on the surface of the wings 
in addition to those on the veins; and (3) some species with the 
2nd longitudinal vein forking before (proximad of) the origin of 
the 3rd longitudinal vein whilst other species have the fork 
beyond (distad of) the origin of the 3rd vein. None of these can 
by any means be regarded as generic characters, but serve very 
well as primary divisions of the species into groups. 

The genital organs have not occupied much attention hitherto, 
but Dr. Annandale is engaged to some extent on their examina- 
tion. In dried specimens their exact observation is always 
difficult. 

The presence of the spiral chaetae (so conspicuously devel- 
oped in Brunettia superstes) in Psychoda distincta, Pericoma spini- 
cornis and margininotata, though much inferior in size, affords 
a further point on which to separate or substantiate species, 
but they must not be regarded as of more than specific value. 
Dr. Annandale has also discovered in Psychoda bengalensits and 
nigripennis a previously overlooked peculiarity in the shape of a 
pair of very small bifid chaetae on each joint (except the last) of 
the flagellum, having the appearance of two pairs of cow’s horns. 
They may, quite possibly, be present in some other species, but 
require very minute examination, being almost transparent. 


> 


| «« Conspicuous ’’ is emphasized, as very small chaetae (differing only in size 
from those in Parabrunettia), are found by Dr. Annandale to be present in some 
species of Pericoma. They appear to be absent in most species of Psychoda. 

2 A number of genera in Psychodidae, created on variations of scales on the 
wings, seems wholly undesirable, as tending to reduce the family to the level of 
taxonomic absurdity at present happily to be found only in the Culicidae. 


IgII.] E. Brunetti1: New Oriental Nemocera. 201 


PSYCHODA and PERICOMA. 


A closer study of the Indian species in these two genera 
(which represent the only Oriental ones available) results in several 
characters being found which were not made use of in my pre- 
vious paper on this family. 

Firstly there is the presence or absence of distinct scales on 
the wings—nearly always of some shade of brown, although 
sometimes appearing whitish or greyish when viewed from certain 
directions. These scales appear in only one species (apicalts) of 
Psychoda, but characterise several species in Pericoma, whilst 
in Parabrunettia they are present in some species on both the 
upper and lower surfaces of the wing, and in others on the lower 
surface only. 

In referring to these scales, allusion is not made to a certain 
number of scale-like hairs or even true scales, few in number and 
irregularly placed, at the base of the wing, and which may be 
present in many species, possibly adventitiously ; but only to 
cases where the scales are numerous, eminently conspicuous and 
arranged in regular rows, covering thickly the whole of such 
,portion of the wing-surface as they may occupy. 

The second character, which, like that of scales, is quite con- 
sistent in its occurrence or absence, is that of hairs, comparatively 
soft or at least less bristly than usual, closely covering the sur- 
face of the wing as well as the veins and arranged longitudinally 
or nearly so, so that they do not form the ‘“‘ criss-cross”’ pattern 
obvious in the case of all those species in which the hairs are 
confined to the veins. It is quite obvious at a glance to which 
category any wing belongs except in the densely scaled ones. 

The third consistent character is the position at which the 
2nd longitudinal vein forks (always near the base of the wing) : 
whether before or after (proximad or distad of) the origin of the 
3rd longitudinal vein. 

Further characters appear less definite and should best be 
applied only to the separation of closely-allied species, being 
liable to occasional variation. 

The three principal characters above specified can be used in 
both Psychoda and Pericoma.! Under Brunettia and Parabrunettia 
(two at present rather indefinitely characterised genera) are given 
some considerations relating, chiefly or wholly, to those species at 
present referred to them. 


PSYCHODA, Latr. 
Table of specves. 


A Surface of wing either without scales, or 
(when such are present) limited to a 
1 Probably they will be found to apply equally well to at least Pavabrunettia 


also, when this genus is more closely characterised and a larger number of its 
species better understood. 


292 Records of the Indian Museum. [Vors lve 


few small tufts forming spots placed 
at the tips of the veins, on the 
margin of the wing or at the forking 
of the veins: never covering any 
appreciable extent of the wing-sur- 
face.| Sometimes a few, irregular- 
ly placed, at the base of the wings. 
B Wings with the hazrs on the veins only, 
arranged in two divaricate depress- 
ed rows, one on each side, and 
generally overlapping those of the 
contiguous veins, presenting a 
““criss-cross’’ appearance to the 
whole wing. No hatrs whatever on 
the surface of the wing. No obvious 
scales, except as forming the small 
spots. 
Wing with distinct black hair-spots at 
the tips of many or all of the veins. 
D The 2nd longitudinal vein forks very 
closely proximad or distad of the 
base of the 3rd vein. 
EK Fore metatarsi wholly white scaled .. geniculata, sp. nov. 
EE Fore metatarsi never wholly white 
scaled (at the tip only in albonigra, 
and not at all in bengalensis). 
F Wings with thick black bushy hair on 
costa, with small spots (each com- 
posed of one or two white scales), 
placed along a considerable section 
of it, at apparently regular inter- 
vals. (The 2nd longitudinal vein 
forks immediately proximad of the 
base of the 3rd vein; the fork of 
the anterior branch of the 2nd vein 
opposite that of the 4th vein.) .. albonigra, Brun. 
FF Wings with only normally blackish 
grey hair; scale-spots wholly absent. 
The 2nd longitudinal vein forks 
barely distad of the base of the 3rd 
vein, practically simultaneously. 
The fork of the anterior branch of 
the 2nd vein a little distad of that 
of the 4th vein: .- AP : 
DD The 2nd longitudinal vein forks con- 
siderably distad of the base of the 
3rd vein. 


bengalensis, Brun. 


! An apparent exception is P. orbicularis, in which scales are present on the 
basal portion of the veins only, but never on the surface of the wing, that is to 
say, emanating from the surface. 


Coe] E. BRUNETTI : New Oriental Nemocera. 


G White scale-spots absent from tips of 

veins. ‘Tips of tibiae and metatarsi 

not white scaled 

GG White scale-spots present ‘at tips of 

veins. ‘Tips of tibiae and metatarsi 

with white scales ¥ 

CC Wings without hair-spots ait tips ei 
veins. 

H Veins of wings wholly without rows of 
distinct scales on any part of their 
length, nor distally in the form of 
spots. 

I The 2nd longitudinal vein forks distad 
of the base of the 3rd vein. Hairs 
on veins uniformly blackish—no 
conspicuous lighter sections of hairs. 
The middle veins (2nd, 3rd and 4th) 
nearly straight or gently bisinuate. 

J Thorax with blackish grey hair. An- 
terior branch of 2nd longitudinal 
vein forking a little beyond the 
middle of the wing, the 4th forking 
a little before the middle : 

JJ Thorax with bright yellowish brown 
hair. Anterior branch of 2nd vein 
forks as in ntgripennis, but the 4th 
vein forks near base of wing 

Il ‘The 2nd longitudinal vein forks proxt- 
mad of the base of the 3rd _ vein. 
Hairs on veins mainly blackish but 
with a transverse band of light 
brown hairs, formed by a short row 
placed beyond the middle part of 
each vein, in the distal portion of 
the wing, The middle veins (2nd, 
3rd, 4th) distinctly arcuate 

HH Veins of wings with distinct scales on 
the basal fourth of all of them 

BB Wing with the surface closely covered 
with hairs as well as the veins, the 
hairs placed much more longitudin- 
ally than in the species of B section, 
giving a longitudinal appearance to 
the pubescence instead of a criss- 
cross network. 

K Distinct spots at least near the margin 
of the wing, white or black or both, 
composed of small scales or stiff 
hairs. 


. distans, sp. nov. 


. albopicta, sp. nov. 


nigripennis, Brun. 


. orbicularis , 


. fulvohirta, sp. nov. 


. vittata, Brun. 


sp. nov. 


204 Records of the Indian Museum. [VOE Ly. 


I, The white spots on wing chiefly con- 
fined to the wing-border, or at fork- 
ing of 2nd and 4th veins. 
M Anterior branch of 2nd longitudinal 

vein forks opposite fork of the 4th 

vein. A few scales near the base of 

the wing. The small white scaly 

hair-spots placed at the tip of each 

vein, wherever they occur .. albonotata, Brun. 
MM Anterior branch of 2nd longitudinal 

vein forks a little proximad of the 

fork of the 4th vein. The white 

spots composed of white hairs, not 

scales, and placed between the tips 

of the veins, not at the tips . distincta, Brun. 
MMM Anterior branch of 2nd longitudinal 

vein forks near middle of wing, a 

little distad of the fork of the 4th 

vein. All three terminations of the 

2nd, and that of the 3rd vein with a 

pair of small spots placed very closely 

together, composed of a few elong- 

ate yellow bristly hairs. Lower 

branch of 4th vein, and the 5th 

vein, at their tips with conspicuous 

black hair-spots. Tips of other 

veins with more or less inconspi- 

cuous black hair-spots, or a tend- 

ency thereto 4. . decora, sp. nov. 
LIL, The white spots (20 or more) distri- 

buted irregularly along all the veins, 

giving the entire wing a eee 

appearance Af . maculipennts, sp.nov. 
KK _ No distinct spots near or at the wing- 

margin. With or without two 

transverse narrow bands of grey 

scale-like hairs. 

N With such transverse bands .. transversa, sp. nov. 
NN Wing wholly unmarked .. .. hirtipennts, sp. nov. 
AA Surface of wing wholly covered with 

closely-placed imbricating dark scales apicalis, sp. nov. 


Psychoda geniculata, mihi, sp. nov. 


@. Ceylon. Long. about 14 mm. 


Very near P. albonigra, mihi. 

A dark brown species with dark brown wings and conspicuous 
white spots on the legs. Wings with black and white spots near 
tips of veins. 


I9OII.] E. BRUNETTI: New Oriental Nemocera. 295 


Body clothed with thick blackish brown hair, with which is 
intermixed on the thorax, some fan-shaped tufts of whitish 
grey hair. White bristly, almost scale-like, hair on vertex and 
bristly black hair on frons and face. Antennae with very elongate 
flask-shaped joints, verticillate as in P. bengalensits; scape with 
milk-white elongate scales; chaetae present on flagellum, but 
small, curved, not S-shaped ; pubescence of flagellum greyish white. 

Legs with light brown scales and some bristly hairs. Knees 
narrowly but distinctly white, through the presence of a few snow- 
white small elongate scales. ‘Tips of tibiae and of the metatarsi 
with a circlet of small elongate snow-white scales. Front meta- 
tarsi almost wholly white scaled. Some pale stiff bristles on the 
legs but apparently no actual isolated elongate scales as in albo- 
nigra. 

Wings with hairs situated on the veins only, rich brown in 
colour, showing golden brown with violet iridescence in certain 
lights. No surface scales on the wings, the only ones present being 
the smail ones forming the spots at the tips of the veins. Black 
bristly hair-spots at tips of practically all the veins ; some white 
scaly hair-spots near tips of some of the veins, and a few on each 
of the two lower branches of the 2nd longitudinal vein and both 
branches of the 4th vein, all placed at a little before their tips. A 
larger white scaly hair-spot near the base of the costa, and some 
isolated erect white hairs placed irregularly here and there on the 
veins. The 2nd longitudinal vein forks immediately distad of the 
base of the 3rd vein, the upper branch of the former forking barely 
beyond the middle of the wing, and very little distad of the fork 
of the 4th vein. 

Described from one @ in the Indian Museum from Pera- 
deniya, Ceylon, taken by Mr. E. E. Green in October IgIo. 

It is in perfect condition and is a distinctly good species. 


Psychoda albonigra, mihi. 


To the description of this species may be added, ‘‘ knees with 
a very few small white scales; tips of the metatarsi with a circlet 
of white scales.” 

The type still remains the only known specimen. 


Psychoda bengalensis, mihi. 


In the figure of the wing of this species, no auxiliary vein is 
shown, although it is, of course, distinctly present, extending at 
least to a point beyond the fork of the 2nd longitudinal vein. 

Additional data.—Darjiling (5,000 ft.) , 3—4-vii-08 [ Annandale] ; 
(7,000 ft.), 5—8-viii-og [Paiva]; Kurseong, 24-iii-10; 25—26-vi-10 
[Annandale]; Phagu (9,000 ft.), I1-v-09; Barogh (5,000 ft.), 
I0-v-I0; Kasauli (6,300 ft.), 15-v-oS8; Dharampur (5,000 ft.), 
13-v-08 (all four localities, Simla hills, taken by Dr. Annandale) , 
Naini Tal (6—7,000 ft., Western Himalayas) [Lloyd]; Calcutta; 


296 Records of the Indian Museum. [Vor lV: 


common vi, vii, viii [dunandale and others}; Dum Dum, near 
Calcutta, 29-vii-og [Lord]; Madhupur, Bengal, 22-x-0g [Paiva]; 
Port Canning (Ganges delta), 6-xit07 [Annandale|; Bangalore, 
Mysore State (3,000 ft.), 16-x-10; Trivandrum, 13-xi-o8 ; Kulatu- 
puzha (W. base of Western Ghats, Travancore), 19-xi-o8 ; Madda- 
thorai, same district, 17-xi-08 (all three, South India, taken by Dr. 
Annandale); Peradeniya, Ceylon, 8-vi-1o [Gravely|; Moulmein, 
Lower Burma, 25-11-08 [Annandale]. 


Psychoda distans, mihi, sp. nov. 


9. South India. Long. barely 1 mm. 


Body covered with dark brown hair, mixed here and there 
with grey, the ground colour of the thorax yellowish brown, with 
dark greyish hair. Legs brown, the tarsi a little lighter, no white 
tips *> tibiae or metatarsi. 

Wings with the two divaricate rows of hairs on the veins 
only. The 2nd longitudinal vein forks considerably distad of the 
base of the 3rd; the anterior branch forking just beyond one 
third of the wing, and nearly opposite the fork of the 4th vein. 

A distinct and rather large black bristly hair-spot at the tip of 
each vein. No white scale- or hair-spots on the wing. A distinct 
though not conspicuous spot composed of black hairs, at the fork 
of the 2nd longitudinal vein and the fork of its anterior branch 
also. 

Described from a single @ in the Indian Museum from 
Maddathorai, Travancore State, South India, taken by Dr. Annan- 
dale, 16-xi-08. 

N.B.—The genital organs are not easily seen but the specimen 
appears to bea @. 


Psychoda albopicta, mihi, sp. nov. 


2. Bengal. Long. 13 mm. 


Very near P. distans, but differing from that species, by very 
distinct though small white scale-spots at the tips of most of the 
veins, including the 3rd vein. A conspicuous bunch of black 
scaly hairs at the fork of the anterior branch of the 2nd longi- 
tudinal vein, and the fork of the 4th vein. Some of the black 
hair-spots at the tips of the veins are much less distinct than in 
distans. 

Fringe of wing dark brown on costa, lighter brown on 
posterior margin, the whole appearing golden brown in certain 
lights. The hairs on the wing are brown, darker towards the 
base. 

The body is dark, with brownish yellow hair, which is darker 
brown on the thoracic dorsum, and almost yellow in places. 
Antennae brown, flagellar joints flask-shaped with long necks, 
the verticillate hairs widely expanded. Long sinuous chaetae 


LORE) E. BRUNETTI: New Oriental Nemocera. 207 


distinctly present. The tips of the tibiae and of the metatarsi 
with a narrow circlet of white scales: the legs moderately dark 
brown. 

Described from a single type 2 in the Pusa collection, taken 
at Pusa, Bengal, 8-1-08. 


Psychoda nigripennis, mihi. 


Several specimens recently acquired by the Indian Museum 
through the generosity of Mr. C. W. Beebe, are in all probability 
this species, taken by that gentleman ten miles south of Kuching, 
Sarawak, Borneo, 25-vi-10. 

Being mounted on microscopic slides their absolute identifi- 
cation is impossible, in the absence of any previous examination. 

Additional data respecting this species —Darjiling, 3-vii-08 
[Annandale|; 8—r1I-vili-og [Paiva]; Kurseong, 23—25-vi-08 
[Annandale]; Simla, 9-v-09, and Phagu (Simla district, 9,000 ft.), 
iI-v-0g | Annandale]; Calcutta, common in June, July, August ; 
24-11-10, hatched from partially dried freshwater sponge from edge 
of a garden pond; Kichha, Naini Tal (plains), 4-iv-og [Hodgart] ; 
and the following localities from South India, collected by Dr. 
Annandale: Trivandrum, 13-xi-08; Shasthancotta, near Quilon, 
7-x1-08, ‘‘ at light ;’’ Maddathorai, 16—18-x1-08 ; Tenmalai, 22-xi-08; 
Nedumangad near Trivandrum, 14-xi-o8; Ernakulam (Cochin), 
4-xi-08. 


Psychoda fulvohirta, mihi, sp. nov. 


?. Darjiling district. Long. 14 mm. 

Body covered with blackish grey hair, with the exception of 
the thorax where the hair is conspicuously bright vellowish brown, 
that of the alulae being more yellow. Antennae as in nigrifennis. 
Legs blackish, without any ornamentation. 

Wings with the appearance of those of nigripennis. The 2nd 
longitudinal vein forks distinctly distad of the base of the 3rd 
vein, and its anterior branch forks a little beyond the middle of 
the wing very considerably beyond the fork of the 4th vein, which 
occurs towards its base. The 3rd vein is gently bisinuate. 

The hairs on the wings are blackish grey, those towards the 
base being blacker, and those of the posterior marginal fringe 
somewhat greyish. 

Described from two @ @ from Darjiling (7,000 ft.), 7-viii-o9, 
type [Paiva]; and Kurseong, 26-vi-1o [Annandale|; both in the 
Indian Museum. 

N.B.—The bright yellowish brown colour of the hair on the 
thorax makes this species easily distinguishable from all others 
except those specimens of margininotata with similarly coloured 
hair, but from that species the wholly unmarked wing and un- 
ornamented tarsi will readily separate it. 


298 Records of the Indian Museum. [Vous IVE 


Psychoda vittata, mihi. 


A o@ has been taken by Dr. Annandale at Maddathorai, 
Travancore, South India, 18-xi-o8. This sex was not previously 
known, but the genital organs appear normally generic in 
character. The wings in this species are distinctly lanceolate, 
the fringe of the wings is darker on the section contiguous to the 
brownish band of hairs, and also in the neighbourhood of the 
wing tip. A good specific character is the curved nature of the 
middle veins of the wing (2nd, 3rd and 4th, with their branches), 
which distinguish it to some extent from its nearest allies, 
jugripennts and fulvohirta. 

Of the ‘‘ further specimens’’ remarked by me after the origi- 
nal description of the species, as being in the Indian Museum, two 
are now destroyed, and the other two are certainly vzttata. 

Additional data.— Several specimens in the Indian Museum, 
I1—26-vii-08 , and one, 16-xi-08, from Calcutta. 


d 


Psychoda orbicularis, mihi, sp. nov. 


9. Bengal. Long. # mm. 

Very near P. nigripennis, but differing by the presence of 
distinct elongate brown scales on the basal fourth of all the veins. 
The wing is lanceolate in shape, the entire margin very distinct, 
whence its name, the costal fringe blackish, but appearing lighter . 
if viewed from certain directions; the fringe of the posterior 
margin light greyish. The 2nd longitudinal vein forks distinctly 
distad of the base of the 3rd vein, the anterior branch forking 
immediately before the middle. The 4th vein forks distinctly 
before one-third of the wing. 

The body is covered with brown or brownish grey hair. The 
antennae as in P. nigripennis, with grey pubescence. The legs 
yellowish grey, with light greyish pubescence. 

Described from a single @ in good condition from Pusa, ~ 
21-xii-o8. In the Pusa collection. 


Psychoda albonotata, mihi. 


It should be noted that the whole surface of the wing 1s hairy 
in this species, as in all the following species. Moreover there are 
some distinct shortly elongate dark brown scales over the base of 
the wing. The Sylhet specimen noted by me ' is almost certainly 
this species and the few white scales on its abdomen were probably 
accidentally attached, not forming part at all of the insect’s 
vestiture. 


Psychoda distincta, mihi. 


In the original description of this species is an inaccuracy res- 
pecting the scales noted to occur on the basal half of the wing 


1 Rec. Ind. Mus., ii, 374. 


Ig1l1.] E. BRUNETTI: New Oriental Nemocera 299 


{n the type specimen (the only one now existing, as the one dated 
1-i-o5 has been accidentally destroyed) there is absolutely no trace 
of any such scales, either at the base of the wing or elsewhere on 
it, with the exception of the very small ones that compose the 
wing-margin spots. 

The pinned specimen dated 30-1-05 is the type; the third 
specimen (which is mounted on a slide, and which shows the 
scales mentioned), not belonging to this species. 


Psychoda decora, mihi, sp. nov. 


2. South India. Long. I mm. 


Body covered with blackish hair, the thorax covered with 
bright yellowish brown hair. Antennae with the pubescence 
rather closely compressed, as in P. mnigripennis. Legs brown, 
normally scaled and pubescent. 

Wings with dark brown and blackish hairs covering the sur- 
face as well as the veins. Two inconspicuous and incomplete 
narrow transverse lines passing across the wing similar in position 
to those in P. transversa, composed of light brown or yellowish 
brown hairs, placed in small sections. Two distinct black spots 
near the base of the wing, composed of stiff, erect long black 
thickened hairs, one spot on the base of the 2nd vein, just before 
the fork, and the other on the anterior branch just beyond the 
fork. The wing-fringe is greyish, with blacker sections here and 
there, the hair is also darker along the costa. 

Described from one ¢@ in the Indian Museum taken by 
Dr. Annandale at Tenmalai, Travancore State, South India, 
22-x1-08. 


Psychoda maculipennis, mili, sp. nov. 


? @. Ceylon. Long. I mm. 


Body with rich dark brown hair towards sides of dorsum, 
yellowish grey in middle and anteriorly, deep blackish brown on 
abdomen. Antennae with verticillate hairs thick and close, 
brownish, but showing greyish reflection. Palpi black with grey 
scales; grey scales on face, black bristly hair on frons; grey hair 
on vertex. 

Legs brownish, with concolorous bristly hair and _ scales ; 
tarsi with dirty grey scales. 

Wings with surface as well as veins closely covered with dark 
brown hairs. Wing-border with a narrow fringe of black hairs, 
which is tolerably well defined from the longer greyish fringe 
around the whole margin. At numerous and apparently irregular 
intervals these short black hairs are replaced by white or greyish 
ones, and along most of the veins irregularly placed, and at the 
tips of most of the veins are very small spots composed of a few 
white hairs. These small white spots (to the extent of twenty or 
more) give the whole wing a spotted appearance that immediately 
distinguishes it from all other species. 


300 Records of the Indian Museum. [Vous hve 


Described from a single specimen, apparently a o, in 
good condition, from Peradeniya, Ceylon, taken 17-vi-Io by 
Mr. E. E. Green. In the Indian Museum. 


Psychoda transversa, mihi, sp. nov. 


? 9. “Darilines: Wonge tenn. 

Body covered with brownish grey or brownish yellow bristly 
hairs. Antennae as in nigripennis. Legs blackish with the tarsi 
showing a lighter shade in certain lights, the tips of the joints 
with small pale scales. 

Wings densely covered with blackish hairs situated over the 
entire surface of the wings as well as on the veins. 

No hair-spots nor scale-spots at the tips of any of the veins, 
but two narrow transverse bands composed of elongated, thick- 
ened (almost scale-like) greyish white hairs, the first placed just 
before the middle of the wing, the second just beyond three- 
fourths of the wing ; both bands extending from the costa to the 
hind margin. 

Described from a single specimen, which is apparently a 2 
(the genital organ not being clearly visible), in the Indian Mus- 
eum, from Kurseong (4,700—5,000 ft.), 20-vi-10, taken by Dr. 
Annandale. 

N.B.—The two conspicuous transverse bands of grey stiff 
hairs on the wings immediately distinguish this species from all 
other Oriental ones. 


Psychoda hirtipennis, mihi, sp. nov. 


2. Daryiling district;"S. India; Bengal’ Long. “about 
14 mm. 


Body brownish yellow, with yellowish brown or brownish grey 
hair varying both in shade and intensity. Antennae practically 
as in P. mgripennis. Legs with light brown scales, tarsi not 
obviously lighter, except when viewed in certain directions. 

Wings lanceolate, the brown hairs covering the surface of the 
wing and placed mainly longitudinally, no regularly divaricate 
rows of hairs on the veins. Some erect bristly hairs on the basal 
fourth of the wing. 

Described from several specimens in the Indian Museum 
presenting the following data: Kurseong, 5-ix-og; Bangalore, 
15-x-10; Maddathorai, 18-x1-08 ; Trivandrum, 12—13-xi-08 (the last 
three places in South India, taken by Dr. Annandale); Calcutta, 
2 -xA-TO! 

N.B.—One example of the above series (from Maddathorai), 
varies more than the others by having browner legs, brighter 
brown hair on the thorax, and a tendency to patches of darker 
hairs on the wing, with light greyish reflections here and there. 
The specimen seems to come within the probable specific range 
of hirtipennts, which is certainly of variable nature. 


IgII. | E. BRUNETTI: New Oriental Nemocera. 301 


Psychoda apicalis, mihi, sp. nov. 
? 2. South India. Length of wing 2 mm. 


Body with dark brown hair on the thorax, a little lighter here 
and there. The abdomen (subsequently lost) seems to have been 
covered with small elongate whitish scales, which, when viewed in 
certain lights, show prismatic colours. Antennae as in P. nigri- 
pennts, the pubescence very dense, lying rather close, so that the 
outline of the flagellum appears to have parallel sides. Legs closely 
covered with blackish scales; tips of tibiae and of metatarsi, with 
a row of white scales, of which there are a few on the tarsi. 

Wings very lanceolate and narrow, wholly covered with 
closely placed, overlapping dark brown scales. It is difficult to 
see whether hairs are present on the actual surface of the wing or 
not, but apparently (and in all probability) they are present. 
Some are present on the veins. 

The 2nd longitudinal vein forks considerably distad of the 
base of the 3rd vein; the anterior branch forking again distinctly 
beyond the middle of the wing, a good distance beyond the fork 
of the 4th vein, which occurs just before the middle of the wing. 

Wing-border wholly unmarked, bearing a blackish brown 
fringe, darkest on the costa and at the base of the hind margin: 
a small are of nearly snow-white hairs at the tip of the wing. 

Described from a single 2 (2) in the Indian Museum from 
Maddathorai, Travancore State, South India, 16-xt-08, taken by 
Dr. Annandale. 

N.B.—Very distinct from all other species by the densely 
scaled wing, with snow-white fringe at the tip. The abdomen has 
been lost in mounting the specimen for the microscope, after the 
description was drawn up, but the length of the insect was over- 
looked. It is a small species, the wing measuring 2 mm. in 
length. The sex is not quite certain, but was noted originally 

s ‘‘probably ?.” 


PERICOMA, WIk. 
Table of spectes. 


A Wing with two rows of depressed 
scales on all the veins from the base 
to the middle where they area 
develop into bristly hairs . Ssquanunervis, Sp. nov. 
AA Wing without scales, except small ones 
in the nature of tufts forming spots 
at the tips of the veins or at or near 
their forkings. 
B_ Wing surface completely covered with 
comparatively soft black hairs, in 
addition to those on the veins .. annandaler, Brun. 
BB Wing surface wholly destitute of hairs 
(except an isolated one or two near 


302 


Records of the Indian Museum. [Vor iv; 


the margin and a few at extreme 
base of wing). The hairs placed 
exclusively on the veins. 


C Lower branch of 4th longitudinal vein 


with a distinct appendix at its basal 
angle. (The 2nd longitudinal vein 
forks proximad of the base of the 
3rd vein.) Antennae in ~ with six 
prominent erect spines on the upper 
side of the Ist flagellar joint, which 
is considerably lengthened. (An- 
tennae in @ without such spines, 
the rst flagellar joint not abnor- 
mally lengthened.) 


CC Lower branch of 4th longitudinal vein 


FF 


EE 


without such appendix at its basal 
angle. The 1st flagellar joint not 
abnormally lengthened in either sex, 
and always without the prominent 
spines. 


Wing with smail but distinct spots at 


the tips of many or all of the veins, 
composed of black or white hairs, 
scale-like hairs, or true scales, often 
a black and a white spot both 
present at the tip of the same vein. 


A distinct section of the marginal 


fringe at the tip of the wing, com- 
posed wholly of white or whitish 
hairs. 


Metatarsus normally black, except for 


an apical fringe of small white 
scales. Marginal fringe of wing 
with sections composed of grey or 
whitish grey hair. (The 2nd longi- 
tudinal vein forks proximad of the 
base of the 3rd vein.) 


Metatarsus with at least the apical 


half covered with whitish scales.! 
Marginal fringe of wing with only 
one are of whitish grey hair, which 
is apical. (The 2nd _ longitudinal 
vein forks proximad of the base of 
the 3rd vein.) .. : 


the marginal fringe at the wing-tip. 


. spinicornis, Brun. 


(appendiculata, 
Brun.) 


margininotata, Brun. 
(bella, Brun.) 


: .. metatarsalis, sp. nov. 
No distinct section of white hairs in 


1 In two out of the three specimens present before me it is wholly covered 


with the whitish scales, except rather narrowly at the base. 


IQIt.| KR. BruNETII: New Oriental Nemocera. 303 


G Distinct bright yellow hairs in short 
sections on the veins, these sections 
so situated as to form an irregular 
transverse line from the costa to 
near the distal end of the posterior 
margin. Costa with very thick 
black bristly hair. (The 2nd longi- 
tudinal vein forks distad of the base 
of the 3rd vein.) . mxta, sp. nov. 
GG No distinct sections of Brent velo 
hairs on the veins, any hairs of such 
colour being isolated and excep- 
tional. Costa with only normally 
black or blackish hairs, with some 
sections of lighter coloured hairs. 
H ‘Tarsi (apart from metatarsi) wholly 
white. 
I Metatarsi wholly white. The 2nd 
longitudinal vein forks proximad of 
origin of 3rd vein . lactevtarsis, Brun. 
ei! Metatarsi black, with white ‘scales at 
tips. The 2nd longitudinal vein 
forks distinctly distad of the origin 
of the 3rd vein .. . gilvtpes, Brun. 
HH Tarsi black: (apical third of fore 
metatarsi) and tips of posterior 
metatarsi with small white scales .. Proxima, sp. nov. 
DD Wings without distinct hair- or scale- 
spots at tips of veins. Hairs on 
veins long enough to overlap those 
of the adjacent veins. No white 
scale-spots on wings. 
J Wings rather smaller than usual. The 
2nd longitudinal vein forks (appar- 
ently) distad of the base of the 3rd 
vettn (54 ee .. impunctata, sp. nov. 
JJ Wings very large. The 2nd _ longi- 
tudinal vein forks immediately 
proximad of the base of the 3rd 
Welt” 2% os oe .. unicolor, sp. nov. 


Pericoma squaminervis, mihi, sp. nov. 


? 2. Darjiling district. Long. 14 mm. 

Body covered with light brown and greyish hair; ground 
colour of thorax light brown, that of abdomen blackish. An- 
tennae as in Psychoda bengalensis. Legs brownish grey, tarsi dis- 
tinctly lighter, almost as pale as in lactettarsis when viewed from 
certain directions. 


304 Records of the Indian Museum. [Viors IV, 


Wings with a depressed row of elongate brown scales on each 
side of each vein, from the base to beyond the middle of the wing 
where they gradually become narrower until eventually, towards 
the margin of the wing, they are replaced by stiff hairs. They lie 
close together, although not always touching one another, and are 
long enough to overlap those emanating from the adjacent veins. 
A distinct spot of bristly black hairs at the tip of each vein, and 
apparently a slightly clearer spot in the wing immediately in front 
of the vein-tip. 

Described from a_ single specimen (apparently a @) in the 
Indian Museum taken by Dr. Annandale at Kurseong, 4-viii-o8. 

N.B.—A very distinct species from all others, readily dis- 
tinguished by the conspicuous, scaled veins. 


Pericoma spinicornis, mihi. 
P. appendiculata, mihi. 


As noted in the original descriptions these two forms re- 
present the sexes of a single species. There can be no reasonable 
doubt about the point, although no actual pair has been found 
in cop. ‘The number of spines on the first flagellar joint in the 
g is six: the palpi consist of four subequal, oblongo-cylindrical 
joints. : 

The Indian Museum possesses a good series giving the follow- 
ing data: Darjiling (7,000 ft.), 26—28-v-1o and 2-x-08, common 
[Brunetti |; 8—-11-viti-0g [ Jenkins]; 6—10-viii-og [Paiva]; Kurseong 
(5,000 ft.), 20--26-vi-Io; 3—5-vil-o8; 4—5-ix-o9 [Annandale]; 
10—26-ix-09 [Lynch]; Kurseong (4,700—5 ,000 ft.), 24-iii-10 [A nnan- 
dale and Gravely]; Siliguri, base of Darjiling hills, 18—20-vii-07 
[Hodgart|; Simla, r1-v-08, 5-v-10, I0-v-0g, on windows [Annan- 
dale|; Phagu (9,000 ft.), 11-v-og [Annandale]; Naini Tal (6,000 
ft.) , 2-vi-o9 [Hodgart]. 


BI 


B] 


Pericoma annandalei, mihi. 


Eight examples of this species have been acquired by the 
Indian Museum, captured by Dr. Annandale at Kurseong, 
22—29-vi-10, where he found them running over Caladium leaves 
at dusk, in thick jungle. 


Pericoma margininotata, mihi. 


P. bella, mihi. 

Long. 13—1? mm. 

From a good series of this species (with which must be in- 
cluded the form described as bella), recently acquired by the 
Indian Museum, it is possible to emend the original description. 
The species is very variable in its coloration. 

The 2nd scapal joint is subglobular, a little more than half as 
iong as the Ist, which is cylindrical, twice as long as its width. 
The flagellum is composed of either fifteen distinct joints, or of 


IgII.] E. Brunetti: New Onental Nemocera. 305 


fourteen joints with an apical style to the last one. The 4th 
palpal joint is the longest and thinnest. 

At the tip of each vein, normally, is a black hair-spot and 
a small white scale-spot, the latter sometimes almost in the 
marginal fringe. The costal fringe comprises some short sections 
composed mainly of white or whitish hairs, including generally a 
section of some length just beyond the middle; whilst the wing- 
tip is always clothed with white hairs for some distance, there 
being also some few short patches of white hair in the fringe of 
the posterior margin. 

An error has crept into the sentence (Rec. Ind. Mus., ii, 383) 
about the 3rd vein, which should read: “‘ the 3rd vein originates 
in a right angle from the 2nd, just beyond where the latter forks ; 
its basal portion very narrow, but quite distinct in wings denuded 
of vestiture ; and there is a distinct appendix at the angle.”’ 

The study of a good series in the Indian Museum makes it 
clear that the black hair-patches on the wing are tolerably con- 
stant, especially those at the fork of the upper branch of the 2nd 
longitudinal vein, and at the fork of the 4th vein. The hairs at 
the wing-tip are generally wholly white, always mainly so. 

The wing as described under the specific name bel/a in my 
previous paper on this family, may be regarded as the typical 
form ; whilst the form described (/oc. cit.) in the subsequent note 
is not at all uncommon. 

The colour of the hair on the dorsum of the thorax varies 
from greyish white to rather bright reddish brown, all interme- 
diate shades being represented in different individuals. 

The tarsi are best described as variable; generally pale 
yellowish at base, darkening to dark brown at the tips; with 
long, irregularly placed bristly hairs, which are pale on the basal 
half of the tarsi and dark brown on the apical half, being concolor- 
ous with the ground colour of the limb. The rest of my descrip- 
tion of the tarsi is correct, and may be supplemented by “* the 
distance covered by the white apical scales at the tips of the basal 
joints of the tarsi, varies, especially on the 2nd tarsal joint, which 
is in some examples all white, the colour in all cases being that of 
the scales, as the ground colour of the whole tarsus is always black.”’ 

The Indian Museum has this species from Darjiling (7,000 ft.), 
26-v-10 [Brunetti]; 11-viii-og [Dr. Jenkins]; 5—8-viii-o9 [ Parva}; 
Kurseong, 25—27-vi-to [Annandale]; Simla, 25-iv-07, II-v-08, 
g—io-v-0g [Annandale]; Phagu (9,000 ft., Simla district), 
II—15-v-09 [Annandale]; Pallode (20 miles N. KE. of Trivandrum, 
South India), 15-xi-o8 [Annandale). 


Pericoma metatarsalis, mihi, sp. nov. 


9. Western Himalayas. Long. 1} mm. 


This species differs from the bella form of margininotaia in 
only two characters, but these are practically consistent in the 
three examples examined. 


306 Records of the Indian Museum. [Vou. IV, 


The metatarsus is, with the exception of its immediate base, 
covered rather thickly with whitish scales (in the type and in one 
other specimen), or at least for more than the apical half (as inthe 
third specimen). 

In margininotata the last three tarsal joints often have a 
greyish or blackish grey appearance, but in the present species 
they are all uniformly intensely black. 

The second character is that the fringe of the wings possesses 
no admixture of whitish hairs either singly or in short sections, 
with the exception of a broad section at the tip, comprised 
between the lower branch of the 2nd longitudinal vein and the 
upper branch of the 4th vein. 

The 2nd longitudinal vein forks a short distance before the 
base of the 3rd vein, at which spot is placed the anterior cross- 
vein; the fork of the anterior branch of the 2nd vein is distinctly 
proximad of the fork of the 4th vein. 

In view of the close affinity of Jlactettarsis and gilvipes to 
margininotata, although they appear to be perfectly good species, 
there seems no reason to refrain from establishing the present 
form as a distinct species. 

Described from three 2 @ from Simla district, two from 
Simla, 9-v-0g (type) and 12-v-o09, the third from Phagu (9,000 ft.), 
II-v-09, all taken by Dr. Annandale. 

Type and the other specimens in the Indian Museum. 


Pericoma mixta, mihi, sp. nov. 


@. Western Himalayas. Long. 2 mm. 


Of the general appearance of that form of margininotata, 
which was described as bella. 

Body with blackish and grey hairs, the former predominating 
chiefly on the dorsum of the thorax and at the base of the wings. 

The wings have a dark brown appearance. The 2nd longitu- 
dinal vein forks some little distance beyond the base of the 3rd 
vein, aithough still quite near the base of the wing, as the latter 
vein begins sooner than in most species, its exact origin near the 
root of the wings being obscured by the pubescence. The fork of 
the upper branch of the 2nd vein and the fork of the 4th vein are 
approximately opposite one another. ‘The veins are closely covered 
with a double row of black or dark brown hairs, with a distinct 
black hair-spot at the tip of each vein; a few small pale hairs in 
front of some of these black hair-spots. Some erect short, snow- 
white hairs towards the bases of the veins, and others on the three 
branches of the 2nd vein, placed some little distance before their 
tips ; also to aless extent in short sections on the veins in other 
parts of the wing. 

A number of bright yellow hairs, distributed (1) along the 
veins, apparently thickest along the costa, especially at its base, 
intermixing with the normal, thick black or dark brown hairs 
forming the fringe, (2) at the bases of the veins, but disposed in 


191 r.] EK. BRUNETTI: New Oriental Nemocera. 307 


small sections separate from the snow-white ones in that region 
of the wing, and (3) on the three branches of the 2nd longitu- 
dinal vein proximad of the white hairs. To an apparently less ex- 
tent they also occur on the veins in other parts of the wing. The 
wing-fringe on the posterior margin is really dark brown, although 
in some lights it appears light brown or even greyish here and there. 
The tip of the wing bears a few whitish hairs but of less white 
colour and to an extent of much less obvious demarcation than 
in the bella form of margininotata. 

Legs dark yellowish grey, thickly clothed with stiff bristly 
hairs; the tibiae, the metatarsi and 2nd joint of tarsi with a 
narrow circlet of whitish scales at the tips of each. 

Described from a single @ in good condition from Simla (7,000 
ft.), taken by Dr. Annandale, 6-v-og. 

Type in Indian Museum. 


Pericoma lacteitarsis, mihi. 


This form, originally described by me asa variety of my 
margintnotata, with the suspicion that it might prove distinct, 
is now, to my thinking, sufficiently established as a good species 
through the acquisition of several specimens by the Indian Museum. 

The black hair-spots at the tips of the veins, the relative posi- 
tions of the forks of the 2nd and 4th longitudinal veins, and the 
coloration of the tarsi are constant. The 2nd longitudinal vein 
forks proximad! of the base of the 3rd vein; the anterior branch 
of the 2nd vein forks before the middle of the wing and a little 
distad of the fork of the 4th vein. 

There are no white hair-spots in the fringe of the wing, nor on 
the absolute margin of the wing (at or very near the tips of the 
veins) as in typical marginenotata, which character alone is almost 
sufficient to distinguish the two forms. The whole wing has a 
more brownish appearance, without the variegated appearance of 
margininotata, and the absence of the white fringe at the wing-tip 
will at once identify it from that species. 

A second good character that I believe separates it from all 
other Oriental species is that of the whole of the tarsi being milk- 
white. In the allied species, margininotata, giluipes, proxima and 
mixta, some portion of the tarsus is always black, often the major 
portion. 

A specimen in the Indian Museum, taken by Dr. Annandale 
at Quilon, Travancore, 9-xi-o8, has the hairs of the body, on 
the tibiae and metatarsi (except the tip of the latter) darker 
brown, also the lower branch of the 2nd longitudinal vein runs 
almost directly to the wing-tip. ‘The blackish hair-spots on the 


| This is the case in the original type specimen (a ?). In a second specimen 
the pubescence obscures a vein of the base of the wing. In a specimen mounted 
for the microscope, the 2nd vein forks distad of the base of the 3rd, and for 
this reason it is doubtful if it is of this species. 


308 Records of the Indian Museum. [Vorrive 


disc of the wing, on the fork of the anterior branch of the 2nd 
vein, and on the fork of the 4th vein, are well marked. 


Pericoma gilvipes mihi. 


This form also, is certainly a good species, distinct from 
margininotata, as a questionable variety of which I first described 
it. ‘The distinguishing character is the absence of white hairs at 
the wing-tip and of white hair-spots on the margin of the wing, 
also of isolated small patches of erect white hairs on the veins. 
Another good character is the distad forking of the 2nd longi- 
tudinal vein in reference to the origin of the 3rd vein, instead of 
the proximad forking as in margininotata. From its nearest ally, 
lacteitarsis, it is known by the nearly wholly black metatarsi, and 
by the distad instead of proximad forking of the 2nd vein, in 
relation to the origin of the 3rd vein. The anterior branch of the 
2nd vein forks a little before the middle of the wing and a little 
distad of the fork of the 4th vein. 

Additional specimens in the Indian Museum afford the 
following data :—Calcutta, 9-ii-10, 17—18-vii-o7, 28-vii-o8, 8— 
23-vili-o8, 1—26-ix-08 [all Annandale, some taken ‘“‘ at light’’]; 
Madhupur, Bengal, 17-x-09 “‘ atlight’’ [Paiva]; Ernakulam, Cochin, 
Malabar Coast, 4-xi-o8 [Annandale]; Quilon, Travancore, 9-xi-08 
[Annandale]. 


Pericoma proxima, mihi, sp. nov. 


> @. Ceylon. Long. 14 mm. 


Body with dark brown hairs, intermixed with grey hairs here 
and there ; bushy white hair on head above, and black hair below 
also on palpi. Abdomen with black hairs. Antennae with black 
scales on scape ; flagellum like Psychoda bengalensis , the verticillate 
hairs widely spread out, the hairs whitish. 

Legs dark, with blackish scales, black bristles and paler stiff 
hairs; fore metatarsi with apical third bearing white scales, 
posterior metatarsi with white scales at tips only. 

Wings with hairs only on the veins, rich, moderately dark 
chestnut-brown. Very distinct and moderately large black hair- 
spots at tips of veins, where there are also some small white scaly 
hair-spots. Isolated groups of a few white scaly hairs placed 
apparently irregularly on the veins. 

A distinct are of white scaly hairs on the wing-fringe between 
the lower branch of the 4th longitudinal vein, and the 5th vein, 
but the wing-tip itself has distinctly rich brown hairs and no trace 
of white hairs. ‘The wing-fringe brownish grey. 

A distinct spot of black bristly erect hairs at the fork of the 
anterior branch of the 2nd longitudinal vein, and another at the 
fork of the 4th vein, both spots with a few white hairs beyond 
them. 

Anterior branch of 2nd vein forking approximately opposite 
the fork of the 4th vein, distinctly but not greatly before the 


IQII.] E. BRUNETTI: New Oriental Nemocera. 309 


middle of the wing. ‘The pubescence hides the base of the wing 
sufficiently for it not to be clearly seen whether the 2nd longitudi- 
nal vein forks before or after the origin of the 3rd. 

The example appears to be a o ; the genitalia are covered 
with bristly bright yellow hair. 

Described from a single specimen in good condition in the 
Indian Museum collection, taken at Peradeniya, Ceylon, viii-1gIo, 
by Mr. Green. 


Pericoma impunctata, mihi, sp. nov. 


Sex ? South India. Long. 15 mm. 


Body thickly clothed with long, very dark brown bristly hair ; 
the surface of the body itself also dark brown. Antennae brownish 
yellow. 

Legs with dark brown bristly hair ; tarsi light brown but 
without traces of any pale scales at tips of joints. 

Wings thickly clothed on all the veins with a double row of 
dark brown bristly hairs, denser and more bristly along the costa 
and at the base of the wing. In certain lights the fringe of the 
wing and some of the stiff hairs on the basal part appear greyish, 
but the true colour of practically every part of the insect is dark 
brown. 

Described from one specimen (sex uncertain) from Tenmalai 
(west side of Western Ghats), Travancore, South India, 22-xi-08, 
captured by Dr. Annandale. 

Type in Indian Museum. 

N.B.—The 2nd longitudinal vein apparently forks beyond the 
base of the 3rd vein but the root of the wing is too closely covered 
with hairs to speak with certainty. 


Pericoma unicolor, mihi, sp. nov. 


Sex? Darjiling district. Long. nearly Lj mm. 


This species is wholly brown in colour, only the tarsi being 
rather lighter. The antennae (partly broken) have flask-shaped 
flagellar joints. The wings are very large, and bear almost wholly 
dark brown hairs which appear rich golden brown in certain lights 
and show a violet tinge when viewed from different directions. 
The 2nd longitudinal vein forks immediately proximad of the 
base of the 3rd vein, the anterior branch of the 2nd forking again 
near the middle of the wing, much distad of the fork of the 4th 
which occurs towards the base, only shortly after the almost 
invisible posterior cross-vein, which latter is situated in a line with 
the basal section of the 3rd vein. The hairs at the tips of some 
of the veins are blacker but do not form distinct spots. 

Described from a single specimen (sex uncertain) in the Indian 
Museum, from Kurseong, taken by Mr. D’Abreu in November, 
IgIo. 

N.B.--Although the actual difference in length between this 
species and impunctata is so little, the present form has a much 


310 Kecords of the Indian Museum. [Vor. IV, 


larger appearance owing to the considerable enlargement of the 
wings, which in zmpunctata are smaller than usual. 


BRUNETTIA, Annand., and PARABRUNETTIA, gen. nov. 


The characters of Brunettia (as defined by its author) which 
appear to me to be most strikingly generic are the closely scaled 
wings and the presence of frominent S-shaped chaetae on the 
flagellar joints. 

To these may be added the lesser ones of broader wings than 
in the other genera in this family, but this character is almost 
certain to be a somewhat variable one. 

The original species was superstes, Annandale, which was at 
first placed by its author in Diplonema, Lw. (an extinct genus), and 
afterwards removed to the present genus, which was established 
by Dr. Annandale for its reception; and to this new genus he 
added a second species from South India, travancorica. 

As superstes was placed first (of the two species) in the new 
genus, it must be regarded as the generic type, as indeed, was 
intended and expressed by its author. Moreover, as superstes has 
a venation identical with that of Pericoma,! that type of venation 
must stand good in all species of Brunettia. 

B. travancorica, Annand. (which was afterwards recognised as 
synonymous with my Psychoda atrisquamis), has, however, 
Psychoda-like venation, and therefore this species cannot remain 
in Brunettia, for it would be illogical to allow the two types of 
wing in a single genus, when Psychoda and Pericoma themselves 
are mainly differentiated by this difference in the venation. 

It therefore appears that the genus Dr. Annandale has flatter- 
ingly named after me, still contains but the original species 
superstes, the consequence being that a new genus must be formed 
for all those other species of the present group that possess 
Psychoda-like venation. 

This new genus it is proposed to call Parabrunettia, and it 
differs from Brunettia primarily in the different venation. 

Its other characters may be regarded as— 

(2) the presence of closely placed dark imbricating scales on at 
least some considerable portion of the wings, on both their upper 
and lower surfaces, or on the under side alone?; (3) the surface 
of the wing generally rather thickly covered to some considerable 
extent with more or less longitudinally placed hairs’; (4) chaetae 
present on the flagellar joints (possibly not on all of them, 
irrespective of the apical joint, which, even in B. superstes, is 
devoid of them). They are not so large nor so conspicuous in any 


| That is to say, the 3rd longitudinal vein ends a little below the wing tip, 
instead of at the tip. : 

2 Any species (if discovered), with scales on the upper side of the wing only, 
would logically fall in the new genus. : 

8 These are absent in two species provisionally placed here, but this may be 
better regarded perhaps as a subgeneric character. as species both with and 
without hairs on the surface of the wings occur equally in Psychoda and Pericoma. 


IQII.]| E. BRUNETTI: New Oriental Nemocera. SLE 


of the species as they are in swperstes, and appear to vary a good 
deal in size, according to the species.! 

In many species there is a rather noticeable patch of smooth 
depressed silky long hairs extended posteriorly from the alulae, 
and which may probably figure as a secondary character of the 
new genus. 

Both Brunettia and Parabrunettia belong distinctly to the 
Psychodinae subfamily of Psychodidae, the 7th longitudinal vein 
being conspicuously present, and the 2nd longitudinal vein forking 
quite near the base of the wing. 


PARABRUNETTIA, mihi, gen. nov. 


To this new genus must be admitted the three species placed 
by me in Psychoda but divided off from the others by the presence 
of large scale-covered areas in the wings: squanupennis, atrisqua- 
mts and argenteopunctata. 

Brunettia travancorica, Annand., has been shown to be 
synonymous with my airisquamis. 

To these three species are now added four new ones, 
albohumeralis, 9-notata, flavicollis and longichaeta. 

Owing to the denseness of the vestiture of the wings (the 
basal hairs, the surface hairs and the opacity of the scales), it has 
been impossible to note the exact position of the forking of the 
2nd longitudinal vein in some of the species, but in all those in 
which it has been noted, it occurs distad of the origin of the 3rd 
longitudinal vein. 


PARABRUNETTIA. 
Table of spectes. 


A Hairs thickly present on swv/ace of wing. 
B Upper surface of wing as well as lower 
surface covered to a considerable 
extent with small dark imbricating 
scales. 
C No white spots on wing-border. (The 
2nd longitudinal vein forks distad 
of origin of 3rd vein.) .. squamipennts, Brun. 
CC White spots on wing-border .. atrisquamis, Brun. 
BB Upper surface of wing without any con- 
siderable area covered with scales 
(a few may be present at the base 
of the wing). 
D Under surface of wing wholly covered 
with dark scales. 


1 Similar chaetae but much smaller, have been detected by Dr. Annandale in 
Pevicoma margininotata and Psychoda ‘distincta, so they cannot be considered of 
generic importance. 


312 Records of the Indian Museum. [VoL. IV 


E A _ silvery white scale-spot below 
shoulder ny, ate .. albohumeralis, sp.nov. 
EE Such silvery white spot absent .. argenteopunctata , 
Brun. 
DD Under surface of wings with scales 
covering at most the basal half. 
(The 2nd longitudinal vein forks 
apparently distad of the base of the 
ard vein.) ih “ . g-notata, sp. nov. 
AA Hairs wholly absent from the surface 
of the wing. (The 2nd longitudinal 
vein forks distad of the origin of the 
3rd vein.) 
F eee flavicollis, sp. nov. 
FF Bene longichaeta, sp. nov. 


Parabrunettia atrisquamis, mihi. 


Psychoda id., mihi. 
Brunettia travancorica, Annand. 

The brown or blackish short and moderately short scales on the 
upper surface of the wing are thickly present on the upper 
as well as the lower surface, but on the basal half only. They 
appear silky black when viewed in some directions and dull 
greyish white viewed from other directions. On the lower surface 
these dark scales cover the whole area. 

The very small white hair-spots at the tips of the veins are 
placed on the absolute edge of the wing, almost in the adjacent 
fringe. 

Some specimens taken by Mr. E. E. Green at Peradeniya, 
Ceylon, afford the data 18-vii, 14-vii1 and October (all 1910). 

N.B.—A specimen in the Indian Museum, unfortunately in too 
bad condition to describe, certainly represents an undescribed 
species allied to atrisquamis, from which it differs by the scales on 
the under surface of the wings covering the basal half only. No 
obvious patch of silky hairs on the alulae. Traces of small white 
spots on the extreme border of the wing. 


Parabrunettia albohumeralis, mihi, sp. nov. 


@. Ceylon. Long. 14-14 mm. 

Body covered with dark blackish brown hair, which, at least 
on the abdomen, appears dark greyish in certain lights. A small 
but very conspicuous tuft of snow-white scaly hairs at the sides of 
the thorax, nearly below the shoulders, a little in front of the base 
of the wing. 

Antennae with general appearance of those of Psychoda 
bengalensis. egs with brown hairs and scales, which are darker 
on the tarsi. Tips of tibiae and of metatarsi with white scales. 


IgII.| E. Brunetri1: New Oriental Nemocera. 313 


Wings with surface closely covered with dark brown hairs ; 
the upper surface without any covering of scales except a very 
few at the base and the small ones forming the white spots. 
Under side of wing closely covered with dark brown imbricating 
small scales, extending almost to the margin. Conspicuous though 
small spots, composed of elongate snow-white scales, appear to be 
placed normally at the tips of nearly all the veins, but in the 
three examples present there is a little variability in their exact 
position. In the type they occur as follows: at a little distance 
before tip of Ist longitudinal vein; shortly before the Ist ending 
of the 2nd vein; at tips of the other two endings; at tips of 3rd 
vein, both branches of 4th, the 5th and 7th. A single white hair 
still remaining shows another spot to be present on the 7th vein 
some little distance before its tip. Tip of 6th vein without a spot. 
In one example there is a white spot behind the tip of the 6th 
vein, in another specimen the spot is at the vein tip. Venation 
Psychoda-like. 

Described from three specimens, females, from Peradeniya, 
taken by Mr. E. E. Green, ix- and x-1910. 

Type in Indian Museum. 

N.B.—The white shoulder-spots immediately distinguish this 
from all other species. 


Parabrunettia g-notata, mihi, sp. nov. 


9. Orissa Coast. Long. 14 mm. 


Body covered with rich dark brown bristly hair on thorax, 
blackish brown on abdomen, with a few white hairs at the abdo- 
men tip. 

Antennal scape with dark elongate scales; flagellum with 
pear-shaped joints and brown verticillate hairs, which appear 
greyish when viewed in certain directions. Distinct long, curved 
chaetae present. 

Legs blackish ; femora with a considerable number of greyish, 
elongate, depressed scales ; tibiae and metatarsi with whitish grey 
scales at tips; a few scales of a similar nature are also present at 
the tips of the tarsi. 

Wings with the whole surface as well as the veins covered 
with blackish hair, which is much thicker and denser on the 
costa, where it is distinctly prominent and its colour distinctly 
black. Around the remainder of the border of the wing it is dark 
brown. A very conspicuous spot at the extreme edge of the wing- 
border, at the tips of every vein, composed of small white scale- 
like hairs. A number of white erect hairs on most of the veins in 
the basal half of the wing Anterior branch of 2nd longitudinal 
vein forks opposite the fork of the 4th vein. The pubescence pre- 
vents a clear view of the base of the wings, but apparently the 
2nd longitudinal vein forks distad of the base of the 3rd vein. 
Venation Psychoda-like. 


314 Records of the Indian Museum. PVon. ive 


Described from one @ from Puri, Orissa, east coast of India, 
12-xi-Io, taken by Dr. Annandale on a window pane; the speci- 
men is now in the Indian Museum. 

N.B.—The conspicuous, curved chaetae on the flagellum 
suggest that this species belongs here, and the hairy surface of the 
wing is a second character of the genus. However, the wing is des- 
titute of scales, except the small ones forming the spots, and the 
species is placed here provisionally, pending the satisfactory 
elucidation of this group of forms. 


Parabrunettia flavicollis, mihi, sp. nov. 


a7. Ceylon. Long. about 14 mm. 


Vertex with bushy yellow bristly hair; frons and face with 
black bristly hair. Antennae with flask-shaped flagellar joints, 
long necked, the brown verticillate hairs widely spread: long 
S-shaped chaetae distinctly present: scapal joints with scales. 

Thorax with rather bright yellowish hair, with some grey hair 
intermixed. Abdomen with moderately dark brown hair. Genital 
organs with close long bristly hair, apparently normal in !orm. 

Legs covered with dark browm scales; some yellowish white 
bristly hairs on tibiae, longest on hind pair, which latter are 
thickened at their tips and bear a circlet there of yellowish grey 
scales. Tips of anterior tibiae and tips of all the metatarsi with 
natrow circlets of yellowish white scales. 

Wings with the 2nd longitudinal vein forking considerably 
distad of the base of the 3rd. Anterior branch of 2nd vein forking 
distinctly before the middle of the wing and a little beyond the 
fork of the 4th vein. Hairs placed only on the veins, not on the 
surface of the wings. A black hair-spot at tips of the veins and a 
few stiff yellowish grey hairs at or just before the tips of the veins. 
A black bristly hair-spot at fork of anterior branch of 2nd longi- 
tudinal vein. Venation Psychoda-like. 

Described from a single ~ in the Indian Museum collection 
from Peradeniya, Ceylon, taken August, 1910. 


Parabrunettia longichaeta, mihi, sp. nov. 


¢. South India. Long. 14 mm. 


Thorax with dark ground colour and yellowish bristly hair. 
Abdomen with dark ground colour with light brownish grey hair. 
Hair on head brown Palpi dark brown. Antennae generally as in 
Psychoda bengalensts; the verticils of hair widely spread out, light 
brown: the chaetae very long and conspicuous, although pale, 
S-shaped. 

Legs light brown, with concolorous scales and bristles, also 
whitish scales on knees and on base of tibiae, on tips of tibiae and 
of metatarsi, and very minute yellowish white scales on the tips of 
the remaining joints. Femora and tibiae with short stiff black 
hairs here and there. 


Ig1I.] EK. BRUNETTI: New Oriental Nemocera. 315 

Genitalia large and distinct, very pubescent. 

Wings (rather rubbed) with hairs on surface only; light brown 
or greyish, with small patches of light erect hairs irregularly dis- 
posed. The 2nd longitudinal vein forks distinctly distad of the 
base of the 3rd vein; the anterior branch forking a little beyond 
the fork of the 4th vein, both very near the middle of the wing. 
Venation Psychoda-like. 

Described from a single * in the Indian Museum, taken by 
Dr. Annandale at Maddathorai, Travancore State, South India, 
17-x1-08. 

N.B.—Reterred doubtfully and provisionally to the present 
genus. There being no hairs on the surface of the wings and 
no scales, it is questionable whether a new genus should not be 
erected for it. The species, however, is quite a good one. 


BLEPHAROCERIDAE. 
Apistomyia trilineata, mihi, sp. nov. 


@. Darjiling district. Long. 4 mm. 


Head.—Eyes closely, microscopically pubescent, upper facets 
very distinctly larger than lower ones, divided by a distinct 
natrow space, the upper ones being about one-fourth the total 
height of the eyes. Frons one-third the width of the head, bare, 
the eye-orbits narrowly silvery ; ocellar triangle distinct, elevated 
the three ocelli conspicuous, reddish brown. Face whitish grey, 
with silvery reflections. Antennae black, bare; Ist scapal joint 
much broader at tip than at base, 2nd joint twice as long as the 
Ist, much broadened at tip, where it is produced on the under 
side into an elongate blunt point. Flagellum of 8 joints, the Ist 
barely half as long as the 2nd scapal joint, the following joints of 
thickened bead shape, the apical joint ovate. 

The proboscis consists of a long elongo-conical rostrum, 
moderately stout at the base, whitish in colour, with blackish 
dorsum, thence produced into a tapering, very sharply pointed 
brownish yellow horny piece quite separate from the proboscis 
proper, which is very long, black, narrowly cylindrical, genicu- 
lated before the middle, beyond which it is bilobed, the ends 
curled up. 

Thorax ash-grey, the central part of the dorsum up to 
beyond the middle occupied by three broad velvet-black stripes 
almost contiguous, extending laterally almost to the sides; the 
median stripe produced forwards to the anterior margin, which 
is wholly black to the shoulders, leaving a narrow grey space in 
front. Pleurae, scutellum and metanotum blackish. 

Abdomen velvet-black, the anterior corners of most of the 
segments a little reddish brown with more or less silvery reflec 
tions. Belly reddish brown. Genitalia moderately small, oval, 
blackish, not conspicuous. 


316 Records of the Indian Museum. [Vot. IV, rg1t.] 


Legs.—Coxae wholly and the femora more or less at the base, 
brownish yellow: remainder of anterior legs blackish; tibiae and 
tarsi of hind legs brownish yellow, extreme tips of joints black. 
Anterior femora distinctly clubbed at the tips, the hind femora 
moderately thickened towards tips. The hind femora and tibiae 
are each twice as long as the anterior ones, and the hind tarsi are 
fully as long as the hind tibiae. 

Wings absolutely clear, costa very narrowly black; a 
small brownish infuscation at extreme tip of wing. Venation 
normal, agreeing with Kellogg’s figure.| Halteres black, stems 
yellowish. 

Described {from a single specimen in the Indian Museum 
taken by Dr. Annandale at Kurseong (5,000 ft.), 4-ix-09. 


Blepharocera indica, mihi, sp. nov. 


@” 9. ° Western Himalayas. Long. 4—5 mm. 


Head.—Frons narrow,” dark grey or blackish; ocelli large and 
conspicuous. -Face whitish. Proboscis brownish yellow, blackish 
at the base on upper side ; elongate, pointed, about as long as the 
height of the head. Palpi elongate, four joints of about equal 
length, pale brownish yellow, with some stiff black hairs. 

Thorax.—Dorsum blackish. Sides, scutellum and metanotum 
brownish yellow. A very small yellowish mark behind each 
shoulder, 

Abdomen blackish; the base of each segment very narrowly 
yellowish white. Genitalia inconspicuous. 

Legs brownish or brownish yellow: the hind femora slightly 
thickened towards the tips, about one-and-a-half times as long 
as the anterior femora. 

Wings very pale yellowish grey ; unmarked; venation nor- 
mal. Halteres: stem yellowish, club black. 

Described from two 7 @ anda single 2 in the Indian Mu- 
seum from Phagu (9,000 ft.), Simla district, 12—15-v-09, taken 
by Dr. Annandale. 

N.B.—Both sexes are presumed to be present from the 
appearance of the abdomens, which in the two examples I consi- 
der males is blunted, with an exceedingly small projecting piece ; 
and in the supposed female is widened before the tip, with a 
short pointed ovipositor-like termination. 


1 Plate ii, fig. 20, in Gen. Ins., Fasc. 56. 

2 The headin each of the examples is either damaged or shrunken in drying, 
so that the proportionate width of the frons is not easily gauged. It is appar- 
ently quite narrow. 


MISCELLANEA. 


SYNONYMY IN CORETHRINAE.—Many recent authors have 
followed Theobald (Gen. Ins., Fasc. 26) (1905) in referring to 
Sayomyia, Coq., most of the species till recently incorporated 
under Corethra, Mg., but the American author’s genus must give 
way, on his own admission,! to Chaoborus, Lichtenstein, estab- 
lished as long ago as 1800; so that a few synonymical notes may 
be useful. 

Corethra, Mg. (1803), was originally erected (Illig. Mag., i, 
260) for the Tipula culiciformis of De Geer (1776); and for many 
years it contained only that species, with pallida, F. (1781), and 
plumicornis, F. (1794). It was not until 1823 that another species 
was added ,—punctipennis, Say., followed by flavicans, Mg., in 1830 
and others of more recent date. 

Mochlonyx, Loew (1844), was formed for Corethra velutina, 
Ruthe, and to this genus has been added effoetus, Wlk., and 
actually culiciformis, De Geer (!), the very type species of Corethra; 
even Prof. Kertesz, in his catalogue of the world’s diptera (vol. 1, 
1902), making the same error. 

Theobald (Gen. Ins.) admitted Corethrinae as a subfamily of 
Culicidae and correctly replaced culiciformis in Corethra, with 
velutina (effoetus, Wik., being added as synonymous with this) ” 
and a third, recently described species, cincttpes, Coq. Therefore, 
unless velutina can be generically separated from culicifornis (and 
I have no means of forming an opinion on this), Mochlonyx must 
sink in favour of Corethra, Mg. 

Schiner (Fauna Austr., ii, 624) placed velutina and effoetus (the 
latter not described, not being Austrian) in Mochlonyx ; and placed 
culiciforymis (also undescribed for the same reason), with the names 
of five other undescribed non-Austrian species, after his description 
of the three Austrian species (pallida, plumicornis and fusca) of 
Corethra. It is quite possible that he may not have seen all these 
species, and therefore his generic separation of velutina and culict- 
formis may not have represented an individual opinion. 

Chaoborus, Lichtenstein, was erected for ‘‘ antisepticus sp. 
nov.,’’ which proved synonymous with crystallina, De Geer, this 
latter, queried by Theobald (Gen. Ins.) as synonymous with plum- 
cornis, F., being definitely given by Kertesz as identical with it. 


1 Proc. U. S. Nat. Mus., vol. 37, p. 603 (1910). 
2 As Mr. Theobald has presumably examined the type of Walker’s species, 
some Importance attaches to this opinion. 


318 Records of the Indian Museum. [VOE EVs 


Now p/lumicornts, F., pallida, F., and punctipennis, Say. (a 
North American species), with others, are retained by Kertesz 
under Corethra. This-is obviously wrong, since Loew’s principle 
in establishing Mochlonyx was a sound one, ?.e., ‘‘ metatarsus 
several times shorter than the next tarsal joint,’’ as contrasted 
with those species in which the metatarsus is longer than the 2nd 
tarsal joint.! The separation of these groups is justified : Loew 
lost his genus through unfortunately giving the name Mochlonyx to 
that group of species containing the type species of Corethra, Mg. 
Of course, in the days in which he wrote it was sometimes not 
easy to distinguish which species was intended by an author as 
the type of his genus* and this may have authorised him to split 
off any group, or particular species at will. 

Corethra, Mg., must therefore always stand for culiciformis, 
with its congeners. 

Coquillett in 1903 erected Sayomyia for ‘‘ Corethra puncti- 
pennis’’ Say., which he admitted as congeneric with plumicornis, 
F. (the latter species possessing crystallina, De G., and others as 
synonyms), both of which species have now to be placed in 
Chaoborus. 

The synonymy of the species immediately concerned wili 
stand thus :— 


Corethrinae (Subfamily of Culicidae.) 
CoRETHRA, Mg., 1803. 
(Mochlonyx, Loew, 1844.) 


1. culiciformis, De Geer (Tipula id.). TYPE of genus. 
2. velutina, Ruthe (Mochlonyx id., Loew ; TYPE of Mochlonyx) 
(effoctus W1k.). 


3. cinctipes, Coq. 
CHAOBORUS, Lichtenstein, 1800. 
(Savomyia, Coq., 1903.) 
1. plumicornis, F. (Ttpula id.). Type of genus. 


2. punctipennis, Say. (TYPE of Sayomyta). 
3. All other species referred to Sayomyza since Theobald’s ac- 
ceptance of the genus, but previously placed in Corethra. 


The three other genera recorded by Theobald appear to me 
built on very weak characters and I should prefer to regard them 
as subgenera of Chaoborus only. Each contains but one species. 
They are Pelorempis (Joh.) americana, Joh., Corethrella (Coq.) 
brakeleyt, Coq., and Eucorethra (Underwood) underwoodi, Underw. 


E. BRUNETTI. 


1 Theobald (Gen. Ins.) is distinctly wrong in terming the joint following the 
metatarsus as the first, since the metataysus itself is the ist taysal joint, the suc- 
ceeding joint being the second. 

2 T have seen it stated somewhere that Meigen placed what he considered all 
the most typical species in the middle of the genus, and those tending to aberration 
at one end or the other of it. 


1911. | Miscellanea. 319 


FuRTHER Notes on INDIAN Philebotomt.—The recent acquisi- 
tion of a number of specimens of Phlebotomus from different parts 
of India and especially from Ceylon enables me to amplify the 
notes published in these ‘‘ Records’’ last year (vol. iv, No. ii, 
pp- 35——52), andI have been helped greatly to gain a true apprecia- 
tion of certain characters by frequent discussions with Mr. E. 
Brunetti. 

Hitherto I have laid great stress on the relative lengths of the 
different parts of the second longitudinal vein as a specific charac- 
ter, but these lengths are evidently more variable than I thought. 
They would provide a method of separating a species such as 
P. himalayensis from one such as P. argentipes ata glance, but in 
the latter the length of the upper branch of the vein, and conse- 
quently that of the other parts also, varies within fairly wide limits. 
I have had, therefore, to recognize that the form recently described 
by me as P. marginatus is not a ‘‘ good’’ species but merely a 
colour variety of P. argentipes. 

Regarding the sanitary importance of Phlebotomus attention 
may be directed to a paper just published by Major F. Wall, 
I.M.S.! The facts cited suggest (1) that troops moved into certain 
barracks in Chitral become infected with a fever akin to or 
identical with what is called Papatacifieber in Austria, these 
barracks being frequented by P. papatasi and P. babu, and (2) 
that men from districts in which these two species occur are less 
liable to suffer from this fever (having perhaps become to some 
extent immune) than those from districts in which the common 
species are P. major and P. limalayensis. 


Phlebotomus argentipes, Annandale and Brunetti. 


Rec. Ind. Mus., iv, p. 44, pl. iv, fig. 3, pl. vi, fig. 6; Spolia Zey- 

lanica, vii, p. 59. 

I have recently examined a good many specimens from Pera- 
deniya, which is situated in the interior of Ceylon at an altitude of 
about 1,500 feet. The species occurs all over the plains of India 
except in the extreme north-west. Apparently it does not occur 
in the Himalayas. 


Var. marginatus, Annandale. 
P. marginatus, Spolia Zeylanica, vii, p. 62, fig. 7. 

This form cannot be regarded as morethan a colour variety, 
distinguished from the typical form of P. argentipes by the fact 
that the dorsum of the thorax is brown instead of black and the 
sides of the thorax rather darker than in the typical form. 


The var. marginatus occurs in Calcutta as well as at Pera- 
deniya and is apparently not a seasonal form. 


1 Ind. Med. Gazette, xlvi, p. 41 (TOIT). 


320 Records of the Indian Museum. [Vou IV, tort.] 


Phlebotomus major, Annandale. 


Rec. Ind. Mus., iv, p. 46, pl. v, fig. 4, pl. vi, fig. 4. 

Specimens appear to be invariably larger than those of 
P. argentipes, from which both the typical form and the variety 
here described may be distinguished by the fact that the dorsum of 
the thorax is never black or brown. 


Var. grisea, nov. 


Distinguished from the typical form by the general greyish or 
brownish (instead of golden) colour. 

I took several specimens in a house at Kurseong in the Dar- 
jiling district (alt. 4,700 feet) in June, rg10, and also saw the 
variety in the same house in April, 1911. The two varieties are 
easily distinguished by the naked eye. 


Phlebotomus babu, Annandale. 


Rec. Ind. Mus., iv, p. 49, pl. iv, fig. 1, pl. vi, figs. 3, 3a; Spolia 

Zeylanica, p. OI. 

As I have pointed out in a recent paper, this species is prob- 
ably identical with ‘‘ Hebotomus’’ minutus, Rondani, but it is 
impossible to state that it is synonymous without a comparison of 
specimens. Ihave recently obtained specimens of P. babu from 
Peradeniya in Ceylon and from Drosh in Chitral in the Hindu- 
Kush Mountains (4,700 ft.). The species is probably distributed 
all over the plains of India and ascends the Western Ghats to an 
altitude of at least 2,000 feet, but is not known from the Hima- 
layas. 

Var. miger, nov. 


Darker than the typical form and as a rule larger. 

This form was at first regarded as a distinct species and is 
marked as probably being so in Mr. F. M. Howlett’s collection. 
I cannot, however, distinguish any constant difference in its vena- 
tion or genitalia. 

As yet the variety has only been taken in Bihar. I have not 
seen it in Calcutta. 


Phlebotomus papatast (Scopoli). 


Rec. Ind. Mus., iv, p. 51, pl. iv, fig. 4, pl. vi, fig. 2. 
I have recently received specimens from Drosh, Chitral (Major 
F. Wall, I.M.S.), and from Quetta, Baluchistan (Lt.-Col. Wimber- 
ley, I.M.S.). The species appears to be common, together with 
P. babu, all over the north-west of India, ranging as far east as 
Pusa in Bihar. 
N. ANNANDALE. 


Part V.—Revision of the Oriental Leptide.¥% Revised and annotated Catalogue of 
Oriental Bombylide, with descriptions of new species. 


Vol. III, 1909. 
Part I.—The Races of Indian Rats. 


Part II.—Notes on Freshwater Sponges, X. Report on a collection of aquatic 
animals made in Tibet by Capt. F. H. Stewatt in 1907, IJ. Note on some 
amphibious Cockroaches. Description de quelques nouvelles Cécidomyies des 
Indes. Description of new land and marine shells from Ceylon and S. India. 
Description of two new species of Cavanx from the Bay of Bengal. Remarks on 
some little known Indian Ophidia. Remarks on some forms of Dipsadomorphus. 
A pelagic Sea-Anemone without tentacles. Rhynchota Malayana, II. 


Part III.—Notes on the Neuroptera in the collection of the Indian Museum. New 
Indian Leptide and Bombylide, with a note on Comastes, Os. Sac., v. Hetero- 
stylum, Macq. Notes on the Trichoptera in the collection of the Indian Museum. 
Diagnoses of new species and varieties of Freshwater Crabs, 1—3. Report on a 
small collection of Lizards from Travancore. Descriptions of three new Cicinde- 
line from Borneo. The relation between fertility and normality in Rats. Des- 
cription of a Barnacle of the genus Scalpellum from-Malaysia. The Hemipterous 
family Polyctenide. Notes on Freshwater Sponges, XI. Descriptions of two 
new shells from S. India. Preliminary note on a new genus of Phylactolezematous 
Polyzoa. Miscellanea, 


Part IV.—Description of a minute Hymenopterous insect from Calcutta. The Insect 
Fauna of Tirhut, No. 1. Descriptions of new species of Botia and Nemachilus. 
New Oriental Sepsinz. A new species of /vedericella from Indian lakes. Diagnoses 
of new species and varieties of freshwater crabs, No. 4. On some new or little 
known Mygalomorph spiders from the Oriental region and Australasia. 


Vol. IV, 1910-1911. 


No. I.—Second report on the collection of Culicide in the Indian Museum, with des- 
criptions of new genera and species. 

Nos. II and III.—The Indian species of Papataci Fly (Philebotomus). Taxonomic 
values in Culicide. 

No. IV.—Revision of the Oriental blood-sucking Muscide (Stomoxine, Philema- 
tomyta, Aust., and Pristivhynchomyia, gen. nov.). 


No. V.—A new arrangement of the Indian Anopheline. 


No. VI.—A revision of the species of Tabanus from the Oriental Region, including 
notes on species from surrounding countries. 


Vol. V, 1910. 


Part I.—The Hydroids of the Indian Museum, I. Notes on Freshwater Sponges, XII. 
Descriptions of new Shells in the collection of the Indian Museum from Burma, 
Siam and the Bay of Bengal. Materials for a revision of the Phylactolamatous 
Polyzoa of India. Studies on the aquatic Oligocheta of the Punjab. An undes- 
cribed Burmese Frog allied to Rana tigrina. Miscellanea. 


Part I7,—Description d’Ophiures nouvelles provenant des derniéres campagnes de 
‘‘V Investigator ’’ dans l’Océan Indien. Description d’Holothuries nouvelles 
appartenant au Musée Indien. ‘The races of Indian rats, II. Description of a new 
species of Sca/pellum from the Andaman sea. Descriptions of five new species 
of marine shells from the Bay of Bengal. Notes on fish from India and Persia, 
with descriptions of new species. 


Part III.—A new genus of Psychodid Diptera from the Himalayas and Travancore. 
The Indian barnacles of the subgenus Smiliuwm, with remarks on the classification 
of the genus Scalpellum. On a sub-species of Scutigerella unguiculata, Hansen, 
found in Calcutta. The distribution of the Oriental Scolopendride. Notes on 
Decapoda in the Indian Museum, I. Description of a new species of Nemach*lus 
from Northern India. Notes on the larve of Toxorhynchites immisericors, Wk 
Description of a South Indian frog allied to Rana corrugata of Ceylon. Contri- 
butions to the fauna of Yunnan, Introduction and Part I. Miscellanea. 


Part IV.—Notes and descriptions of Indian Microlepidoptera. On some aquatic 
oligochaete worms commensal in Spongilla cartert. On Bothrioneurum tris, 
Beddard. Notes on nudibranchs from the Indian Museum. On the classification 
of the Potamonidae (Telphusidae). Catalogue of the pheasants, peafowl, 
jungle fowl and spur fowl in the Indian Museum, On certain species of Palaemon 
from South India. Alluaudella himalayensis, a new species of degenerate (¢") 
cockroach, with an account of the venation found in the genera Cardax and 

; Alluaudella. Rhynchota Malayana, III. 


Voie Vion: 


Part I.—Note on a Rhizocephalous Crustacean from fresh water and on some speci- 


Pavt II.—Some Sponges associated with gregarious Molluscs of the family Vermetidae. . 


No. 


mens of the order from Indian seas. Notes on Decapoda in the Indian Museum, 
TI. Contributions to the fauna of Yunnan, Parts. II to V. Notes on Pedipalpi 
in the collection of the Indian Museum, I and II. Descriptiors of six new 
species of shells from Bengal and Madras. Miscellanea. 


Report on a collection of aquatic animals made in Tibet by Captain F. H. Stewart 
in 1907, III. Notes on Cyprinidae from Tibet and the Chumbi Valley, with a des- 
cription of a new species of Gymnocypris. Preliminary descriptions of new species 
and varieties of Crustacea Stomatopoda in the Indian Museum. Notes on the 
development of some Indian Ascalaphidae and Myrmeleonidae. Miscellanea. 


MEMOIRS 
of the 


INDIAN MUSEUM 


Vol. I. 


. I.—An account of the Rats of Calcutta. By W.C. Hossack. Rs. 5-8. 
. 2.—An account of the Internal Anatomy of Bathynomus giganteus. By R. E. 


TOM anG we 


. 3 A and B.—The Oligocheta of India, Nepal, Ceylon, Burma and the Andaman 


Islands, with an account of the anatomy of certain aquatic forms. By W. 
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. 4.—Investigatoy sicarius, a Gephyrean Worm hitherto undescribed, the type of a 


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. 1.—Report on the Fishes taken by the Bengal Fisheries Steamer ‘‘ Golden Crown.’’ 


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. 3.—A description of the deep-sea fish caught by the R.I.M.S. Ship ‘‘ Investi- 


? 


gator ’’ since the year 1900, with supposed evidence of mutation in Malthopsis, 
and Illustrations of the Zoology of the R.I.M.S. Ship ‘‘ Investigator,’’ Fishes, 
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; 4.—Etude sur les Chironomides des Indes Orientales, avec description de quelqyes 


nouvelles espéces d’Egypte. Par J.J. KIEFFER. Rs. 2. 


Vol. III. 


. I.—Report on the Fishes taken by the Bengal Fisheries Steamer ‘‘ Golden 


Crown.’’ Part II.—Additional notes on the Batoidel. By N. ANNANDALE. Part 
III.—Plectognathi and Pediculati. By N. ANNANDALE and J. T. JENKINS. 
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2.—Studies in post-larval development and minute anatomy in the genera 
Scalpellum and Ibla. By F. H. STEWART. Rs. 4. 


Other Publications edited and sold by the Superintendent of the Indian 
Museum (also obtainable from Messrs. lriedlander & Sohn) issued 
by the Director of the Royal Indian Marine. 


. Illustrations of the Zoology of the R.I.M.S. ‘‘ Investigator ’’ 1892. Fishes, Plates I to VII. 
Crustacea, Plates I to V, 1894. Fishes, Plates VII to XIII. Crustacea, Plates VI to VIII. Ech- 
inoderma, Plates I to III, 1895. Echinoderma, Plates IV and V. Fishes, Plates XIV to XVI. 
Crustacea, Plates IX to XV, 1896. Crustacea, Plates XVI to XXVII, 1897. Fishes, Plate XVII. 
Crustacea, Plates XXVIII to XXXII. Mollusca, Plates I to VI, 1898. Fishes, Plates XVIII to 
XXIV. Crustacea, Plates XXXIII to XXXV. Mollusca, Plates VII and VIII, 1899. Fishes, 
Plates XXV and XXVI. Crustacea, Plates XXXVI to XLV, 1900. Fishes, Plates XXVII to 
XXXV. Crustacea, Plates XLVI to XLVIII. Index, Part I, 1901. Crustacea, Plates XLIX 
to LV. Mollusca, Plates IX to XIII, 1902. Crustacea, Plates LVI to LXVII. Crustacea, Plates 
_LXVIII to LXXVI._ Fishes, Plates XXXVI to XXXVIII, 1905. Crustacea (Malacostraca), Plates 
LXXVII to LXXIX. Crustacea (Entomostraca), Plates I and II. Mollusca, Plates XIV to 
XVIII, 1907. Fishes, Plates XXXIX to XLIII. Crustacea (Entomostraca), Plates III to V. 
Mollusca, Plates XIX and XX, 1908.—Re. 1 per plate. Mollusca, Plates XX1 to XXIII, 1909.— 
As. 8 per plate. — i : 


RECORDS 
of the 


INDIAN MUSEUM 


Vol. I, 1907. 


Part I.—Coutributions to the Fauna of the Arabian Sea. Hemiptera and Hymenop- 


tera from the Himalayas. Indian Freshwater Entomostraca. The Fauna of 
Brackish Ponds at Port Canning, Lower Bengal, I—III. A Sporozoon from the 
heart of a Cow. Miscellanea :—The appendicular skeleton of the Dugong. An 
egg laid in captivity by a Goshawk. Melanic specimens of Barbus ticto. Two 
barnacles new to Indian seas. Mosquitoes of the genus Anopheles from Port 
Canning. Anopheles larvae in brackish water. Mosquitoes from Kumaon. 
Peculiar habit of an earthworm. 


Part II.—Revision of the Oriental Stratiomyidae. “An Oligochaete Worm allied to 


Chaetogaster. The Fauna of Brackish Ponds at Port Canning, Lower Bengal, IV. 
A Polyzoon from the Himalayas. Batrachia, Reptiles and Fish from Nepal and 
the Western Himalayas. The Fauna of Brackish Ponds at Port Canning, Lower 
Bengal, V. Oriental Diptera, I and II. Miscellanea :—Gecko verticillatus in 
Calcutta. The distribution of Kachuga sylhetensis. The distribution of Bufo 
andevsont. Note on Rutilia nitens. Some Indian Cerambycidae. Some Indian 
Hemiptera, A preoccupied specific name in Macrothyix. An enemy of certain 
Pearl Oysters in the Persian Gulf. The distribution in India of the African snail, 
Achatina fulica. Statoblasts from the surface of a Himalayan pond. Notes on 
Hislopia lacustris. 


Pavit III.—Marine Polyzoa in the Indian Museum. The Fauna of Brackish Ponds at 


Port Canning, Lower Bengal, VI. Earwigs (Dermaptera) in the Indian Museum. 
Oriental Diptera, III. A new snake from Nepal. Marketable fish from Akyab. 
Freshwater Oligochaete Worms from the Punjab. Phosphorescence in Marine 
Animals. The rats of Dacca, Eastern Bengal. Freshwater Sponges, I—V. 
Miscellanea :—The original home of Mus decumanus. Colour change in Hylo- 
bates hoolock. Eggs of Tylototriton verrucosus. The hosts of Tachaea spongil- 
licola. A second species of Dichelaspis from Bathynomus giganteus. 


Part IV.—Nudiclava monocanthi, the type of a new genus of Hydroids parasitic on 


Fish. Three new Nycteribiidae from India. Annotated Catalogue of Oriental 
Culicidae. Oriental Diptera, IV. Freshwater Sponges, VI, VII. A new Cyprinid 
Fish of the genus Danio from Upper Burma. Mzscellanea :—A colour variety of 
Typhlops braminus. Reptiles and a Batrachian from an island in the Chilka 
Lake, Orissa. 


Vol. II, 1908-1809. 


Pavt I.—Retirement of Lieut.-Col. Alcock. The Fauna of Brackish Ponds at Port 


Canning, Lower Bengal, VII. A new Dictyonine Sponge from the Indian Ocean. 
Freshwater Sponges, VIII. Remarkable cases of variation, I. A new species of 
Lizard of the genus Salea, from Assam. The Fauna of Brackish Ponds at Port 
Canning, Lower Bengal, VIII. A new Cavernicolous Phasgonurid from Lower 
Siam. New species of Marine and Freshwater Shells in the Indian Museum. 
Oriental Syrphidae, I. A new variety of Spongilla loricata. Oriental Diptera, V. 
Miscellanea :— Remarks on Simotes splendidus. Corrections to No, IV of ‘‘ Notes 
on Oriental Diptera.’’ The Isopod genus Tachaea. The habits of the Amphipod, 
Quadrivisio bengalensis. New varieties of Nanina berlangeri and Corbicula 
fluminalis. Recent. additions to the Entozoa in the Indian Museum. A sub- 
fossil polyzoon from Calcutta. Corrections as to the identity of Indian Phylac- 
tolaemata. A peculiar form of Euglena. 


Part II,—Gordiens du Musée Indien. The Fauna of Brackish Ponds at Port Canning, 


Lower Bengal, IX. A new species of Danio from Lower Burma. Rhynchota 
Malayana, I. Czmex votundatus, Signoret. Freshwater Sponges, IX. Fruit 
Bats inhabiting the Andaman and Nicobar Archipelagos. A new species of Sun- 
Bird obtained near Darjiling. Three Indian Phylactolaemata. Two new species 
of Eagle-Rays (Myliobatidae). A new species of the genus Sesavma, Say., from 
the Andaman Islands. New species of Land, Marine, and Freshwater Shells from 
the Andaman Islands. 


Payt I[J.—The Fauna of Brackish Ponds at Port Canning, Lower Bengal, X, XI.- 


Oriental Solifugae. The difference between the Takin (Budorcas) from the Mishmi 
Hills and that from Tibet. Caridina nilotica (Roux) and its varieties. A new 
species of Chavaxes from the Bhutan Frontier. First report on the collection of 


Witics Awe VESTON. OF . Loe ORLEN TAT, 
SPE Gino OF THE GENERA IGF FEE 
Ravin, ve ‘TAB ANI DUA OULEE R 
a HU ACN TA ee aeine Sy. 


By GERTRUDE RICARDO. 


Family TABANIDAE. 
Subfamily TABANINAE. 


Hind tibiae with no spurs. Ocelli usually absent. 


The genera found in the Oriental Region, besides the large 
genus of Tabanus proper, are: Haematopota, Meigen, small flies 
with peculiarly marked wings; Udenocera, Ricardo, formed for a 
species from Ceylon with the long antennae situated on a tubercle ; 
Diachlorus, Macquart, including chiefly species from S. America 
distinguished by the simple antennae not situated on a tubercle, by 
the brown or yellowish markings of the wings and by the curved 
dilated fore tibiae, and slender build. The one species described 
from the Philippines is unknown to me. A new genus Neotabanus 
is now added, allied to Udenocera in having the antennae situated 
ona tubercle, but the antennae themselves are similar to those 
of the genus Tabanus. 


HAEMATOPOTA, Meigen. 


Illiger’s Magazine, ii, p. 267 (1803); Chrysozona,! Meigen, 
Nouvelle Classification, 23 (1800) ;. Kertesz, Cat. Dipt., iii, p. 201 
(1908). 

The described species from the Oriental region now amount to 
31, including 11 new species described in this paper. 

The following is a list of all the described species and syno- 
nyms :— 

[Notre.—The synonyms are printed in italics. ] 


HAEMATOPOTA annandalei, n. sp. 
assamensis, ll. sp. 
astatica, Rondani = javana, Wied. 
atomaria, Walker. 
bilineata, n. sp. 
borneana, Rondani. 


‘L For reasons against the adoption of this name see Verrall ‘* British Files,’’ 
Vv, p. 772 (1909) Austen, African Blood-Sucking Files, p. 121 (1909). 


Se) 
iS) 
N 


Records of the Indian Museum. [VoL. IV 


brevis, Ricardo. 
cana, Walker. 
cilipes, Bigot. 
cingalensis, Ricardo. 
cingulata, Wiedemann. 
cordigera, Bigot (nomen bis lectum) = fusci- 
frons, Austen. 
dissimilis, n. sp. 
fasciata, n. sp, 
fuscifrons, Austen. 
immaculata, n. sp. 
inconspicua, n. sp. 
irrorata, Macquart. 
javana, Wiedemann. 
lata, Ricardo. 
latifascia, n. sp. 
limbata, Bigot. 
lunulata, Macquart. 
marginata, n. sp. 
nigra, Wiedemann ? = javana, Wied. 
pachycera, Bigot. 
punctifera, Bigot. 
roralis, Fabricius. 
rubida, Ricardo. 
sinensis, n. sp. 
singularis, Ricardo. 
tessellata, Ricardo. 
unizonata, Ricardo. 
validicornis, n. sp. 


The small flies of this genus are easily recognized by the pecu 
liar marking of the wings, three rosettes more or less distinct, form- 
ed by pale marks on the darker ground colour are usually present, 
though in a few species they are absent and only pale streaks are 
present. The great majority of the species have typical paler 
rings of colour on the middle and posterior tibiae, occasionally 
absent, or only present on the middle pair, this character is some- 
times indistinct owing to denudation. In the grouping of the 
species I have taken this character as a means of division, after- 
wards relying chiefly on the shape of antennae (in the females 
only) and on the form of the apical band of wing. As many more 
species will probably be discovered from India and other Oriental 
districts it seems useless as yet to draw up a table of species. 


Synopsis of the Divisions and Groups of Haematopota. 


Division I. Legs uniform in colour with no typical rings on 
the tibiae af =f a3 .. Group” I. 
Division II. Legs not uniform in colour, but with the base of 
the fore tibiae at least white or yellowish, no rings on 
tibiae me Se ie :. Group. TT: 


IQII.| G. Rrcarpo: The Oriental Tabanidae. 323 


Division III. Legs not uniform in colour, but with typical rings 

on the middle tibiae only, all tibiae whitish at base Group I/I. 

Division IV. Legs not uniform in colour, but with typical rings 

on the middle and posterior tibiae Groups IV, V, VI, VII. 

Group IV. First joint of antennae abnormally large and 
stout, and as long as or longer than the third joint. 

Group V. Third joint of antennae broad and flattened on 
the first annulation, the last three annulations forming a 
narrow apex, the first joint incrassate, nearly as long as 
or longer than the first annulation of third joint. 

Group VI. Third joint of antennae not very broad at base, 
the first joint short, often only half as long as or at least 
always shorter than the third joint which is usually long 
and slender. 

Group VII. Antennae long and slender, the first joint as long 
as the second and third together. 


DIVISION I. 
Group I. 


Legs uniform in colour, with no rings on the tibiae or pale 
bases to fore tibiae. 


Haematopota roralis, Fabr. 
(Plate xvii, fig. 24.) 


Syst. Antl., 107, 2 (1805) ; Wied. Dipt. Exot., 97, 1 (1821) ; 2d., 
Auss. zweifl. Ins., i, 215 (1828); Bigot, Bull. Soc. Zool. France, 
xvi, 78 (1891). 

Wings with white spots, abdomen brown, with three stripes 
and spots, white. Habitat—Tranquebar. Related to H. pluvialis. 
Antennae elongate, yellow. Head ashy grey with two frontal 
black spots. Thorax dark, with an ashy grey stripe. Abdomen 
elongated, brown, with three white stripes and spots. Median 
stripe and spots linear. Wings dark with very numerous white 
spots and little lines. Legs testaceous. Fabr., Syst. Antl., 107. 

Brownish grey. Thorax and abdomen with white stripes. 
Wings with hyaline little spots. @ 4%1. From Tranquebar. 

The head is gone in the Fabrician type and the abdomen is 
crushed ; but it is easily distinguished from H. plwtalis, by the 
legs being unicoloured, blackish ' ochre-brown, not variegated, and 
by the chestnut brownish costal border and stigma of the wings ; 
segmentations of the abdomen reddish, but of the under side greyish 
white. In the Fabrician collection. Wied., Auss. zweifl. Ins., 1, 
p. 215. 

In Brit. Mus. coll. a male from Velverry, a female from Hot 
Wells, Trincomalee, and another female from Pankullam Road, 
Trincomalee ; Ceylon (Yerbury), 1891. 


1 In Dipt. exot, Wiedemann describes the legs as ochre-brown. 


324 Records of the Indian Museum. [VoL. IV 


In Indian Museum, males and females from Calcutta; and 
Goalbathan, E Bengal. 

In Howlett coll , a series of females from Gorakhpur, United 
Provinces; and from Belgatchia, Bengal, ‘‘ on cow.” 

A species distinguished by its testaceous legs with no darker 
bands, or base of fore tibiae lighter, and by the absence of any 
spots on the face. 

The female is long, slender, greyish brown, the abdomen 
marked with a median grey stripe and with lateral grey spots on 
each segment. Antennae yellow, long, the first joint nearly as long 
as the third one which is blackish at apex. Forehead with the usual 
paired spots, no unpaired spot present. Frontal callus yellow- 
ish, transverse. Wings viewed from the base usually appear 
pale in the basal cells and in the basal portions of the discal, first 
posterior and submarginal cells, elsewhere greyish brown, with 
three rosettes distinct, the apical band simple, short, a row of 
short whitish disconnected bands run from it round posterior 
border of wing. 

The males are more reddish yellow on the abdomen, the 
first joint of antennae short, stout. 

Length of males g—1r mm., of females 8}—1I mm. 

9. Face greyish with some short white pubescence, the 
fovea alone appearing darker. Palpi pale yellow with black 
pubescence, stout, ending in an obtuse point. Antennae reddish yel- 
low, the first joint yellowish cylindrical, slightly incrassate, 
barely as long as the third joint, the second joint very small, yel- 
low, both the two first joints with black pubescence, the third 
joint slender reddish, blackish on the annulated apex. Forehead 
same colour as face or a shade darker, with some white pubes- 
cence, broad, hardly narrower at vertex, the frontal callus red- 
dish yellow, shining, reaching the eyes, produced to a short point in 
the middle of its upper border, a narrow spot proceeds from its 
lower border to between the antennae, the paired spots blackish, 
not touching eyes. Thorax blackish brown with three grey stripes 
and sides grey, the scanty pubescence consists of short greyish 
yellow hairs: scutellum blackish brown, largely covered with greyish 
tomentum. Breast greyish. Abdomen blackish brown or brownish, 
with a distinct ashy grey median stripe, and grey roundish spots 
on each segment not reaching the anterior or posterior border, 
these markings are only distinct in well-preserved specimens, the 
pubescence chiefly consists of very fine short yellowish hairs, 
under side uniformly greyish. Legs reddish yellow, coxae greyish, 
base of femora sometimes darker, tibiae not incrassate or hardly so. 
Wings greyish, paler at base and at base of submarginal, first pos- 
terior and discal cell, though not always very distinctly so, the 
stigma and veins yellow, an appendix present on fork of third 
longitudinal vein, the two upper rosettes distinct, the third small, 
its outer ring incomplete, continued from above across the anal 
cell into the axillary lobe of wing, reaching the border, the short 
pale bands begin from the anal cell and continue round the border 


TGLE:| G. Ricardo: The Oriental Tabanidae. 325 


of wing to the apical band which is small and often inconspicuous, 
crossing the anterior branch of third longitudinal vein, but not 
always reaching the second longitudinal vein. 

@. Similar, but the abdomen is largely reddish yellow, darker 
at apex. Eyes with the large facets occupying two-thirds of sur- 
face, coppery coloured, the basal small facets blackish. Face more 
hairy. Palpi pale whitish yellow. Antennae with the first joint 
incrassate only about half as long as the third. Frontal triangle 
with small yellowish callus, the spot between the antennae brown. 
Hind tibiae fringed with black hairs on each side. Fore tibiae 
more filiform than in female. 


Haematopota limbata, Bigot. 


Bull. Soc. Zool. France, xvi, p. 78 (1891) ; 7d., Mem. Soc. Zool. 
France, v, p. 626 (1892) ; Ricardo, Ann. Mag. Nat. Hist. (7), xviii, 
p. I15 (1906). 

Antennae, palpi and proboscis dull fawn coloured ; beard grey; 
face whitish below, above dark shining chestnut; forehead 
whitish with two side black spots; the first segment of antennae 
hardly incrassate; thorax dull reddish; the dorsum with four 
diffuse blackish stripes, scutellum blackish, sides whitish with 
hairs of the same colour (or, in the Latin, pleurae ashy grey) ; 
abdomen brownish (in Latin, chestnut coloured) with a wide dor- 
sal stripe very regular and greyish white, all the segments narrowly 
bordered with yellow, with a lateral blackish spot; pleurae 
and halteres pale fawn coloured ; legs uniformly pale fawn coloured ; 
wings very pale yellow, the stigma narrow, reddish, and all the 
veins bordered with a pale reddish colour, some indistinct very 
pale white markings. India: one specimen, 9,11 mm. Bigot, 
Mem. Soc. Zool. France, v, p. 626. 

The author in his first notice of the species in his table for 
Haematopota in Bull. Soc. Zool. France, xvi, p. 78, describes the 
antennae as almost entirely yellowish, abdomen chestnut coloured 
with a wide whitish dorsal stripe ; wings yellowish with pale indis- 
tinct spots. 

From the type kindly lent me by Mr. Verrall in 1906 I made 
the following desctiption, published in the ‘‘ Annals’’ as above :— 

Type (female) from Bengal, and another female from Khasi 
Hills. A fair-sized species, easily distinguished by the prominent 
bluish grey median stripe of abdomen, with large black spots on 
the upper part of the face. 

Brown. Face grey ; a large irregular-shaped black spot on each 
side of antennae, reaching to the eyes. Frontal callus yellow, 
shining, narrow, concave on the posterior or lower border; the 
spot between the antennae black. J orehead grey, the paired spots 
black and distinct, the unpaired spot brown, indistinct. Antennae 
yellow ; the first joint stout, not so long asthe third joint, with 
black pubescence ; the second joint very small, with black hairs ; 
the third joint broad, becoming narrower where the annulations 


) 


326 Records of the Indian Museum. [VOL sive 


begin, and tapering to a point. Palpi yellow, with dense 
black pubescence. Thorax blackish brown, lighter coloured at 
the sides, with faint narrow grey stripes, the breast with hoary 
markings. Abdomen brown, the posterior borders of the segments 
narrowly yellowish; some faint black markings on the sides of 
abdomen; the under side brown, covered with grey tomentum. 
Legs yellow, with fine black pubescence ; the coxae grey pollinose. 
Wings grey, with yellow veins and along appendix; most of the 
veins are faintly shaded with darker colour ; the typical markings 
are faint : there is one rosette apparent, enclosing the appendix, 
and another beyond. Length 11} mm. 

In Indian Museum are two females from Khasi Hills, Assam. 
One of these islabelled H. limbata, n. sp., Bigot, in his handwriting 
and no doubt may be considered a para-type. Both agree in. all 
particulars with the descriptions, the apical band of wing is simple, 
narrow, but curved on the part crossing the anterior branch of 
third vein. 


DIVISION II. 
Group II. 


Legs not uniform in colour, with no rings, but with the base 
of fore tibiae ateleast white or yellowish. 


(A) Wings with no typical rosettes but with a pale band and streaks. 
Haematopota cingulata, ? , Wied. 


Auss. zweifl. Ins., i, p. 216 (1828); Bigot, Bull. Soc. Zool. 
France, xvi, p. 79 (1891); Ricardo, Ann. Mag. Nat. Hist. (7), 
XVili, p. I15 (1906). 

Thorax brown, with a wide yellow stripe ; abdomen blackish 
with whitish segmentations; wings brown, with base, band, and 
two curved streaks, hyaline. 9? 44 lines. From Java. 

Antennae leather-yellow, with the apex of third joint blackish. 
Palpi blackish. Face blackish brown, faintly greyish white at sides 
with black dots ; cheeks blackish brown, forehead shining blackish 
brown, above in certain lights grey with two brown contiguous 
spots. The yellow stripe of thorax is as broad as the full clear 
brown sides and is continued on to the scutellum, breast sides full 
clear brown, breast greyish. Abdomen blackish brown, the first 
segment with a broad triangular whitish spot which reduces the 
ground colour at the base to a triangular spot; second segment 
with a narrower, triangular spot which attains the base and at its 
own base unites with the white segmentation; the following seg- 
ments merely with white segmentations; perhaps in perfect un- 
rubbed specimens these may have white hairs and. triangular 
spots. Under side the same colour, likewise with white segmenta- 
tions. Wings at the base as far as the middle cross-veins fairly 
hyaline, only yellowish between the last two veins and on the 
costal border ; otherwise blackish brown; the hyaline band lies a 


IgII. | G. Ricarpo: The Oriental Tabanidae. 327 


little behind the middle cross-veins, is somewhat oblique, and is 
narrower towards the fore border where it meets the posterior end 
of the stigma, on the inner border it is abbreviated, behind it lies 
the first streak which starting from the fore border, decreasing, 
curves forward bow shaped and ends in the band; the second 
streak nearer the apex begins from the fore border close to the 
apex of the second vein and runs curved forward, so that it gradu- 
ally approaches the first one and ends on the inmost part of the 
band: the inner border of the wing is brown much further towards 
the base than to the middle. Halteres white. Legs blackish brown, 
base of tibiae white, posterior femora brown at the base, at the 
apex to far beyond the half, honey yellow. In the Leyden 
Museum. Wied., Auss. zweifl. Ins., 1, p. 216. 

The reference given by Kertesz in both of his Catalogues, w7z., 
Cat. Tabanidarum, 1900, and Cat. Dipt., vol. iti, 1908, to v. d. 
Wulp, Tijd. v. Entom., xxxiv, p.197, 1891, isincorrect. In Wulp’s 
Cat. of Oriental Diptera, 1896, there is no such reference against 
H. cingulata, and I have not been able to find any reference to 
this species by v. d. Wulp, and it is unknown to me. It should be 
an easily identified species by means of the wings which apparently 
have not the usual rosettes, but are allied to those of H. vubida and 
singularis, Ricardo. fi 


(B) Wings with the usual rosettes, the apical band double. 
Haematopota punctifera, Bigot. 
(Ploxyi fig: 20") 


Mem. Soc. Zool. France, v, p. 629 (1892); Ricardo, Ann. 
Mag. Nat. Hist. (7), xviii, p. 125 (1906). 

Antennae incomplete, blackish at base, first joint very short ; 
face and palpi whitish ; forehead wide, dull black with grey tomen- 
tum, and a large shining black callus above the antennae; thorax 
dark chestnut coloured with four greyish white stripes ; scutellum 
brown, greyish in the middle; abdomen blackish brown, sides of 
segments, a narrow, interrupted, median stripe, and two spots on 
each segment, greyish ; squamae grey, halteres brown ; legs brown, 
tibiae pale yellow, black at apex, tarsi black, anterior tibiae almost 
wholly black; wings grey, with fine white marking (specimen 
damaged). The shortness of the first antennal joint makes the 
exact position of this species doubtful. Length 11 mm.: Java, one 
specimen. Bigot, Mem. Soc. Zool. France, p. 629. 

The following description was published by me after seeing 
the type :— 

The antennae are incomplete; the first joint red, short, and 
incrassate, the second one red, small. 

Face grey, with white hairs, no spots, but a dark brown band 
between the eyes and the antennae. Palf: reddish, with black 
pubescence. Frontal callus black, shining, reaching the eyes, 
narrow, curved on the posterior border and produced to a point in 


328 Records of the Indian Museum. [Vor. IV, 


the centre ; a black spot between the antennae. Forehead (denu- 
ded) blackish, with grey tomentum and some golden pubescence. 
Thorax (denuded) brown, with grey stripes and tomentum and some 
black and grey pubescence ; sides grey, with traces of fulvous and 
grey pubescence ; scutellum brown, with median grey stripe and 
greyish pubescence. Abdomen brown, with grey median stripe, 
grey spots on each side, and grey sides; the greyish pubescence 
rather dense. Legs reddish brown, the tibiae yellower, the an- 
terior ones whitish at the base, brown at the apex ; the anterior 
tarsi brown, the middle and posterior tarsi yellow, darker at the 
joints and apex ; coxae with rather long white pubescence ; femora 
with short white pubescence; tibiae and tarsi with some black 
pubescence. Wangs greyish, with appendix; veins yellowish brown ; 
stigma brown; a darker spot is visible above the stigma. Length 
8 mm. Ricardo, Ann. Mag. Nat. Hist. (7), xviii, p. 125. 

(C) Wings with the usual rosettes, the apical band single. 

Haematopota indiana, 2 , Bigot. 
(Pl: xvii, figs r9,) 


Bull. Soc. Zool. France, xvi, 78 (1891); 7zd., Mem. Soc. Zool. 
France, v, p: 026 (1892); Ricardo. Ann: Mage Nat.) mist (7) 
XVili, p. 124 (1906). 

Antennae double aslong as the-head, first and second joints 
reddish yellow, the third brown, at the base a little reddish yellow, 
the first cylindrical, the third not incrassate; palpi chestnut col- 
oured, proboscis brown; cheeks and face white below ; forehead 
ashy grey with two black spots, and with a black, shining, anchor- 
shaped spot below ; thorax chestnut coloured with four ashy grey 
stripes, scutellum chestnut coloured, pleurae ashy grey; abdomen 
brown, with white segmentations and with a median broad white 
stripe; anterior and intermediate femora pale chestnut, paler at 
their apices, the posterior ones dull brown, tibiae not with rings, 
black, at the base widely white, anterior tarsi black, intermediate 
and posterior ones white at base; pleurae brown, halteres reddish 
yellow with apex black ; wings pale cinereous, at the apex diffusely 
pale brown, and ornamented with numerous white spots and lines. 
Bigot, Mem. Soc. Zool. France, v, p. 627. 

In Indian Museum one female from Tezpore, Assam. 

The following description was published by me in the 
“Annals ’’ after an examination of the type :— 

A brown species, distinguished by its long slender antennae 
and all the tibiae white at the base. 

Face grey, dark brown in the centre, immediately below the 
antennae ; the stripe between the antennae and the eyes brown, 
but broken upinto dots and spots. Palpi reddish, rather long, with 
brown pubescence and some white hairs; beard white. Frontal 
callus narrow, long, reaching the eyes, shining brown, both borders 
nearly straight ; the spot between the antennae black, bordered 


d 


IQII.] G. Ricarpo: The Oriental Tabanidae. 329 


with grey; the paired spots dark brown, large, reaching the eyes 
and almost touching the callus ; the unpaired spot brown, small ; 
forehead yellowish, with brown markings and grey tomentum and 
short black pubescence. Awtennae long, slender, yellow; the 
third joint dusky, reddish at base, black at apex ; the first joint 
as long as the part of the third joint which is unannulated, the 
second joint small, both with black pubescence. Thorax brown, 
with grey shoulders and three narrow grey stripes ; short yellowish 
pubescence is visible on the dorsum; the sides brown with black 
hairs, the breast grey. Abdomen a redder brown, with light nar- 
row borders to the segments, darker at the apex; the pubescence 
on the light borders short and yellowish, on the other parts black ; 
a hoary grey median stripe is very distinct from the second to the 
fifth segment ; the under side brown, grey at the sides. Legs red- 
dish brown, the fore tibiae and tarsi darker brown, with black pu- 
bescence, the base of all the tibiae whitish, on the middle tibiae the 
white extends further ; the basal joint of the middle and posterior 
tarsi whitish. Wings brown, with yellowish brown veins and 
stigma and an appendix ; the apical band single, broad; the first 
second, third, and fifth posterior cells light coloured at their apices ; 
the dark spot on the fifth posterior cell (as shown in figure) is only 
noticeable in the wing in certain lights. Lengthg mm. The type 
is labelled ‘‘ Margherita, 5373. 8.” 


Haematopota borneana, 2 , Rondani. 


Ann. Mus. Civ. Genova, vii, p. 461 (1875); Bigot, Bull. Soc 
Zool. France, xvi, p. 78 (1891). 

Length 6—7 mm. @°. 

Antennae with the first joint yellowish brown; the second 
pitchy blackish with the greater part of the third, whose apex is 
black. 

Forehead dull brown, with two rather small black spots and 
a wide shining ferruginous brown anterior callus. Face and palpi 
reddish with two small impressed spots at sides. Thorax with the 
scutellum above red, the intermediate stripe and side spots brown- 
ish, not very distinct ; pleurae paler coloured. Abdomen at the 
base yellowish brown, then black, with white segmentations to the 
posterior segments, wings brownish, the costal border towards the 
apex browner, white spotted, the spots small, and a white trans- 
verse stripe distinct at apex. Halteres with a white stalk and 
ferruginous club. Legs with anterior and posterior femora pitchy 
black, the intermediate ones yellowish brown ; all the tibiae white 
with the apex blackish ; anterior tarsi wholly black, the posterior 
ones reddish at base. Rondani, Ann. Mus. Civ. Genova, vii, 
p. 461. 

I saw the type in the Museum at Genoa in the spring of Igo9, 
it is not in good condition, 64 mm. long. A small species with a 
red thorax marked with a broad black median stripe and traces of 
lateral ones. Jace dull reddish, convex, no dark spots on it, the 


330 Records of the Indian Museum. IAYCo) oe Oe 


frontal callus reddish brown, shining transverse, produced in the 
middle on posterior border near antennae. Forehead reddish, no 
other spots visible. Antennae yellow, the first joint nearly as long 
asthe third. Palpi yellow. Wings with the apical band single, the 
first rosette distinct. 


Haematopota cana, 2 , Walker. 


List Dipt. Brit. Mus., i, p. 207 (1848) ; Bigot, Bull. Soc. Zool. 
France, xvi, p. 79 (1891); Ricardo, Ann. Mag. Nat. Hist. (7) 
XVili, p. I16 (1906). 

Body hoary ; head white beneath ; eyes bronze colour ; feelers 
black, a little longer than the head; first joint tawny towards the 
base; mouth black; palpi hoary, tawny towards the tips ; chest 
with four indistinct whitish stripes ; abdomen with a broad brown 
stripe on the back; hind borders of the segments pale tawny ; 
legs piceous ; shanks (tibiae) tawny, with piceous tips; four hin- 
der feet (tarsi) tawny at the base; wings very pale grey, adorned 
with numerous colourless segments of circles ; wing ribs and veins 
tawny, the latter piceous towards the tips of the wings ; poisers 
tawny. Length of the body 34 lines ; of the wings 7 lines. 

N. Bengal. From Miss Campbell’s collection. Walker, List 
Dipt. Brit. Mus., i, p. 207. 

Type (female), Northern Bengal, 42, 25 (Lieut. Campbell). 

A small yellow-brown species, with large grey spots at the 
sides of abdomen and an indistinct median stripe. 

The type is in very poor preservation, which makes any des- 
cription of it incomplete. 

Face grey, no spots; the frontal callus brown, both its borders 
somewhat irregular; a small brown spot is situated between the 
antennae ; the paired spots are brown, large, oblong, the unpaired 
one is small. Forehead grey. Antennae brown, the first joint yel- 
lowish, stout, not quite so long as the third; the falfi vellow. 
with black pubescence. Thorax blackish, with three grey stripes, 
the sides and breast greyish. Abdomen yellowish, darker at the 
tip, with large hoary grey spots on the sides; the median stripe 
appears incomplete, the hind borders of the segments narrowly 
yellow. Legs yellow; the fore coxae very long ; the femora brown- 
ish; the fore tibiae brown, yellow at the base: the tarsi brown at 
the tips. Wings grey, the veins yellow, with an appendix; the 
light rosettes and spots fairly distinct. Length 8 mm. Ricardo, 
Ann. Mag. Nat. Hist. (7), xvili, p. 116. 

This type being so deteriorated it is impossible to place it even 
with any certainty inthis group. The wings bear some resem- 
blance to those of H. voralis, Fabr., the basal cells being largely 
pale, the apical band is single lineal, curved towards the apex 
where it joins the first small transverse band, which joins the 
succeeding one, all these bands forming a nearly continuous line 
round the border of wing, no pale openings are visible in the pos- 
terior cells, three rosettes are distinct ; only one wing remains. 


d 


IgII.| G. Ricarpo: The Oriental Tabanidace. 331 


Division III. 
Group ITI. 


Legs not uniform in colour but with rings on the middle tibiae 
only, all tibiae white at base. 


(A) Apical band of wing double. 
Haematopota dissimilis, @ , n. sp. 


(Plate xvii, fig. 22.) 


In British Museum type (female) and others from Baste, and 
Gersoppa, N. Canara, India, 1907 (T. R. Bell). 

A species allied to H. bilineata, n. sp., in the wing but distin- 
guished from it by the first joint of antennae being cylindrical 
hardly incrassate, yellowish or yellowish brown in colour, the frontal 
callus is also broader. Tibiae not incrassate, the hind tibiae with 
a white-haired ring at the base, the middle tibiae only with typical 
rings, in this allied to. atomaria, Walk., but distinguished from it 
by the cylindrical antennae and narrow frontal callus. Apical band 
of wing double, the upper branch proceeding from the inner bor der, 
the low: er and larger one from the outer border. Length 81mm. 

Face ashy grey with some short black pubescence, a brown 
band is present on upper part, broken up into spots, so that the 
ground colour appears yellowish brown, and the spots are dark 
brown or black. Palfz with the second joint swollen at base end- 
ing in a narrow apex, obscurely reddish, with black pubescence. 
Ajiteanae reddish yellow, slender , long,” the first joint fully as long 
as the first annulation of third joint, shining with black pubes- 
cence, hardly incrassate, the second joint very small cup shaped, 
likewise with black pubescence, the third joint slender, only slightly 
broader at base, dusky in colour at tip; a large black spot is pres- 
ent between the antennae. Forehead brownish, the frontal callus 
dark blackish brown, shining, almost straight on both borders. 
in the type produced slightly on the upper border, reaching 
the eyes anteriorly and posteriorly, occupying in breadth about 
a third of the width of forehead; the paired spots large, 
black, touching the eyes and the frontal callus, the paired spot 
small, triangular. Thorax brown with some yellowish brown 
tomentum and short pale pubescence, shoulders with some ashy 
grey tomentum. Scutellum similar to thorax. Abdomen choco- 
late-brown with narrow grey ‘segmentations, under side similar, 
pubescence on dorsum scanty, white on segmentations, elsewhere 
black. Legs blackish, the base of fore tibiae whitish and the usual 
tings on the middle tibiae reddish yellow or whitish, often indis- 
tinct, hind tibiae with only a white ring near base and sometimes 
a tuft of white hairs beyond, the white base of fore tibiae is more 
correctly a white ring near the base, fore tibiae hardly incrassate, 
femora reddish yellow, the basal joint of middle and posterior tarsi 
pale. Wings very similar to those of H. bilineata, n.sp., the apical 


332 Records of the Indian Museum. [ Vier. stv, 


band in type has the upper branch proceeding from the inner 
border very insignificant consisting of a very short narrow line 
not reaching .the anterior branch of the third vein, but in the 
other specimens it usually crosses it or at least reaches it, the lower 
and larger branch is wide at its base, becoming narrower as it 
proceeds, crossing the upper branch of third vein. 


(B) Apical band of wing single. 


Haematopota atomaria, 9 , Walker. 
(Plate xvi ois. 11.) 


Proc. Linn. Socs London; 4; p. 112 >1(1856) ;) Ricardo,” Ant 
Mag. Nat. Hist. (7), xviii, p. 117 (1906). 

In British Museum coll. 

Type (female), Sarawak, Borneo (Wallace), 56, 44, and two 
other females from Sarawak, 57, 36. 

A small dark species, with brown wings, distinctly marked 
with the usual rosettes and spots; the first joint of the antennae 
incrassate. 

Face grey; two small black spots beneath the antennae ; the 
upper part of the face with a brown stripe, which becomes yellow- 
ish near the antennae; the palpi yellow, with black pubescence ; 
the beard white. Frontal callus pitchy brown, shining, broad, 
reaching the eyes, with the posterior border convex, rounded, the 
anterior border reaching round the antennae; the paired spots black, 
round, reaching the eyes and the frontal callus, with yellow borders. 
Forehead brown, yellower on the vertex; the unpaired spot not 
present. Antennae blackish ; the first joint dark red, shining, with 
black hairs, slightly incrassate, considerably shorter than the third ; 
second very small; third broad, ending in a point. Thorax brown 
with traces of darker stripes and of short white pubescence; scu- 
tellum the same colour ; breast brown. Abdomen brown, with nar- 
row greyish borders to the segments, which are broader on the 
under side. Legs reddish brown, with black pubescence, which be- 
comes fringe-like on the hind legs; the base of the anterior tibiae 
white, the middle tibiae with the typical rings, the posterior tibiae 
brown at the extreme base, then white and brown on the apical 
half. Wings brown, with brown veins and an appendix; the 
apical band single; all the posterior cells and apical cell with a 
triangular white spot at their openings with the exception of the 
fourth posterior cell. Lengthg mm. Ricardo, Ann. Mag. Nat. 
Mist (7), xvillp. Lr. 

The posterior tibiae are white at base. 


Haematopota unizonata, Ricardo. 
(Plates ies -12. 132) 


Ann. Mag. Nat. Hist. (7), xvili, p. 118 (1906). 
Type (male) from Ceylon, 1892 (Yerbury). 


1QII.] G. Ricarpo: The Oriental Tabanidae. 333 


Type (female) from Hakgala, and three females from Niu- 
wara Eliya, Ceylon, 1891 (Yerbury), females from Pundaluoya, 
Ceylon, 1890 and 1898, and one female from Galagedara, Ceylon, 
1897 (Green). 

There is a note by Col. Yerbury on this species, viz., “‘ Very 
common at Niuwara Eliya, May 189If.’’ 

A reddish brown species, with rings on the middle tibiae 
only; the other tibiae white at the base; the antennae long, 
cylindrical. 

Face grey; a yellowish stripe covered with brown dots and 
spots reaches from the antennae to the eyes on each side; palpi 
yellow, with white pubescence; beard white. Frontal callus dark 
brown, shining, concave on the anterior (lower) border, convex on 
the posterior (upper) border; the spot between the antennae black, 
oblong: the paired spots black, oblong, just reaching the eyes, 
with grey borders. Forehead brownish yellow, darker on the 
vertex, grey at sides. Antennae long and slender, reddish yellow, 
the third joint darker; the first joint not quite so long as the 
third, the second small, both with black pubescence. Thorax 
reddish brown, the shoulders, two stripes which reach the suture 
and end in two spots, the base of thorax and a spot on each side 
grey; traces of a short white pubescence on the dorsum; breast 
brown with brown pubescence, then grey with white pubescence ; 
scutellum reddish brown. Abdomen a redder brown, segments 
bordered with narrow whitish bands; pubescence black, white on 
the borders of the segments and at the sides; traces of grey spots 
on the apical segments; the under side brown, with white borders 
to the segments, grey at the sides. Legs dark brown, the middle 
tibiae redder, with two white rings, and the base of the tarsi yellow- 
ish, as are also the posterior tarsi; the middle femora are 
reddish, with white pubescence. Wings brownish, with brown veins 
and an appendix, the light markings distinct, with three rosettes, 
the apical band single. Length 8mm. Ricardo, Ann. Mag. Nat. 
Hast. (7),. x<vili,.p. 118. 

Distinguished from H, atomaria, Wlk., by the narrower frontal 
callus, and by the longer cylindrical first antennal joint. 


DIvIsiIon IV. 


Legs not uniform in colour but with typical rings on the 
middle and posterior tibiae. 


Group IV. 
The first joint of antennae abnormally large and stout, and 
as long as or longer than the third joint. 
Haematopota validicornis, 2 , n. sp. 


(Plate xvii, fig. 23.) 
In Brit. Mus. coll. 


334 Records of the Indian Museum. [Vion EVE 


Type (female) from Biserat, Siam (Robinson and Annandale) ; 
others from Phrapatoon, Siam (Dr. P. G. Woolly); and from 
Siam (W. Palmer), : 

In Indian Museum females from Biserat, Siam (Robinson and 
Annandale). 

This and the following species are distinguished by the very 
large stout first joint of antennae as long as the third joint. The 
antennae in this species are reddish yellow, the face with no black 
band but one spot on each side touching the eyes, about half 
way between the antennae and the oral opening. Frontal callus 
yellowish brown. Thorax with grey stripes stopping short at 
suture, two others meeting them from the base. Abdomen black- 
ish brown with grey spots. Wings with the apical band single, 
broad, legs with the usual rings on tibiae, hind tibiae fringed. 
Length 9 mm. 

Face greyish, where the dark band is usually apparent it is more 
yellowish brown, the black spot irregular in shape, pubescence of 
face whitish. Palfi pale yellow with pale hairs at base, and black 
pubescence. Antennae reddish yellow, the first joint very large, 
incrassate, as broad as the third joint at its widest part, with 
black pubescence, the second joint very small with thick black 
pubescence, the third joint broad and flattened on its basal annu- 
lation with the three last ones forming a small apex. Forehead 
greyish with black pubescence, the paired spots touching the eyes 
and the frontal callus, almost round in shape. Frontal callus 
yellowish brown, shining, almost straight on both borders, and 
reaching the eyes on lower border, its outer angles rounded reced- 
ing from eyes, no well-marked spot between the antennae but 
brown rings surround the base of each antenna. Thorax brown 
with short yellowish pubescence, the grey stripes dispersed as in 
H, javana, Wied., and the base of the thorax grey. Scutellum 
brown with grey tomentum in centre. Abdomen blackish brown, 
or reddish brown with ashy grey segmentations and round lateral 
spots and a grey median stripe is often apparent. Under side 
blackish brown with grey segmentations, Legs blackish brown, 
the fore tibiae at base yellowish, the middle and posterior tibiae 
brownish, each with two yellowish rings, the tarsi on the basal 
joints of middle and posterior pair yellowish, the femora with 
black pubescence, thick below, on the hind pair thickest, above. 
and below atapex. The fore and hind tibiae slightly incrassate, all 
with black pubescence, which is thickest on the hind pair, forming 
fringes. Wings brownish, the apical band in type is continuous, cross- 
ing the whole width of apex, crescent-shaped, concave towards the 
apex, in the other specimens often broken in the middle, the small 
white spot above stigma has a dark centre, which is occasionally 
absent, another white spot meets it, partly surrounding the ap- 
pendix and forming the centre of the first rosette, the second 
rosette is distinct, encircling the small transverse vein and apex 
of discal cell, the third and fourth rosettes are fairly distinct 
below, on the hind border of wing white marks are present in the 


IgiI.] G. Rrcarpo: The Oriental Tabanidac. 335 


first, second, third and fifth posterior cells but are small, that in 
the first cell being largest, below them short bars of white colour 
appear. 


Haematopota annandalei, @ , n. sp. 


(Plate xvii, fig. 21.) 

In Brit Mus. coll. 

Type (female) from Govt. Gardens, Shillong, Assam; and 
another from Khasi Hills. 

In Howlett coll. females from Govt. Gardens, Shillong, and 
from Umling, Assam, ‘‘ on cattle.’’ 

Distinguished by the very stout large first joint of antennae, 
differing from H. annandalet by the presence of a black band 
on face, and the antennae are shining black, the third joint 
is narrower. The apical band of wing is single but a different 
shape, being short and curved, not reaching the posterior border. 
Frontal callus black, shining. Length 83 mm. (type), other females 
7—8 mm. 

Face greyish with rather thick long white pubescence, the band 
between eyes and antennae a deep black, extending across the face 
below antennae, the palpi small, greyish, with white pubescence, 
darker at tip. Antennae black shining, the first joint longer than 
the third one, very stout along its whole length, the second joint 
very small, the third joint much narrower than the first joint, the 
first annulation being hardly broader and the last three annuli 
only slightly narrower. Forehead a darker grey colour, almost 
brown with some grey tomentum, the frontal callus narrow, black 
and shining, reaching the eyes on its lower border, above with 
rounded angles receding from the eyes, almost straight on both 
borders, the paired spots large, black, touching the frontal callus 
with their apices, and the eyes with their bases, the unpaired spot 
narrow, distinct. Thorax blackish brown with scattered short 
yellow pubescence, two grey stripes are apparent ending at the 
suture, two similar stripes proceed from the base of thorax 
towards them; sides of thorax a little grey. Scutellum same 
colour as thorax. Abdomen blackish brown with grey seg- 
mentations, round small side spots are visible in some of the 
specimens ; under side dark with grey segmentations. Legs black- 
ish, the base of fore tibiae and two rings on the other tibiae, 
and basal joints of middle and posterior tarsi yellowish or white, 
pubescence on femora chiefly black, scanty, a little thicker in pos- 
terior pair on whose upper side some white pubescence is visible, 
tibiae slightly incrassate on fore and posterior pair, with black 
pubescence on the dark partsand white on the pale rings, the hind 
tibiae fringed with thicker black hairs, tarsi with black pubescence. 
Wings brown with the apical band simple, short, beginning just 
above apex of second vein crossing the first branch of third vein 
ending just beyond it, it has an outward curve towards the base 


! This is perhaps a mistake for va/iditcornis.—Ep, 


336 Records of the Indian Museum. [Vor <tVi; 


of wing and is concave on its apical border, the round white spot 
above the stigma is distinct and with no brown centre in the type, 
in other specimens it is present as a half circle, three rosettes are 
clearly formed and pale spots appear in the apices of the first, 
third and fifth posterior cells, and smaller ones occasionally in 
the others. 


Haematopota pachycera, 2 , Bigot. 


Nouv. Archiv. Mus. Hist. Nat. Paris (3), ii, p. 206 (1890) ; 7d. 
Bull. Soc. Zool. France, xvi, p. 76 (1891); Ricardo," Ann. Mag. 
Nat. Hist (8), i: p. 59 (1908). 

Length 8} mm. 

Antennae twice the length of head, yellow, the first joint some- 
what incrassate, cylindrical, apex brown ; second joint stout, pyri- 
form, compressed, apex brown ; others yellow, small. Palpi, beard 
and face dull ashy grey. Forehead ashy grey, at base with a 
shining transverse yellow callus, brown at vertex with two frontal 
black spots. Thorax ashy grey, with wide indistinct brown stripes ; 
scutellum brown, abdominal segments with narrow grey borders, 
and side spots irregular in shape, of same colour. Wings almost 
black, with white markings. Legs brown, fore tibiae dull white at 
base, intermediate and posterior tibiae pale yellow, at base apex 
and ring brown. 

Antennae at least twice as long as head, yellow, the first joint 
cylindrical, rather thick, black at apex, the second shorter, thick, 
pyriform, truncated, laterally compressed, likewise black at its 
apex, the last joints indistinct and very small, bright yellow ; palpi, 
beard and face dirty grey, forehead same shade, with a large red- 
dish shining transverse callus situated above the base of antennae, 
and with two small black spots at border of eyes ; thorax blackish, 
indistinctly striped with grey lines, sides grey; scutellum brown ; 
abdomen blackish, border of segments and two series of little in- 
distinct spots greyish ; calyptrae and halteres yellowish grey ; wings 
blackish with rather extensive white markings and spots; legs 
brown, naked, fore tibiae whitish at base, intermediate and poster- 
ior ones pale yellow with the base, apex and a median ring 
blackish. 

Laos, one specimen. 

The unusual form of antennae, notwithstanding the general ap- 
peavance, seems to authorize the establishment of a new genus for 
this and the preceding species. Bigot, Nouv. Archiv. Mus. Hist. 
Nat: Paris: 1690, 11, p. 200: 

The following notes were published after I had seen the type 
in the Paris Museum :— 

This type from Cambodia (the description gives Laos) is also 
in the Paris Museum. 

This species would come under heading ‘‘ 15’ in my table of 
Indian and Ceylon species next to H. lata, Ricardo, from which it 
is distinguished by the hind tibiae not being fringed, and the third 
joint of the antennae is wider and shorter. It resembles H. rubida, 


IQIT. | G. Ricarpo: The Oriental Tabanidae. 3 
9 Jan 


Ricardo, in the antennae. Bigot seems to have overlooked the 
second joint, which is very small, and described the third joint as 
the second one; the first joint is long, cylindrical, and the third 
joint very wide, the basal division being large and swollen, the re- 
maining divisions very small. The abdomen is black, with the 
segmentations grey ; the two rows of spots mentioned by Bigot are 
hardly noticeable. The legs have two rings of light colour on the 
middle and posterior tibiae, and the base of the fore tibiae is 
yellowish. The wings have two indistinct rosettes. Ricardo, Ann. 
Mag. Nat. Hist. (8), i, p. 59. 

This species probably belongs to this group with the first joint 
of antennae very stout and it is possible that my new species 
validicornis is identical, though it is apparently distinguished by 
the presence of fringes of hair on hind tibiae, said to be absent in 
Bigot’s species ; a further examination of Bigot’s type with regard 
to the markings of wing is imperative before coming to a 
decision. 

Since writing this paper I have received from M. Surcouf of 
the Paris Museum a copy of his paper in Bull. Mus. Hist. Nat. de 
Paris, 1909, No. 7, p. 453, in which he establishes a new genus 
Potisa for this species and suggests my H. rubida will belong to it 
also, but as already remarked I have left this species near H. 
stngularvis owing to the peculiar markings of wings. The confor- 
mation of the first joint of antennae in these species H. pachy- 
cera, validicornts and annandalet hardly seems to justify the estab- 
lishment of a new genus divided off from Haematopota, as the 
shape of the antennae in Haematopota varies very considerably, but 
only in small gradations, while the typical markings of wings and 
other characteristics remain the same. 


Group V. 


The third joint of antennae broad and flattened on its first 
annulation, the last three annulations forming a narrow apex; the 
first joint incrassate, nearly as long as or longer than the first annu- 
lation of the third joint. 


This group is allied to the genus Parhaematopota established 
by Grunberg (Zool. Anzeig., xxx, p. 360) in 1906 for a new 
East African species which he named P. cognata. Speiser in Hy- 
men, Dipt., vii, p. 360 (1907), added H. vittata, Loew, to this new 
genus [Mr. Austen states this is incorrect, see African Blood-Suck- 
ing Flies, p. 122 (1909)] and remarks that H. decora, Wik. (H. dor- 
salis, Loew, is now asynonym of this species), and its related species 
—differing from other Haematopota species by the peculiar form of 
the third antennal joint—form a connecting link with Parhaema- 
topota, the establishment of which as a genus is only justified pro- 
visionally, in the present imperfect state of our knowledge. It 
certainly seems advisable to refrain for the present from creating 
new subgenera, and to rest content with grouping allied species 


338 Records of the Indian Museum. [Vor. IV 


together. The figure of the antennae of Parhaematopota is not 
unlike those of this group as regards the third joint, but the pro- 
duced upper angle of the first joint is not present in these Oriental 
species. 


(A) Wings with a pale oblique transverse streak and no rosettes. 
Haematopota rubida, Ricardo. 
(Plate xvii, fig. 18.) 


Ann. Mag. Nat. Hist. (7), xviii, p. 120 (1906). 


Type (female) from Burma, 57, 16 (Mrs. Waring). 

An easily distinguished red species, with the hind tibiae in- 
crassate ; the hind femora with a white bunch of hairs above and a 
fringe of black hairs on the under side. 

Face grey, the whole upper part deep black ; the palpi yellow 
with black pubescence and white hairs below. Frontal callus shining 
brown, protuberant, rounded, the posterior border produced, the 
anterior border with a deep incision in the middle, filled by the 
black spot usually present between the antennae, which is large 
and square with yellow borders; the frontal callus is short, not 
reaching the eyes; forehead grey, darker in the centre; the paired 
spots are black, small and isolated, the unpaired spot not present. 
Antennae are of an unusual form ; the first joint very much incras- 
sate and large, nearly as long as the third joint, yellow and shining, 
the second very small and narrow, yellow, both joints with black 
pubescence ; the third joint very broad, ending in an obtuse point, 
reddish yellow, darker at the tip. Thorax reddish brown, with 
lighter stripes, darker at the sides; the breast grey, with white 
hairs; the scutellum the same colour. Abdomen reddish brown, 
with very narrow yellow borders to the segments, darker at the 
apex ; the under side yellow, with grey tomentum. Legs reddish 
brown ; the anterior and posterior pairs the darkest, the anterior 
tibiae white at base, the middle tibiae yellowish brown, the two 
yellow rings not well defined; the posterior femora broad, with 
the fringe of hairs on the under side black, on the upper side black 
on the basal half, on the apical half a tuft of white hairs are pres- 
ent; the posterior tibiae very stout and broad, with ill-defined 
rings as on the middle pair, and fringed with black hairs above 
and below; the basal joints of the middle and posterior tarsi pale 
yellow. Wangs reddish brown, the veins yellow, with a long ap- 
pendix, the apical band single; the pale streak across the middle 
of the wing is very noticeable, and at once distinguishes the 
species from H. lata, n.sp. Length 10mm. Ricardo, Ann. Mag. 
Nat. -rist. (7) 7 xvill p. 120. 

This species in the shape of its antennae is very nearly allied 
to my two new species H. validicornis and annandalei, and might 
be almost included in that group, but is left for the present here 
owing to the similarity of the wing to H. singularis, Ricardo. 


IQII.] G. Ricardo: The Oriental Tabanidae. 339 


Haematopota singularis, @ , Ricardo. 
CPlL xvill,; fig 27.) 


Ann. Mag. Nat. Hist. (8), i, p. 58 (1908). 

Type (female) from Nhatrang, Annam, 22-x-1905 (Dr. Vassal). 

This species I had placed under H. cilifes, Bigot, in my paper 
on Haematopota in Ann. Mag. Nat. Hist. (7), xviii (August 
1906), p. 126; but since that was published I have had the oppor- 
tunity of examining the type of H. cilipes in the Paris Museum, 
and find this is quite a different species, not previously described ; 
itis related to H. rubida, Ricardo, from Burma, in the wings 
and also in the fringed incrassate hind tibiae, but differs from it in 
having all the femora and tibiae fringed, in this resembling H. ci/i- 
pes, Bigot, and H. lata, Ricardo, with which it should be placed in 
the table on p. 114. It is a handsome small black species with 
densely hairy legs, and is immediately noticeable by the pale 
streak across the brown wings, which have no rosettes apparent, 
thus differing with H. rubida considerably from the other described 
species of Haematopota from the Oriental Region. In the shape of 
the antennae it resembles H. rubida, with the long incrassate first 
joint, the very small second joint, and the broad basal division of 
the third joint, the last division being small and short. 

Face grey, the whole upper part deep black, the lower half 
grey, with white hairs ; the beard and under part of head also with 
white hairs. Palft yellow, with short black pubescence and some 
longer white hairs. Frontal callus black, shining, narrow, almost 
reaching the eyes ; forehead brownish, with grey markings surround- 
ing the two black spots and continued across the forehead ; there 
is a tuft of white hairs on each side of the forehead on the outer 
border of the black spot and reaching the frontal callus; on each 
side of the forehead bordering the eyes there is a narrow grey bor- 
der of tomentum. Antennae yellow, in shape as described above, 
with long black pubescence on the first and second joints ; the third 
is bare. Thorax black, with three short grey stripes, the centre 
one shortest, none reaching the posterior border; the sides grey ; 
the posterior border of thorax is fringed with white hairs; the pu- 
bescence on the dorsum black, short, with some white pubescence 
on the stripes and on the sides ; above the root of the wings there 
is a black tuft of hairs, with some long white hairs below. Scutel- 
lum appears yellowish, bordered with a fringe of white pubescence. 
Abdomen deep black, the second, third and fourth segments with 
white segmentations ; the pubescence black, with white hairs on 
the sides of the dorsum of second segment, on the white segmenta- 
tions, and on the sides of the first segment, on the sides of the 
other segments chiefly black; under side the same, but more 
hairy. Legs all with fringes of black hairs, which are least 
thick on the middle femora; they are brown in colour, with two 
yellow rings on the middle and posterior tibiae, and the base of the 
fore tibiae is yellow ; the middle and posterior femora are largely 
yellow ; the fore tibiae and femora with wholly black fringes, long 


340 Records of the Indian Museum. [Von 7EVe 


on each border of the femora;long on the outer border and short 
on the inner border of the tibiae; the middle femora with whitish 
pubescence forming a scanty fringe on their lower borders, with 
some thick black hairs at apex; the middle tibiae with a fringe of 
coarse black hairs on each border and white and black pubescence 
on the dorsum ; hind femora with very thick black and white pu- 
bescence; the hind tibiae broad, flat, with fringes of short black 
hairs on their borders and black and white pubescence on the dor- 
sum; tarsi with short black pubescence ; the middle and posterior 
tarsi are almost wholly yellow. Wings dark brown, the pale streak 
is continuous from the round pale spot above the stigma to beyond 
the apical cell; the pale markings of the apex and the external 
border are so placed that, viewed by the naked eye, a second pale 
streak is seen divided from the first by a brown parallel band, 
with a few brown markings on it, but no rosettes are apparent. 
Length 8 mm. 


(B) Wings with no pale oblique transverse streak but with the usual 
vosettes apparent. 
(a) Apical band of wing double. 


Haematopota javana, ~ 2? , Wiedemann. 
(Plate xvii, fig. 25.) 


Dipt. exot., p. 100(1821); 24., Auss. zweifl. Ins., i, p. 218 (1828) , 
v. d. Wulp, Sumatra Exped. Dipt., 19, pl. i, fig. 12 (1881). 

? Haematopota nigra, Wiedemann, Dipt. exot., p. ror (1821). 

? Haematopota astatica, Rondani, Ann. Mus. Civ. Genova, vii, 
p. 461 (1875). 

Black ; posterior border of thorax, scutellum and segmenta- 
tions of abdomen whitish (male); or grey-brownish. Abdomen 
with a whitish stripe of spots, segmentations and femora a little 
reddish (female) ; wings smoky grey with whitish markings. 4 lines 
o7 9. From Java. 

Very similar to H. pluvialis. 

Male. Antennae ochre-brown; face greyish white, with white 
hairs, a deep black spot on each side between the eyes and the 
antennae; forehead near the antennae with a black spot. Thorax 
black and black-haired, brown at the sides, posterior border and 
scutellum whitish haired. Abdomen at the base slightly ochre- 
brown; the under side almost wholly reddish ochre-brown. Wings 
with a white median band, convex towards the apex, with pale 
brownish spots enclosed in it; behind the stigma with an oblique 
lambdoidal mark, and with several whitish little lines and spots. 
Halteres yellowish with a brown club. Legs alternately banded 
yellow and black. 

Female. Antennae and frontal callus ochre-brown; face grey 
haired, with a deep black spot on each side, another behind the 
antennae, and the usual two eye-spots on the forehead. Thorax 
brownish grey, sides and three lineal stripes whitish. Abdomen 


IQII.] G. Ricarpo: The Oriental Tabanidae. 341 


somewhat deeper brownish, posterior border of the first segment 
more broadly reddish, with a triangular grey-haired spot in the 
middle, round which the reddish colour is conspicuous ; second and 
third segment with a narrow triangular median, and grey-haired 
side spots; the others with only a median spot. Under side grey- 
haired with a very broad blackish median stripe and narrow red- 
dish segmentations. Markings of the wings hardly different from 
those of H. pluvialis. Femora very pale reddish, white haired, 
apex brown above ; hind tibiae with three brown bands ; the fore 
tibiae brown, only whitish at the base. In Westermann’s collec- 
tion. Wied., Auss. zweifl. Ins., i, p. 218. 

Two males caught in May and July at Rawas and Silago 
(Sumatra). ‘The first joint of antennae distinctly thicker than 
the following ones, and very shining black at the end. The fa- 
cettes of the eyes with the exception of those of the lowermost 
third are very large. ‘Tibiae very broad and flat, on each side 
with thick black-brown pubescence, which is longer on the poster- 
ior pair. The markings on the wings are, according to Wiede- 
mann, very distinct ; I will only add that the light band at the 
apex is double and that a large square pale spot is present on the 
posterior part which lies in the middlemost basal cell and traverses 
the middle of the lowest basal cell. V.d. Wulp, Sumatra Exped. 
DUPt.5 P19: 

. The plate given by Wulp shows the apical band double, and 
light markings below the stigma (which is not shown itself). 

Bigot in Bull. Soc. Zool. France, 1891, p. 74, where he gives 
a table of Haematopota species, remarks that he does not include 
H. asiatica (Rondani) in it, as the author attributes it to Wiede- 
mann, but Bigot cannot find in what paper Wiedemann published 
it, neither canI. Probably it is an error of Rondani’s—he gives 
the first two lines of the description of ‘‘H. javana,” from Dipt. 
exot., p. 100, and then adds a few further particulars. V.d. Wulp, 
inhis Cat. Dipt. S. Asia, says ‘‘asiatica” is a lapsus calam. 1 
saw the type in the museum at Genoa ; it was in bad condition, but 
the face appeared to have no spots, so that it is very probable that 
it is not a specimen of H. javana at all. 

The following is the description of H. negra, Wiedemann, given 
as a synonym by Kertesz in his Cat. Dipt., 1906, on what autho- 
rity I do not know, and without an examination of the type it is 
impossible to decide :— 

Black, the posterior margin of thorax, scutellum and segmen- 
tations of abdomen white; wings smoke coloured, with white spots. 
Length 4 lines, from Java. 

Antennae ochraceous, hypostoma greyish white, white haired 
and on each side between the antennae and eyes is a large black 
spot; forehead near antennae with a black spot. Thorax black 
and black haired, sides brown, the posterior border and the scu- 
tellum white haired. Abdomen a little ochraceous at base; the 
under side almost wholly reddish ochraceous. Wings with a median 
dark band, convex behind, enclosing an obsolete white spot, with 


342 Records of the Indian Museum. [V or, IV, 


a transverse lambdoidal design beyond the stigma and with many 
other white lines and spots. Halteres yellowish, the club brown. 
Legs yellow with alternate black bands. Mus. Westermann. Wiede- 
mann, Dipt. exot., p. Ior (1821). 

In Brit. Mus. coll. a series of females from Darien Tipus, 
Federated Malay States (Stanton coll.). 

In London School Tropical Medicine coll. females from Jelebu, 
Negri Sembilan, Federated Malay States (Stanton); and one from 
Federated Malay States (H. C. Pratt). 

In Indian Museum males from Pusa, Calcutta, Purneah, Raj- 
mahal, and Goalbathan, Bengal; females from Rajmahal (at light 
on banks of Ganges), Port Canning, Bhogaon in Purneah district, 
Khoolna and Goalbathan, Bengal. 

In Howlett coll. males and females from Pusa, and female 
from Calcutta ‘‘on cattle.” 

In Kertesz coll. a female from Tushai Hills. 

These females belong to this species I believe, though Wiede- 
mann’s description is wanting in particulars, but the face with the 
deep black spots on each side and the similar spot between the 
antennae, the ochre-brown frontal callus, the yellowish red an- 
tennae, the whitish or greyish markings on base of thorax and on 
scutellum besides the usual stripes, and the pale reddish femora 
white haired, brown at apex, serve to distinguish the species with 
the help of the figure of wing given by v. d. Wulp, to which the 
above specimens correspond. ‘The wing appears to the naked eye 
distinctly marked with brown and white, the brown colouring 
being so disposed as to appear a dark square blotch surrounding 
the stigma with another similar one above, separated from it by a 
pale interval ; the apical band is double, its lower larger half united 
to a pale blotch in the first posterior cell in Wulp’s figure, but in 
these specimens it is usually disconnected. Length of specimens 
93—I10 mm. 

9. face greyish with silvery white pubescence; the large deep 
black spots very distinct between the eyes and the antennae, the 
centre of face just below antennae is also blackish. Palpz pale 
yellow with short black pubescence and some longer white hairs at 
base and on upper side. Antennae yellowish red, the first joint a 
little incrassate after the base, but not broader than the basal 
annulation of third joint, which it about equals in length ; it has 
dense black pubescence, as has also the second joint which is ex- 
tremely small, the third joint devoid of pubescence, the basal 
annulation large, with the three apical annulations very small and 
narrow. forehead brownish grey, the two black paired spots large, 
not touching the frontal callus which is shining yellowish brown, 
barely reaching the eyes, its upper border straight but produced 
in the middle, its lower border concave, receiving the black spot 
between the antennae; no unpaired spot present or only represented 
by a small brown spot. Thorax brownish grey with short pale 
fulvous pubescence on its dorsum, the three grey stripes distinct, 
narrow, the lateral ones ending in a grey spot at median suture of 


TQII.] G. Ricarpo: The Oriental Tabanidae. 343 


thorax, from the base of thorax other lateral stripes proceed and 
the base of thorax is grey, this is only apparent in unrubbed speci- 
mens; sides of thorax grey, pubescence whitish, breast grey with 
whitish pubescence. Scutellum wholly covered with grey tomentum 
and with some short fulvous pubescence. Abdomen not very well 
preserved in any of the specimens, blackish or reddish brown, 
the first two segments usually reddish, the grey stripe apparent 
and the segmentations grey, side spots not always apparent; under 
side appears lighter at least at sides. Legs brown, the femora 
pale reddish or yellowish, white haired below, but the fore femora 
have a fringe of short black hairs; the middle pair has a few at its 
apices, and the hind femora have a tuft of thick black hairs at 
the apex, thickest on the under side, tibiae with two yellow rings, 
the fore tibiae whitish yellowish at base with white hairs, incrassate 
on the dark part with fringes of short black hairs, the middle 
tibiae with black pubescence chiefly on the dark rings, the poster- 
ior tibiae broader with fringes of longer black hairs intermixed 
with white hairs on the yellow rings, tarsi blackish, first joint of 
hind tarsi pale. Wangs dark brown, with the rosettes not very well 
defined, but two fairly distinct ones appear, the apical band is 
double, its upper part narrowest, the lower part concave on its 
upper border, continued when perfect across the second branch of 
third vein joining a white mark in the apex of first posterior cell, 
above the stigma is a large white mark consisting of a round white 
ring with a dark centre, the upper rosette joins it and encloses the 
appendix, the second rosette is nearer the posterior border of wing, 
encircling the upper part of discal cell and bases of second and 
third posterior cells; below this appear a series of short white 
bands reaching the fore border of wing below stigma, and continued 
to the anal cell border, and into the basal cells, representing the 
third rosette but irregular, forming however a very perceptible 
mass of pale colouring, the anal cell has a curved white streak, 
and on the posterior border a pale mark appears in the species of 
most cells. 

#. ‘The specimens from India in Indian Museum are more red- 
dish coloured than the females, the abdomen largely reddish yellow, 
blackish at apex. Thorax scutellum lighter coloured. 

The antennae are wholly reddish yellow, not black at apex of 
first joint as v. d. Wulp states is the case with his specimens, the 
first joint stouter and shorter than in the female. In the wing 
the pale markings in the fourth and fifth posterior segment almost 
fill up the basal halves of these cells as shown in Wulp’s figure 
which is taken from his male specimens. 


(b) Apical band of wing single. 
Haematopota assamensis, ?, 1. Sp. 
(Plate xviii, fig. 29.) 


In British Museum coll. type (female) and others from Nang- 
poh, Khasi Hills, Assam (H. Maxwell Lefroy) ; and from Shillong , 


344 Records of the Indian Museum. [Ver ING 


Assam (H. Maxwell Lefroy). In Howlett coll. females from 
Nangpoh. 

This species is very nearly allied to Haematopota javana, Wied., 
but is at once distinguished from it by the apical band of wing 
which is single, short; starting from the costal border just above 
where the second vein ends, it crosses the anterior branch of 
the third vein, curving downwards, and ends just beyond; it is 
broad at its base, gradually becoming narrower; a pale indistinct 
spot sometimes appears beyond it; the first posterior cell is in some 
specimens more largely filled at its apex with white markings. 
The frontal callus is black, shining, centre of face with no black 
spot. The first joint of antennae not quite so incrassate or so long 
and the third joint not quite so wide. The grey stripes of thorax 
and grey colour of scutellum is here ashy grey, abdomen blackish 
brown with whitish segmentations, only median spots sometimes 
apparent on second and third segments, and side spots not present. 
The hind tibiae are fringed with black hairs. Length 10 mm. 


Haematopota lata, 2 , Ricardo 
(Plate xvii, fig. 26.) 


Ann. Mag. Nat. Hist. (7), xvili, p. 121 (1906). 

Type (female) from Khasi Hills district, India (Chennell), 1878 ; 
five females from N. Chin Hills, Burma, iv-93 (Watson). 

A dark brown broad-bodied species distinguished by the broad 
hind tibiae fringed with black hairs. 

Face grey, with some brown marks below the antennae, the 
stripe between the antennae and the eyes yellow, with some brown 
dots ; the beard white : the palpi yellow, with white pubescence and 
a few black hairs intermixed. Frontal callus brown, shining, nar- 
row, reaching the eyes; the anterior border irregular with yellow 
edges ; the posterior border nearly straight ; the paired spots black, 
large, touching the eyes; the unpaired spot small, bordered 
with yellow; forehead grey, yellower on the vertex with short 
black pubescence. Antennae very similar to those of H. rubida, 
long, robust, yellowish red, darker at the tips with black pubes- 
cence ; the first joint shorter than the third, stout, incrassate, the 
second very small, the third broad, ending in an obtuse point. 
Thorax brown, with a narrow, indistinct, grey median stripe con- 
tinued to the scutellum; the side stripes grey, broader, ending 
at the suture in triangular spots; the shoulders and _ posterior 
border of the thorax grey; a short grey stripe from each side of 
the base of the thorax running up to the suture outside the usual 
side stripes ; the sides and breast grey, the latter with white hairs, 
traces of silvery white pubescence on the dorsum, with some black 
pubescence. Scutellum brown, grey on its anterior border. Ab- 
domen brown, with distinct greyish white borders to the segments : 
the sides of the first four segments grey ; from the fourth segment 
large grey spots are apparent on each side; a grey median stripe 


IgII. | G. Ricarpo: The Oriental Tabanidae. 345 


is here indistinct, but apparent on the other specimens ; the pubes- 
cence brown, with some white hairs, especially at the sides; under 
side grey. Legs yellowish, the fore tibiae white at base, dark 
brown on the apical half; the fore tarsi brown, the middle and pos- 
terior pale at base; the anterior and middle femora with whitish 
pubescence, the hind ones with a fringe of black hairs; the tibiae 
yellowish, with a brown ring in the middle and brown at base, thus 
appearing as yellow rings on the tibiae; the hind tibiae with a 
heavy fringe of black hairs extending two-thirds of the length to 
the apex. Wangs greyish, with yellow veins and stigma and a long 
appendix ; the apical band single, divided in half, the two upper 
rosettes distinct ; in the corner of the wing is a round circle, and 
above this a double concave circle extending into the anal cell ; 
above the stigma is a small round circle ; the basal half of the first 
posterior cell is wholly pale; only the upper part of the third 
rosette is distinct. Length 9 mm. 

In the females from Burma the wings are browner and the 
white markings more numerous at the opening of the cells. Ricardo, 
Ann. Mag. Nat. Hist. (7), xviii, p. 121 (1906). 

This species is distinguished from H. assamensis,n. sp., by the 
shape of the first joint of antennae which is much stouter, and as 
long as the first annulation of the third joint, whereas in H. assam- 
ensis, it is shorter and not muchincrassate. The face has hardly 
a trace of black spots and there is no spot between the antennae. 

The apical band in the type does not reach across the wing, 
but extends only a short way beyond the anterior fork of third 
vein, in other specimens it reaches across but is broken in the mid- 
dle, and in others it is broader, extending the whole width of apex 
of wing. 


Haematopota sinensis, ? , 1. sp. 
(Pi xvii, tig. 28.) 


In Brit. Mus. coll. type (female) and others from Wei Hai 
Wei, China (Dr. W. M. Muat), 1907; and two females from 
Shanghai (Walker coll.), 1892. 

A species distinctly allied tothe Oriental species, distinguished 
by the short antennae with the disk-like broad third joint, 
on the upper and lower border of which at base appears a percep- 
tible fringe of black hairs, an unusual characteristic ; it is allied 
to H. assamensis, n. sp., from which it may be distinguished by the 
paler coloured wings and by the much broader third antennal 
joint; from it and from H. lata, Ricardo, it is also distinguished 
by the hind tibiae not being heavily fringed and by the shorter 
first antennal joint. 

The face has no distinct band, the frontal callus is triangular 
in shape, the thorax distinctly striped, and the abdomen with 
lateral spots and a medianstripe. Wings with a single apical band 
and the fifth posterior cell largely pale on its apical half. Length of 
type 9 mm., others from 9—11$ mm. 


340 Records of the Indian Museum. [Vou. IV, 


Face greyish with white pubescence, a dark brown spot is 
present usually, between the antennae and the eyes, but in the 
series of specimensis often absent or indistinct, the palpi long and 
slender, pale yellow with black pubescence. Antennae reddish 
yellow, short, the first joint short, stout, not so long as the first 
annulation of the third joint, the second joint very small, with its 
upper and lower borders produced to a narrow process crowned 
with black hairs, both joints with black pubescence, the third 
joint with the first annulation remarkably broad, flattened out, 
on its upper border with a distinct angle about half-way up and 
distinct black hairs on the upper side nearly the whole length 
(these are denuded to a great extent in the type), they are also 
visible on the lower border, the three last annulations almost bi- 
equal, forming a short apex, some black hairs also visible on them ; 
between the antennae a dull brown spot is visible. Forehead 
greyish with some black markings. Frontal callus triangular in 
shape, its lower margins touching eyes, shining, brown; the 
paired spots large, black, not reaching the frontal callus and only 
touching the eyes at one point in the middle of their outer border, 
the unpaired spot small, round. Thorax brown with three distinct 
grey stripes, the outer ones interrupted in the middle, and the 
sides also grey, breast brown with grey tomentum. Scutellum 
covered with grey tomentum. Abdomen brown, same shade as 
thorax, with a narrow median grey tomentose stripe most distinct 
on the first three or four segments and with round grey tomentose 
lateral spots, pubescence whitish, short, but when not denuded 
fairly abundant ; under side brown with grey tomentum and pu- 
bescence. Legs pale reddish yellow, the femora with some grey 
tomentum, the fore tibiae brownish, hardly incrassate, pale at base, 
the middle and posterior tibiae brown with the usual pale 
rings, the hind tibiae with no distinct heavy fringes of hair, the 
fore tarsi brown, paler at apices, middle and posterior tarsi 
more largely pale, the pubescence on femora and pale parts of legs 
whitish, otherwise black. Wings pale brown, the stigma darker, 
yellowish brown, veins brown, three rosettes distinct, the apical 
band single, lineal, starting from a pale blotch at apex of the 
second vein and reaching across apex of wing in a sinuous course to 
the posterior branch of third vein, the first four posterior cells with 
small pale markings at their apices, the fifth with a large pale 
irregular-shaped blotch filling up the apex, occupying more than a 
third of the whole space of cell, three more pale bands are visible 
in the cell towards the base, the small transverse bands at inner 
border distinct and regular, in axillary angle the curved pale line 
is V-shaped. 

Group VI. 


Third joint of antennae not very broad at base, the first joint 
short, stout or cylindrical, often only half as long as the third joint 
or at least never longer than third joint which is usually long and 
slender. 


IgII.] G. Ricarpo: The Oriental Tabanidae. 347 


(A) Apical band of wing broad, double, almost filling up the apex 
of wing. 


Haematopota marginata, o ?,n. sp. 
(Plate xviii, fig. 32.) - 


In Howlett coll. type (female) and another from Pusa, Bengal, 
and another from Tezpore, Assam. 

In Indian Museum type (male) and others from Goalbathan, 
E. Bengal ; and females from same locality. 

A species distinguished by the wide double apical band which 
almost reaches the extreme apex of wing, and continues round the 
posterior border reaching the second branch of fork of third vein, a 
pale band continues from it unbroken round the posterior border 
to the end of wing. The female is blackish brown with a grey 
median stripe and spots, the frontal callus very narrow, the paired 
spots large, a black spot present between eyes but no spots or 
band on face. Antennae reddish yellow, the first joint cylindrical, 
curved. Male with abdomen yellow at base, blackish at apex, legs 
pale yellowish, no rings apparent, the first joint of antennae con- 
siderably incrassate. Length male, type 8 mm., others 7$—8 mm. ; 
female, type 74 mm., others 7—9 mm. 

9. Face grey with hardly any white pubescence. Palfi red- 
dish, covered with grey tomentum, and with black pubescence. 
Antennae slender, reddish yellow, apex black, the first joint not so 
long as the first annulation of the third joint, the second small, 
roundish, about one-fourth the length of the first joint, both with 
some black pubescence, the third joint long slender, only slightly 
broader at base. Forehead blackish brown with lighter circles 
round the spots and some grey tomentose marks, the frontal callus 
very narrow, shining, blackish brown, its upper border somewhat 
curved, on its inner border concave, the band at ends is broader 
and produced on its inner border forming almost a half circle ; a 
small black spot is present between the antennae; paired spots 
large, black, oval, not quite reaching the eyes or the frontal callus. 
Thorax blackish brown, with three narrow grey stripes ; scutellum 
same colour. Abdomen blackish brown with indistinct grey to- 
mentose median stripe and grey tomentose lateral spots ; under side 
darker. Legs reddish brown, the pale rings of tibiae yellowish, 
not always very distinct, fore tibiae at base yellowish, very slightly 
incrassate, posterior tibiae not incrassate, with some black pubes- 
cence. Wings brown, the pale markings very numerous, three 
rosettes distinct, the small transverse bars on posterior border are 
zigzag and almost touch the pale border, stigma very distinct, 
dark brown, two zigzag pale stripes cross the axillary angle of 
wing, the lower branch of the apical band is very narrow and not 
always continued to end of the upper broad part of band. 

@. Eyes with the lower third composed of small facets. An- 
tewnae yellowish, the first joint very stout, almost as long as the first 
annulation of third joint, the second very small, the third joint 


348 Records of the Indian Museum. [Von. IV, 


slender, only slightly wider at base, the first two joints with black 
pubescence ; the black spot between the antennae present and a 
small shining yellow frontal callus. Abdomen with the first four 
segments reddish yellow, the posterior border of fourth and other 
segments blackish. Legs rather paler than in female, and the ring 
of tibiae not always apparent. Wungs identical in design, the 
small transverse bars smaller, reduced to spots. 

This species is allied in the wing to H. pallens, Loew, Neue Bes- 
chreib. Europ. Dipt., ii, p. 61 (1871). The six specimens of Loew’s 
species in the Brit. Mus. coll. recorded by me in the ‘‘ Annals” (7), 
Xviii, p. 96 (1906), are females, not males as erroneously printed ; the 
Indian females of my species have the antennae reddish and no 
band on the face, whereas in the Algerian specimens the antennae 
are blackish, only the third joint being pale yellow, black at apex 
and there is a black band on the face broken up into spots. The 
Indian males have this band but the first joint of antennae is 
wholly yellow, incrassate but less so and shorter than in H. pallens. 
The spot between the antennae in both males and females of my 
species is small and brown, in the Algerian females it is large and 
black. 


(B) Apical band of wing double, but not almost filling up the apex 
of wing. 
(a) Both branches of apical band joined at their base, diver- 
ging later. 


Haematopota tessellata, 2 , Ricardo. 
(Plate xvi, fig. 14.) 


Ann. Mag. Nat. Hist. (7), xviii, p. 120 (1906). 

Type (female), Hot Wells, Trincomalee, 8-xi-g1 (Yerbury), 
g2, 192. 

A grey species with brown wings, the white markings very 
clearly defined, so that the wings have a chequered appearance; 
the middle and hind tibiae with rings. 

Face grey, only a trace of a brown stripe between the antennae 
and the eyes. Frontal callus black, short, not reaching the eyes, 
very much produced on the posterior border, ending in a point; 
the anterior border almost straight; the paired spots large, black, 
not reaching the eyes; the forehead is apparently grey, and no 
paired spot visible. Antennae yellow; the third joint dusky, the 
first joint incrassate, rather shorter than the third, the second 
very short, the third long, tapering to a point. Thorax brown, 
with three grey stripes; the shoulders, base, and sides of thorax 
grey; a short white pubescence on the dorsum; the breast grey. 
Abdomen brown, with rather wide white borders to the segments, 
grey spots on the posterior segments, and a faint grey median 
stripe. Legs brown, the femora lighter, the middle and posteriot 
tibiae with rings, the basal joint of the tarsi of the middle and 


IQII. ] G. Ricarpo: The Oriental Tabanidae. 349 


posterior legs whitish. Wings brown, with yellowish brown veins 
and an appendix; the apical band double, the openings of the 
posterior cells light coloured. 

Length 84 mm. 

A male from Velverry, Ceylon, 26-x-91 (Yerbury), 92, 192, in 
poor preservation, is probably the male of this species. Ricardo, 
Ann. Mag. Nat. Hist. (7), xviii, p. 120. 

The apical band of wing is double, both branches united at 
base, diverging later. The first joint of the antennae is as long 
as the first annulation of the third joint. 


Haematopota brevis, ¢ , Ricardo. 
(Plate xvi, fig. 16.) 


Ann. Mag. Nat. Hist. (7), xvili, p. 122 (1906). 

Type (female), Kanthalla, Ceylon, 19-x-g0 (Yerbury), 92, 192, 
and another female; one female from Velverry, Ceylon, 18-i-91 
(Yerbury), 92, 192; two females from Bangalore, Mysore, June 24 
(Watson), 95, 28, and type (male) from the same locality. 

This species is distinguished from H. cingalensis by the short 
first joint of the antennae, and from H. javana, Wied., by the 
blackish brown abdomen with white incisions and a mere trace of 
a grey stripe. 

A brown species, distinguished by its short first antennal joint 
and by the rings on the middle and posterior tibiae. 

Face grey, with brown pubescence; a dark spot under each 
antenna and some darker colour continued to the mouth; the 
stripe on each side yellowish, with brown dots and spots; the 
beard brown; the palpi yellow, with black pubescence and white 
hairs below. Frontal callus yellow-brown, in some specimens 
darker in the middle, short, broad, not reaching the eyes, the 
anterior border nearly straight, the posterior convex; the spot 
between the antennae small, ill-defined, brown. Forehead yellowish 
brown, darker on the vertex; the paired spots black, triangular, 
the apices touching the frontal callus. Antennae yellowish, densely 
covered with grey tomentum, at the apices black; the first joint 
short, robust, with black pubescence, the second small, with black 
hairs, the third slender, twice as long as the first. Thorax brown, 
with short silvery white tomentum, and three grey stripes all 
ending in a grey spot, but the median one the longest; the base 
of thorax and shoulders grey; scutellum brown, grey on the 
posterior border, pubescence at the sides brown; breast reddish 
brown. Abdomen the same colour as the thorax, the borders of 
the segments whitish, the sides of the first two segments grey; 
there is a trace of a grey stripe on the second segment only. 
Legs brown-black, the base of the tibiae white; the first joint of 
the fore tarsi nearly as long as the four remaining joints, which 
are broad and short; the middle and posterior tibiae reddish 
brown, with yellow rings; the base of the middle and posterior 


350 Records of the Indian Museum. [ VOL; EV; 


first joint of the tarsi yellow. Wings brownish, the veins 
brownish, with an appendix; the apical band double; the openings 
of the posterior cells mostly light coloured. 

Length 9 mm. 

Type (male) differs in the third joint of the antennae being 

broader and shorter, the forehead grey, with a large oblong brown 
spot. The abdomen is redder brown, with a distinct grey stripe; 
the scutellum wholly grey, the fore tarsi not so broad, the wings 
and the base of the tarsi more white than yellow, the apical line 
of the wing broader. Ricardo, Ann. Mag. Nat. Hist. (7), xviii, 
Dieta 2. 
, The short first joint of antennae—shorter than the first 
annulation of the third joint—and the triangular shape of the 
yellowish frontal callus distinguishes this species from H. tessellata, 
Ricardo. 


(b) Both branches of apical band separate, starting from op- 
posite borders of wing and usually overlapping in the 
middle. 


Haematopota bilineata, 2, n. sp. 
(Plate xviii, fig. 34.) 


_ In Brit. Mus. coll. type (female) and another. In Indian 
Museum females, all from Igatpuri, Bombay, 2,000 ft. (H. 
Maxwell Lefroy). 

A species distinguished by the narrow apical band of wing 
composed of two short lineal bands, each starting from opposite 
sides and overlapping at ends, the lower branch often broken up 
into spots, the wing is dark brown, with the pale markings distinct 
chiefly as small round spots, four rosettes are clearly outlined. 
The first joint of antennae is considerably incrassate. Face with 
a band and lower spot. Frontal callus narrow. Paired spots 
large, touching eyes and frontal callus. Abdomen brown with 
grey segmentations. Length type 7 mm., others 8—9} mm. _ Dis- 
tinguished from H. cingalensis, Ricardo, by the shorter incrassate 
first antennal joint. 

Face grey, the band between antennae and eyes dark brown, 
in some specimens partially broken up into spots, an oblong 
brown spot is present below. Palpi pale reddish with black pubes- 
cence. Antennae dusky reddish, the first two joints shining black, 
the first joint incrassate, as long as first annulation of the third 
joint, the second very small, cup-shaped, both with black pubes- 
cence, the third slender, very little wider at base, and the apex 
not very narrow. Frontal callus shining, blackish brown, narrow, 
its upper border straight, the lower one concave in centre, a black 
spot apparent between the antennae. Paired spots large, black, 
oblong, their upper apices touching frontal callus, and their base 
reaching eyes, unpaired spot black, small. Thorax brownish 
with short yellow pubescence, stripes not apparent (denuded). 


IgII.] G. Ricarpo: The Oriental Tabanidae. 351 


Scutellum similar. Abdomen brown with distinct grey segmenta- 
tions. Legs blackish brown, the fore tibiae at base, and the other 
tibiae with two rings, yellowish, the base of posterior tarsi paler. 
Wings dark brown, with apical band as described, posterior cells 
with small pale markings in their apices, the transverse small bands 
small, chiefly appearing as spots, the stigma darker brown than the 
colouring of wing. 


Haematopota cingalensis, ? , Ricardo. 
(Plate xvi, fig. 15.) 
Ann. Mag. Nat. Hist. (7), xviii, p. 119 (1906). 


Type (female), 19th milestone, Candy Road, 1890 (Yerbury) ; 
two females from Tamblegum, 1890 (Yerbury); and a serise 
of females from Anaradhupura, Ceylon; 1899 (Oliver Bartholo- 
mew). 

There is a note with Col. Yerbury’s specimens to the effect 
that the species is common on the road near Tamblegum in Octo- 
ber and November. 

This species is distinguished from H. cordigera, Bigot, by the 
paired spots not coalescing, by the longer first joint of the an- 
tennae, and by the fore legs being lighter in colour. 

A dull greyish brown species. 

Face grey; an oblong black spot under the antennae, and a 
black stripe reaching from the eyes half-way across to the lower 
edge of this spot, the colour above the spot being yellowish. 
Frontal callus pitchy brown, shining, with a concave fore border, 
from which a black spot proceeds to between the antennae; the 
posterior border is produced in the middle; the paired spots are 
black, large, almost toughing the eyes; the unpaired spot ap- 
parent, sometimes indistinct ; forehead yellowish, grey at the sides. 
Antennae yellow ; the third joint darker, its last three annulations 
dull black; the first joint curved, not quite so long as the third, 
the second joint very small. Thorax brown, with three well-marked 
grey stripes and four grey spots on its posterior border, the side ones 
ending at the suture in a spot; there are traces of short white 
pubescence on the dorsum; the breast is grey, with some white 
hairs. Abdomen light mahogany-brown or brownish, with light 
yellow borders to the segments and a well-marked grey stripe 
starting from the second segment; there are also distinct oblong 
grey spots on each side, beginning from the third segment ; there 
is some short white pubescence, thickest on the yellow borders of 
the segments; the under side with grey tomentum. Legs reddish 
yellow, the fore tibiae white at base, or rather, with one narrow 
white ring on the basal half; the apical half black; the middle 
and hind tibiae with well-marked typical whitish rings. Wings 
greyish, with a yellowish brown stigma and yellow veins and an 
appendix ; the rosettes and markings are distinct ; the apical band 
in the type and other specimens is double, but in others the double 


5 


352 Records of the Indian Museum. [Vor.2Vs 


branch is only represented by two small spots, as shown in the 
photograph. Length 8 mm. Ricardo, Ann. Mag. Nat. Hist. (7), 
Vii, Pp. 119. 

The first joint of the antennae is long and cylindrical, nearly 
as long as the third joint. Frontal callus narrow. 


(C) Apical band of wing single, formed of spots. 
Haematopota irrorata, 2 , Macquart. 
(Plate xviii, fig. 30.) 


Dipt. exot., i (1);-p. 167, pl. xix, fis:-3 (1838); V.d. Wulp; 
Sumatra Exped. Dipt., 19, pl. i, fig. 13 (1881) ; Ost. Sacken, Ann. 
Mus. Civ. Genova, xvi, p. 419 (1882); Bigot, Bull. Soc. Zool. 
France, xvi, p. 75 (1891). 

? Haematopota pungens, Doleschall, Naturkand. Tijd. Nederl. 
Ind. x, p. 407, pl.-iv, fig. 3-(1856). 

Red, Antennae with the first joint elongated, cylindrical. 
Legs red, tibiae with white rings. Wings red with white spots 
(ole 19, fie.-3)) Vength 4 lines: os 

Palpi fawn coloured. Face of a yellowish grey colour. Fore- 
head greyish brown, base shining black ; an oval oblique dull black 
spot on each side of forehead, near the eyes and contiguous to 
the black spot at base. Antennae: the first two joints fawn 
coloured; the first one long cylindrical, the third joint wanting. 
Thorax fawn coloured; sides with scanty grey tomentum. Abdo- 
men fawn coloured. Legs fawn coloured; tarsi brown; anterior 
tibiae with a whitish ring at base ; intermediate and posterior tibiae 
with two yellowish white rings and apex brownish. Halteres 
brown. Wings a rather pale reddish brown, with scattered white 
spots; a whitish circle towards the middle of the marginal cell. 
From Java Museum. Macq., Dipt. exot., i (1), p. 167. 

The ring in the figure of wing has a round white circle on 
fore border near stigma and another in the axillary lobe of wing, 
the apical band is only represented by small white spots. The 
type seen by me in Paris Museum in 1906 is almost destroyed, 
only the wings remaining. 

Nine females from Grabak, Alahan and Lebong (Sumatra). 

The specimen caught at Grabak corresponds the most to 
Macquart’s description and figure, owing to its rusty colouring. 
All the others are darker, but cannot be specifically divided from 
it, as they do not differ in any important characters. Even the 
above-mentioned specimen from Grabak has the last segment of 
abdomen black-brown, always with pale red-yellow segmentations 
which indeed are also apparent on the anterior part of the abdo- 
men. ‘The figure furnished by Macquart does not give a very 
exact notion of the insect ; the dark, instead of pale, spots on the 
wings cause the design to be unlike what it is in reality. The 
third antennal joint (which is missing in the specimen described 
by Macquart) is one and a half times as long as the first one, and 


IQgII.| G. Ricarpo: The Oriental Tabanidae. 353 


always black ; the short second joint the same and even the first 
one at least at the end is darker and very shining, in most speci- 
mens wholly black-brown. ‘The pubescence on the face and chin 
is whitish. The fore legs are darker than the posterior ones, even 
in the palest specimens, mostly black, so that the white base of 
the tibiae stands out distinctly. The species can be well identified 
by the wings, which are almost wholly of a dark colour, the pale 
colouring being for the most part confined to small round points, 
which are not present on the border of wing at all; an oblique 
series of these points runs through the cubital and basal cells, 
and the pale band which is present at the apex of wing in other 
species is here hardly outlined by a pair of spots on both sides 
of the upper branch of third vein ; the second basal cell is inter- 
sected by a pale band, a continuation of pale spots in the first 
basal cell, and this band continues strongly curved into the pos- 
terior angle of wing. It is not unlikely that Doleschall had a 
specimen of this species before him when he described H. pungens 
but his remarks are too short to make it possible to decide this 
point ; of the wings he only says that they have white spots and 
points, and his figure bears witness to the contrary, because it 
delineates the rows of points as distinct stripes. As I have al- 
ready remarked, not much value can be attached to them; how- 
ever they are, on the whole, better than those of Macquart. V. d. 
Wulp, Sumatra Exped. Diptera, pl. i, fig. 13, p. 19. 

Four females. Mt. Singalang, Sumatra, July 1878 (Beccari). 
I determine this species in agreement with Mr. V. d. Wulp who 
showed me three closely-allied species from Sumatra, but which 
could easily be distinguished by the arrangement of the spots at 
the tip ofthe wing. Macquart’s description shows slight discrepan- 
cies. H. pungens, Dol. (Java), may be the same species, but the 
description is too incomplete. Osten Sacken, Ann. Mus. Civ. 
Genova, Xvi, p. 419. 

The following is a translation of the description of Haemato- 
pota pungens, by Doleschall :— 

Red-brown, the abdominal segments with paler segmenta- 
tions; eyes brassy brown, wings dark with white spots and 
points, legs with indistinct white lines. Length 4 lines. 

Numerous in the woods of the mountain Oenarang, and very 
troublesome to horses in the rainy season. 

The figure given by Doleschall is useless as a means of identti- 
fication. 

In the British Museum are two females, one from Negri 
Sembilan, Malay Peninsula (H. V. Ridley), 1900, and the other from 
Kobele, N. Borneo (D. Cater), 1895, which I believe belong to this 
species judging from the wings and the rusty or reddish yellow 
colour of abdomen, darker at the apex. The wing as figured by 
V. d. Wulp does not show the round pale circle above the stigma 
or in the axillary lobe of wing, present in Macquart’s type 
and figured by him ; in these specimens the one above the stigma 
is present, though not forming a perfect circle, the second one is 


354 Records of the Indian Museum. [VoL. IV, 


absent; the two spots forming the apical band and the small 
transverse bars usually formed of two spots continued round the 
posterior border of wing, and the absence of any pale markings in 
the apices of cell on posterior border of wing distinguish this 
species from others; two rosettes are fairly distinct. The frontal 
callus is broad, triangular, being produced on its upper border, 
the paired spots touch the eyes but not the frontal callus, an 
indistinct reddish spot appears between the antennae. The first 
joint of antennae is yellowish cylindrical, not quite so long as the 
first annulation of the third joint, the second joint yellow, the 
third is slender, dusky reddish, but black at apex. Legs: in one 
specimen the fore legs are blackish, in the other the same colour 
as the remaining ones, v7z., reddish, white at the base in both 
specimens. 


(D) Apical band of wing single, broad, band-lhke, reaching from 
outer to inner border of wing. 


Haematopota Iunulata, ? , Macquart. 


Dipt. exot. Suppl. iii, p. 175, pl. 1, fig. 6 (1847); V. d. Wulp, 
Sumatra Exped. Dipt.. 20, pl. i, fig. 14 (1881); Ost. Sacken, Berlin, 
Ent. Zeit., xxvi, p. 97 (1882). 

Black. Antennae red at the base. Legs red, tibiae whitish. 
Wings brown, spotted with white, with a lunate spot (pl. 1, fig. 6). 
Length 4 lines. @. 

Palpi brown with white tomentum. Face with white tomen- 
tum. Forehead black with scanty grey tomentum ; anterior callus 
black; a small dull black spot on each side at border of eyes and 
contiguous to the callus. Antennae: first joint fawn coloured, 
cylindrical, a little elongated, second and third black; third joint 
once and a half as long as the first. Thorax with scanty greyish 
tomentum. Abdomen dull brownish black; a little whitish at the 
segmentations ; second segment with a dorsal triangular spot of 
white tomentum. Femora of a fawn colour inclining to brown; 
anterior tibiae incrassate, a little convex on the outer border, an- 
terior half white; posterior half black; anterior and posterior 
white with black apex (sic) ; tarsi black, first joint of anterior pair 
fawn coloured. Wings with white spots and a lunate shaped white 
one atapex. From Java, from M. Payen. Macquart, Dipt. exot. 
Suppl. iii, p. 175. 

One male from Moeara Laboe, and one female from Alahan. 

Both agree so nearly with Macquart’s description as regards 
the design of the wing that I do not hesitate to ascribe them to 
this species ; a male from Silago probably belongs here but is not 
in good condition and is set in such a manner that it cannot be 
distinctly identified. The male from Moeara Laboe is 8 mm. long ; 
the antennae are wholly black or black-brown; the first joint is 
little stouter than the following ones, and very shining; the eyes 


IQII.] G. Ricardo: The Oriental Tabanidae. 355 
g 


have tolerably large facets above, below very small ones, the small 
triangular forehead is almost wholly occupied by the shining black 
frontal callus. 

The legs are very dark brown, the fore legs black, except the 
basal third of the tibiae which is white ; on the middle tibiae the 
white colour extends further, almost three-fourths of the length ; 
the hind tibiae are distinctly incrassate, more red-yellow and 
only white at the base, on both sides with thick black-brown 
fringes of hairs. 


The female is somewhat larger and has more wholly black 
antennae contrary to Macquart’s account, who described the 
female only and called the first antennal joint red-yellow. The 
legs are alSo distinctly darker than Macquart’s description would 
lead one to believe. The facets of the eyes are wholly very small. 
The ground colour of the wings is darker than that of the two 
preceding species, especially on the anterior part ; the pale colour- 
ing consists of a large quantity of spots which are spread over 
almost the whole wing, but are absent at the apex, being there 
replaced by a very broad transverse band which is convex on its 
discal border, commencing at the apex of the second vein 
(radial vein), crossing the upper branch of the third vein and 
across the apex of wing, ending in the posterior border of wing 
(this is probably the moon-shaped form which was the cause 
of the specific name); the pale band intersecting the second 
basal cell ends in the posterior angle of wing, curving like a hook 
and forming with a pale spot situated on the anal vein a more or 
less eye-like form. V.d. Wulp, Sumatra Exped. Diptera, p. 20, 
pl. i, fig. FA; 

Osten Sacken places the following remarks under the heading 

of H. lunulata, Macq. :— 

esSpeie 

All the specimens before me have the design on the wings like 
the above-quoted figure of v. d. Wulp, but some of them have the 
antennae reddish, except the latter part of the third joint which is 
black ; and at the same time two whitish rings on the hind tibiae ; 
the other group of specimens has altogether black antennae and 
only one white ring near base of hind tibiae. Neither of them 
agrees entirely with Macquart’s description. Osten Sacken, Berlin. 
Ents Zett:, xvi, p. 97. 

Type in the Paris Museumseen by mein 1906. A smali species. 
Frontal callus black, convex, reaching eyes, and the two usual 
black spots on forehead present. Legs yellowish with brownrings. 
Macquart’s figure shows the apical band (his ‘‘ lunule blanche ’’) 
single, crescent-shaped, the same width throughout: v. d. Wulp 
figures it as broader and more irregular; in the apices of cells on 
posterior border he shows small pale markings, the third posterior 
cell appears paler with many white spots, the fourth and fifth and 
axillary angle of wing largely paler, only the first rosette at all 
distinct. Macquart’s figure is useless except as regards the 


356 Records of the Indian Museum. [Vers lve 


drawing of the apical band. I have not seen any specimens of 
this species in the collections to which I have had access. 


Haematopota latifascia, ? , n. sp. 
(Plate xvi, fig. 10.) 


Type (¢) in Brit. Mus. coll. from Shillong, Assam (H. Max- 
well Lefroy). 
In Howlett coll. a female from the same locality. 


A species distinguished by the broad band-like apical band of 
wing extending across the whole breadth of apex of wing, by the 
long stout first antennal joint, fully as long as the first annulation 
of the third joint, by the black band of face and the narrow frontal! 
callus. It differs from H. /wnulata, which it approaches in the shape 
of apical band, by the greater length of first antennal joint and by 
the presence of the black band on face. Length 9 mm. 


Face covered with greyish tomentum, more yellowish brown in 
centre, the black band very distinct, reaching across from border 
of one eye to the other and extending more than half-way down 
the cheeks ; pubescence on face white. Palpi fawn coloured, slen- 
der, with black pubescence and some grey tomentum. <Aniennae 
reddish yellow, the first joint stout, the same width throughout, 
the second one very small, both with black pubescence, the third 
joint dusky at apex, the first annulation moderately broad, the 
last three narrower; spot between antennae present, blackish. 
Forehead brownish, with grey circular markings, the pubescence 
black; the frontal callus blackish brown, narrow, almost straight 
on both borders, reaching the eyes, the paired spots large, blackish, 
almost touching it and contiguous to eyes at one point, unpaired 
spot present. Thorax reddish brown, with some reddish yellow 
short pubescence, stripes hardly apparent, sides greyish. Scutellum 
similar to thorax. Abdomen brown with greyish white segmenta- 
tions and some greyish white hairs at sides; under side brown. 
Legs blackish brown, the base of fore tibiae, the rings on middle 
tibiae, the base and ring of posterior tibiae yellowish white, the 
basal joint of middle and posterior tarsi also pale coloured, the fore 
tibiae are slightly incrassate, the hind tibiae broad with thick 
black fringe of hairs on borders, thickest on the outer one. 
Wings brown, with three rosettes clearly outlined, the broad 
apical band starts from the junction of the second vein at border 
and crosses in an almost straight line to the inner border, which, 
commencing from the first posterior cell, has a continuous pale 
edging to the axillary angle; this is not quite so continuous in the 
second female; the small transverse bars between this and the 
rosettes are distinct, each usually taking up the whole breadth of 
the cell in which it is situated ; the white spot above the stigma 
is conspicuous either as a round white circle or a plain spot; 
veins and stigma brown. 


IQII. | G. Rrcarpo: The Oriental Tabanidae. 357 


(E) Apical band of wing single, narrow, not band-ltke, reaching from 
outer to inner border of wing. 


Haematopota fuscifrons, Austen. 
(Plate xvii, fig. 17.) 
Ann. Mag. Nat. Hist. (8), i, p. 41z (1908). 
Haematopota cordigera @ , Bigot, Bull. Soc. Zool. France, xvi, 


p.- 76 (1891); 7zd., Mem. Soc. Zool. France, v, p. 626 (1892) ; 
Ricardo, Ann. Mag. Nat. Hist. (7), xviii, p. 125 (1906). 


Antennae yellow, the first joint cylindrical, the third not at 
all swollen, blackish toward the apex, about twice as long as the 
axis of head: palpi whitish, proboscis brown, face and beard 
white ; a large wide shining black spot above the antennae; and 
beyond this on the pale ashy grey forehead a large dull black 
cordiform spot, vertex brownish ; thorax dull reddish with three 
greyish stripes, sides with white tomentum, scutellum chestnut 
coloured, greyish in the middle; squamae greyish, halteres pale 
yellow; abdomen blackish, and little shining, all the segments 
narrowly bordered with white ; femora yellow, the base and apex 
brownish, the anterior pair blackish, anterior tibiae black with 
the base white and the tarsi black, intermediate and posterior 
tibiae with the base, apex and median ring black, white above the 
latter and reddish below, their tarsi black, yellow at the base; wings 
grey, with numerous whitish spots dispersed in irregular series, 
and at the apex a transverse concave straight simple whitish band. 

Lengthgmm. Bengal, one specimen. Bigot, Mem. Soc. Zool. 
France, v, p. 626. 

Type from India. 

This species is easily distinguished by the colouring of the 
forehead, by the cylindrical antennae, and by the rings on the 
middle and posterior tibiae. 

Face grey, no spots except faint traces of two below the 
antennae. Palpi yellow with black pubescence; beard white. 
Frontal callus dark brown, shining, reaching the eyes, broad, 
bordered in front by a shining yellow narrow band, which is 
continued between the antennae instead of the usual black 
spot. Forehead grey, with a large brown-black heart-shaped 
spot in place of the usual paired spots; it joins the posterior 
border of the frontal callus and almost reaches the eyes at the 
sides; becoming narrower on its posterior half it is continued to 
the vertex as a broad stripe. Awtennae yellow, long, slender, but 
not so long as those of the preceding species (H. indiana, Bigot), 
the first joint being not much more than half as long as the third, 
the second joint short, both with black pubescence ; the third with 
the extreme apex dusky. Thorax brown, with grey shoulders 
and grey stripes, ending in spots at the suture ; grey spots at the 
base ; the sides are also grey, as is the breast which has white 
hairs. Scutellwm brown, grey inthe centre. Abdomen dark brown, 
with distinct white borders to the segments, but no sign of stripe 


358 Records of the Indian Museum. [Vor. IV, 


or spots; the underside brown, with grey tomentum. Legs yellow- 
ish brown ; the fore tibiae white at base, the middle and posterior 
tibiae with the typical rings, and the basal joint of the tarsi 
whitish. Wangs pale brown, with brown veins and an appendix: 
the apical bandsingle, broad and curved. Length 8mm. Ricardo, 
Ann. Mag. Nat. Hist. (7), xviii, p. 125 (1906). 


Haematopota fasciata, 2, n. sp. 


Type (female) from Howlett collection, Shillong, Assam, and 
another in Howlett collection from same locality. 

A species allied to H. latifascia, n. sp., but distinguished from 
it by the narrower, more irregular apical band, by the first joint 
of antennae being less cylindrical but more incrassate or globular, 
as long as the first annulation of the third joint, by the band of 
face being indistinct, broken up into brown spots, and the hind 
tibiae are not incrassate and are less heavily fringed. Length 8 mm. 

Face greyish with white pubescence, the band is represented 
by an upper and alower brown spot. Pali reddish yellow, slender, 
with black pubescence, and some longer white hairs below. 
Antennae reddish yellow, the first joint shining, slightly nar- 
rower at base, incrassate, almost bare of pubescence, the second 
joint small with black hairs, the third joint dull red at base and 
then blackish, narrow; the difference between the antennae and 
those of the preceding species is not very marked. forehead grey- 
ish with black pubescence. Frontal callus blackish brown, narrow, 
almost straight on both its borders, spot between antennae 
black ; paired spots black, touching eyes and frontal callus; unpaired 
spot present. Thorax blackish with short appressed yellowish 
hairs, and some grey tomentum. Scutellum similar. Abdomen 
reddish brown, redder on basal segments, with distinct pale seg- 
mentations clothed with short yellowish hairs which are also 
scattered over dorsum, sides with white hairs; under side reddish 
yellow. Legs reddish yellow, fore pair blackish, the base of fore 
tibiae white, middle and posterior tibiae with faint brown typical 
rings which are fairly distinct however on the other female, apical 
joints of all tarsi blackish. Wings very similar to those of H. 
latijascta, but the apical band is not band-like but narrower and 
irregular, reaching across the whole width of wing however or very 
nearly so, broadest at its base, the inner border with a less pro- 
nounced pale edging. 


(F) Apical band of wing single, narrow, not band-like, short, not 
veaching from outer to inner border of wing. 
Haematopota inconspicua, 9, sp. n. 
(Plate xviii, fig. 31.) 


In Brit. Mus. coll. ‘Type (female) and another from Igatpuri, 
Bombay (presented by H. Maxwell Lefroy). In Indian Museum 
temales from same locality and collector (1904). 


LOLI. | G. Rrcarpo: The Oriental Tabanidae. 359 


A small species with brown wings, the apical band single, 
short, and five distinct small transverse white bars on posterior 
border of wing in the submarginal, first, second, third and fourth 
posterior cells, distinctly divided from the rosettes and almost 
touching the pale marks in the apices of these cells. Frontal callus 
black, shining, narrow, reaching the eyes, the paired spots large, 
touching it. Antennae reddish yellow, the first joint about half 
as long as the third one. Face with narrow black band and a 
black spot between the antennae. Abdomen brown with grey 
segmentations. Length type 7 mm., the other specimens 63—73 
mm. 

Face grey with white pubescence, with narrow black or brown 
band between eyes and antennae and below on each side touching 
eyes a brownish or black spot almost joining or distinct from the 
band. Palpi dull reddish yellow with pale pubescence. Anten- 
nae dull reddish yellow, the first two joints with black pubes- 
cence, the first joint cylindrical, not incrassate. the second 
small, round, about a third of the size of the first joint, the third 
joint slender, the first annulation a little dilated and broader 
than the three last ones. Forehead yellowish grey, broad, the 
paired spots large, almost round, not quite touching the eyes on 
the frontal callus, the unpaired spot small, round. Frontal callus 
black, narrow, a little produced on its upper border ; a black spot 
is present between the antennae. Thorax brown with three distinct 
yellowish grey stripes interrupted at median suture, base of thorax 
with a grey border ; sides of thorax yellowish grey; breast grey. 
Scutellum same colour as thorax or morereddish brown. Abdomen 
reddish brown with distinct grey segmentations, no spots visible, 
pubescence short and fulvous; under side similar. Legs reddish 
brown with base of fore tibiae, two rings on middle and posterior 
tibiae and basal joints of middle and posterior tarsi yellowish, 
the fore tibiae hardly incrassate, pubescence of legs nowhere 
thick or noticeable. Wings pale brown, the pale markings very 
distinct, the apical band single, short, three rosettes distinct, from 
the centre of the lower one situated in the apex of the second 
basal cell a curved white line starts, crossing the anal cell into the 
axillary angle of wing, reaching the posterior border, produced to a 
point in the middle of the axillary angle ; stigma and veins reddish 
brown, 


Haematopota immaculata, 2, n. sp. 
(Plate xviii, fig. 33.) 


In Howlett coll. type (female) and others from Kanara, Bom- 
bay, and from Bombay. 

A species allied to H. inconspicua, n. sp., but distinguished by 
its larger size, the greater length of the first joint of antennae (more 
than half as long as the third joint), by the absence of any black 
spot between the antennae, and of any black band on the face, 
though a spot is usually present below on each side. Frontal 


360 Records of the Indian Museum. (Vou, EVs 


callus is wider and with its borders straight, the paired spots are 
smaller and further removed from the frontal callus. Thorax with 
no stripes visible as a rule, brown with short fulvous pubescence. 
Scutellum similar. Abdomen brown or blackish brown with some 
greyish yellow tomentum which in some specimens appears as 
lateral spots, but in the type the colour of abdomen is uniform with 
some pale yellowish pubescence at sides. Wings very similar to 
those of H. inconspicua, n. sp., the apical pand similar, and the 
transverse bars near posterior border of wing, but on the border 
there are no light markings in the apices of the cells. In all other 
respects similar to H. inconspicua, from which however it is very 
distinct. Length type 8 mm., others 8—9 mm. 


Group VII. 


Antennae long and slender, the first joint as long as the second 
and third together. 


Haematopota cilipes, 2 , Bigot. 


Nouv. Archiv. Mus. d’Hist. Nat. Paris (3), ii, p. 205 (1890) ; 
1d., Bull. Soc. Zool. France, xvi, p. 77 (1891) ; Ricardo, Ann. Mag. 
Nat Hist? (8). p57) (god): 

Length Ir mm. 

Antennae chestnut coloured, elongated, the first segment not 
incrassate, as long as all the others taken together, the third joint 
at base very much longer than the apical ones together : palpi and 
face ashy grey, beard white; forehead black- brown, at the base 
shining chestnut coloured, marked with two lateral black spots ; 
the whole body black-brown. Thorax ashy grey behind, with 
narrow margins, pleurae ashy grey ; calyptrae and halteres brown ; 
wings almost black with white markings and spots ; anterior and 
posterior tibiae fringed below with thick short black hairs, anterior 
legs black, tibiae at base white, intermediate femora and tarsi 
dull chestnut coloured, intermediate tibiae pale fulvous, Vase, apex 
and median ring brown, posterior legs similarly coloured, femora 
above with long white pubescence. 

Antennae rather slender, much longer than the head, brownish, 
at the base slightly reddish, the first joint narrow, the first annula- 
tion of the third joint much longer than the following ones to- 
gether; palpi and face grey; beard white; forehead blackish, a 
transverse irregular callus above the base of the antennae, shining 
red, two black spots near the eyes ; body wholly blackish, with the 
posterior border of thorax and of the abdominal segments very nar- 
rowly bordered with grey colour ; calyptrae and halteres brownish, 
wings blackish with white markings and rather scattered spots ; 
anterior and posterior tibiae shortly and densely fringed black ; 
anterior legs black, the tibiae whitish at base, intermediate and 
posterior legs brownish, the tibiae fawn coloured, black at base 
and apex, ornamented with a median black ring, the posterior 
femora furnished above with white hairs. Laos I specimen. 


TOrr.| G. Ricarpo: The Oriental Tabanidae. 361 


The conformation of the antennae and in particular the hairi- 
ness of the tibiae as well as the posterior femora might establish 
this species as a new genus, divided off from the old genus Haema- 
topota, which perhaps itself is very little homogeneous. Bigot, 
Nouv. Archives Mus. d’Hist. Nat. Paris, 1890 (3), il, p. 205. 

The type came from Laos, a district of Siam, and was seen 
by me in the Paris Museum after my paper on Haematopota had 
gone to the printers. 

It should go in the table published on p. 114 of the Ann. 
Mag. Nat. Hist. (7), xviii (August 1906), under the heading “15"’, 
preceding H. lata, Ricardo, thus:—‘‘ Black species, with all the 
tibiae densely fringed and the posterior femora with white hairs 
on their lower border.’’ The hind tibiae have a very thick black 
fringe, and with the white-haired femora should easily distinguish 
the species. The antennae are long, the first joint being longer 
than the second and third together and is a little incrassate; the 
second is very small, the third rather long and slender. The wings 
are of the ordinary type and have three distinct rosettes, the 
apical band reaching across the apex; the appendix is long. The 
frontal callus is large, reddish, with the black paired spots above. 
Ricardo, Ann. Mag. Nat. Hist. (8), 1, p. 57. 


UDENOCERA, Ricardo. 


Ann. Mag. Nat. Hist. (7), xiv, p. 352 (1904). 


This genus is nearly related to the two South American genera 
Diachlorus, Ost. Sack., and Acanthocera, Macq., and is formed for 
a species from Ceylon which bears a general resemblance to the 
latter genus, but is distinguished from it by the absence of a tooth 
on the third joint of the antennae, which also divides it from 
Dichelacera, Macq. It is distinguished from Dziachlorus by the 
length of the first joint of the antennae, which is nearly as long 
as the third joint, and by the altogether longer slender antennae, 
the third joint being cylindrical; the fore tibiae are neither curved 
nor broader; these differences, together with the presence of a 
tubercular projection situated more than half-way down the face 
on which the antennae are placed, seem to justify the formation 
of a new genus. 

Generic characters.—Antennae long, cylindrical, with no tooth 
or projection on the third joint; the first joint is quite two-thirds 
the length of the third joint, the second more than half as long 
as the first joint, the third joint has the first division as long as 
the four last divisions together, the latter being of equal length, 
with the exception of the last one, which is a little longer, ending 
in a blunt point; they are situated on the lower edge of a protu- 
berant tubercle which reaches the eyes on each side and is itself 
placed more than half-way down the face; the forehead is long, 
gradually narrowing to the vertex. The palpi are Tabanus-like in 
shape, ending in an acute point. ‘The abdomen is long and narrow, 


362 Records of the Indian Museum. [Vor.7 IVs 


the same width throughout. The wings are longer than the body 
and rather large. The legs are long and slender. 


Udenocera brunnea, ? , Ricardo. 
(Plate xvi, fig. 8.) 


Ann. Mag. Nat. Hist. (7), xiv, p. 355 (1904). 


Three females from Kandy, Ceylon (Yerbury). The type is 
numbered 31, 5, 92, the others 18, 5, 92 and 30, 5, 92. 

Brown, Face shining brown below the antennae, forming a 
triangular convex tubercle, with the apex reaching the proboscis, 
and longer than the cheeks; on each side of the face is a brown 
shining stripe bordering the eyes for a short distance and then 
the facial tubercle to its apex; the cheeks are ashy grey, with long 
white hairs. The proboscis short, the palpi nearly as long; the 
first joint short, the second stout at its base, curved, ending in a 
point, yellow, with black pubescence. The first two joints of the 
antennae yellow, with black pubescence, the third black; the 
antennal tubercle brown, shining, grey below; the forehead brown, 
shining, convex in the middle, with a transverse groove near the 
tubercle. Eyes bare, the facets of equal size; on the posterior 
border the eyes project beyond the vertex of the forehead. 
Thorax brownish, shining, with indistinct linear yellow stripes on 
each side, yellow at the sides; the breast brown with black hairs, 
then grey with whitish hairs. Scutellum yellow-brown at extreme 
base, with yellow pubescence. Abdomen of a uniform brown, 
somewhat shining, with brown pubescence, the under side yellowish 
at the base. Legs pale yellow, with the femora, apices of tibiae 
and apices of all the joints of the tarsi except the first one brown ; 
the last joint of the tarsi wholly brown, with pulvilli and claws ; 
the pubescence of the legs is yellow on the yellow parts and black 
on the brown parts. Wings hyaline, the veins yellowish brown, 
the fore border and the apex brown; some light spaces are appa- 
rent in the brown of the apex; the brown shading extends along 
the posterior border faintly into the fourth posterior cell; on one 
wing of the type the apical brown is reduced to a shading of the 
veins. 

Length II mm. 

@. Four specimens from Kandy, Ceylon, are evidently the 
mates of this species though differing in colouring, being reddish 
yellow. 

Eyes.—The large facettes occupy two-thirds of the eye, reach- 
ing the upper end of frontal triangle. Face ashy grey with some 
weak brownish hairs. Beard white. Pa/pi short and stout, yellow 
with greyish tomentum and brown hairs. Antennal tubercle not 
quite so protuberant as in female. Frontal triangle blackish with 
grey tomentum. Thorax bare, chamois-coloured, with a few long 
yellowish hairs on dorsum and shorter ones. Scutellum the same, 
covered with greyish tomentum. Abdomen reddish yellow, shining, 


IQII. ] G. Ricarpo: The Oriental Tabanidae. 363 


darker and narrower at apex, some black short pubescence on 
dorsum, with some yellow hairs at the segmentations. Wzngs with 
the brown colouring fainter. 

The specimen named by Walker Chrysops parallelus, and 
referred by me doubtfully to Diachlorus [see Ann. Mag. Nat. 
Hist. (7), ix, p. 372 (1902)], from Batjan Island (Batchian) (Wal- 
lace coll.), may possibly belong to this genus; but the antennae 
are wanting, the formation of the forehead, of the antennal tubercle 
and face, besides the shape of the abdomen and legs, agrees with 
that of the species described above. 


NEOTABANUS, gen. nov. 


This genus is formed for a species from Ceylon not unlike a 
Tabanus species in appearance, but distinguished from any species 
of Tabanus known to me by the prominent tubercle below the 
forehead where the subcallus is usually situated, bearing the 
antennae, and by the narrow forehead with no frontal callus or 
other spots visible, the narrowest part of the forehead is at the 
vertex. No ocelli present. Hind tibiae with no spurs. Antennae 
are J abanus-like, the first two joints short, the third slender with 
a very slight angle at base, and composed of five divisions. The 
genus therefore belongs to the division Tabaninae, and is allied to 
Udenocera, Ricardo, in the possession of the antennal tubercle, 
but differs in the shape of antennae. 


Neotabanus ceylonicus, ? , n. sp. 


Two co-types @ from Pundaluoya, Ceylon. 

A species not unlike a Tabanus species, with a blackish thorax 
and a brownish abdomen, covered with yellowish grey tomentum 
and some scattered pale yellow pubescence. Antennae and legs 
yellowish red. Wings hyaline, tinged yellow on the fore border. 
Length 12—13 mm. 

Face greyish yellow with some greyish tomentum, the cheeks 
with long blackish hairs, below and on face some pale yellow hairs 
are discernible, face with small foveae on each side below antennae. 
Beard pale yellow. Palfi red or yellow with long pubescence, 
chiefly black, but whitish on outer sides and at base, the first joint 
very small, the second swollen, curved, ending in an obtuse point, 
nearly as long as the short proboscis. Antennae reddish, the third 
joint black, the first two joints with black pubescence, the first 
joint short, stout, nearly as broad as it is long, the second small, 
roundish, barely half as long as the first joint, the third slen- 
der, slightly wider at base, the last four divisions narrower, the 
last division longer than the preceding ones, some black hairs 
present at each segmentation. Forehead brownish with black 
pubescence, the tubercle projecting above the plane of forehead, 
shining reddish brown with traces of grey tomentum, placed below 
the eyes, and rounded off below, divided in the middle, so that 


364 Records of the Indian Museum. [VorLs ave 


each antenna may be said to be surrounded by a tuberculous 
base, the forehead is narrow, about three times as long as it is broad 
anteriorly above the tubercle, at vertex about half as broad as it 
is anteriorly. Thorax blackish brown with traces of grey tomen- 
tum, with some appressed yellowish pubescence, and scattered 
black hairs on dorsum, thick at sides which with shoulders are 
lighter in colour. Scutel/um similar to thorax. Abdomen yellowish 
brown with some grey tomentum, with appressed yellowish pubes- 
cence and some short black pubescence, sides with yellow hairs ; 
under side similar. Legs reddish yellow, pubescence on femora 
chiefly yellowish white, yellow and black on tibiae and tarsi. 
Wings hyaline with the usual neuration of Tabanus species, short 
appendix present, fore border narrowly tinged yellow, veins yellow. 


DIACHLORUS, Osten Sacken. 
Diachlorus flavipennis, 2? , Macquart. 


Dipt. exot. Suppl. iv, p. 339 (1850). 

Thorax black. Abdomen red, with the apex brown. Anten- 
nae and legs red. Wings yellow, unspotted. Length 44 lines @. 
Palpi pale yellow. Face with whitish tomentum, sides shining 
black, as well as the forehead. Antennae pale fawn coloured. 
Thorax (denuded) shining black, traces of whitish tomentum, a 
fawn-colcured spot at base of wings. Abdomen shining (denuded), 
rather narrow and elongated ; the three first segments dark fawn 
coloured, with a dorsal brown stripe; the four others black with 
fawn-coloured segmentations ; under side black with grey tomentum 
and fawn-coloured segmentations. Legs fawn coloured ; posterior 
femora blackish. Wings a little yellowish, at base and on outer 
border yellow ; neuration normal. 

From the Philippines. M. Bigot. Macquart, Dipt. exot. 


Suppl. iv; p- 339. 
Subfamily PANGONINAE. 


Hind tibiae with spurs. Ocelli usually present. 


The following table is drawn up for those genera only which 
are represented in the Oriental Region :— 


1. Third joint of antennae with eight or at least seven divi- 


sions. Proboscis usually prolonged .. Oe 

Third joint of antennae with five divisions. Proboscis 
short BP eZ bis phe 

2. Proboscis long. First posterior cell of wing open. Eyes 
hairy ae : Diatomincura, Rond. 

Eyes bare ys Subgenus Corizoneura, Rond. 

3. First and second joints of antennae short sine 


First and second joints of antennae jong oe -Bh7, 


IQLI. | G. Ricarpo: The Oriental Tabanidae. 365 


4. Face concave in the middle oF aa Wied. 
Face not concave in the middle : 5 

5. Third joint of antennae with an acute spine on the first 
annulation sed .. Gastroxides, Saunders. 

Third joint of antennae simple 6 

6. Second joint of antennae shorter than the first joint. 
Wings usually hyaline Pi Silvius, Meigen. 


7. Second joint of antennae as long or nearly as long as the 
first joint. Wings usually with a black or brown 
design re 2 Chrysops, Meigen. 


The genus Mycteromyia, Philippi, is not included in the table 
owing to the uncertainty of the genus and of the one species re- 
corded from India. 


PANGONIA, Latreille. 


This genus was subdivided by Rondani into four genera: 
Pangonia and Evephopsis with the first posterior cell closed, Dza- 
tomineura and Corizoneura with it open. The only genus as yet 
represented in the Oriental Region is Corizoneura with two species, 
not including the doubtful Pangonia amboinensis, Fabr. 


Corizoneura longirostris, Hardwicke. 


Trans. Linn. Soc. London, xiv, p. 135, pl. vi, fig. 5—6 (1823) 
(Pangonia) ; Wied., Auss. zweifl. Ins., ii, p. 621 (1830) (Pangonza) ; 
Gay in Griffith’s Cuvier Anim. King., xv, pl. cxiv, fig. 3 (1832) 
(Pangoma); Macq., Ann. Soc Entom. France, vi, p. 429, pl. xv, 
figs. 20—23 (1837) (Pangonia); Roder, Stett. Entom. Zeitg., 
xlii, p. 384 (1881) (Pangonta); Ricardo, Ann. Mag. Nat. Hist. 
(7), v, pp. 167 and 168 (1900). 


Pangoma with yellow hairs, thorax ferruginous, abdomen 
black-brown ; margins of segments yellow, wings unspotted (pl. 
vi, figs. 5 and 6). Rostrum about twelve times longer than the 
head. Antennae approximated at the base, short, consisting of 
three pieces, the two lowest of which are very small, the upper 
one long, tapering to a point, and composed of eight small articu- 
lations. Eyes large, very prominent. Head and thorax very hairy 
both above and beneath, longer under the throat ; of a bright yellow 
beneath, brownish above. Wings longer than the abdomen, hya- 
line ; costal nerves brown. Body black ; the margins of each seg- 
ment shining, the sides hairy. Halteres short, having the shafts 
yellow and the knobs ovate, black. Length of the insect from the 
base of the rostrum to the apex of the abdomen ten lines; and of 
the rostrum two inches anda half. Hardwicke, Trans. Linn. Soc. 
London, xiv, p. 135. Hardwicke’s figure of wing has the first 
posterior cell very narrow, but not quite closed at border. 

Gay, in Griffith’s Cuvier Anim. Kingdom, xv, p. 696, pl. 
cxiv, fig. 3, gives in the figure of P. longirostris the wing with the 


366 Records of the Indian Museum. [VOLE AINE 


first posterior cell closed and the proboscis nearly three times the 
length of the body. 

Macquart in Ann. Soc. Entom. France, vi, p. 429, only refers 
to the species ina general article on the genus Pangonza. 

Roder, after some remarks on the descriptions given by other 
authors, more especially by Wiedemann who probably did not 
know this species, adds, from a specimen he had from Allahabad, 
the following particulars :— 

The two first antennal joints are brownish black the third 
quite black. The two first furnished on the upper and lower sides 
with long black hairs. The long face is covered with yellow tomen | 
tum. Palpi yellow. Ocelli absent. The very long proboscis black, 
but not two and a half inches long as Wiedemann gives, but only 
Ir inch 7 lines; my specimen is only g lines long. The beard on 
cheeks and chin is long, yellow, but the forehead clothed with short 
black hairs. The thorax has a brown ground colour, covered with 
longer pale brown hairs, which are quite pale yellow on the lateral 
borders and breast sides, so that two paler stripes thus appear on 
the side borders. The scutellum is brown like the thorax and 
with the same pubescence. The abdomen is reddish at the sides, 
dark brown in the middle, the pubescence is more golden yellow. 
Under side of abdomen yellow, becoming brownish towards the 
apex. ‘The legs are wholly yellow. The first two joints of fore 
tarsi are wider in a peculiar form, the tarsal joints of the fore legs 
increasing in length. 

On the two hind pair the metatarsus is longer than the other 
tarsal joints together, the legs have only very sparse yellow 
pubescence. Wings tinged yellowish, the upper fork of the third 
vein with an appendix. ‘The first posterior cell is rather narrower 
at its apex. Wings longer than the body. Halteres not so short 
as Wiedemann gives, with apale stalk and browner head. Rodder, 
Stett. Entom. Zeitg., xlii, p. 384. 

The males have the prolongation on fore tarsi as in some 
African species. R6der mentions it. One or two of the females 
have long bristles on these joints. The first posterior cell is closed 
in some of the females with a short petiole. Two males, one of 
which was wrongly labelled ‘‘ amboinensis, Fabr.,’’ seem a variety 
of this species, having no prolongation on the fore tarsi; the 
third joint of antennae is bright red, not black. The yellow 
colour on the abdomen is more prominent; the face is shining and 
dark, with hardly any greyish pubescence. 

Hab.—North-West India; Muktesar, North-West Provinces 
(Lingard); Thibet (Landor). Ricardo, Ann. Mag. Nat. Hist. (7), 
Vv, p. 168 (1900). 

This species is easily recognized by the very long proboscis, 
the first posterior cell of wing is usually open, though it varies 
in degree, often being so very narrow that it appears closed at the 
border, but is never pedunculated. The males mentioned above 
as differing slightly from the typical form will belong to the sub- 
species given below. 


IQII.] G. Ricarpo: The Oriental Tabanidae. 3607 


Corizoneura longirostris, Fabr., subsp. varipes, subsp. nov. 


The males with no prolongation on the first two joints of fore 
tarsi and the females with these joints not at all prolonged cup-like 
appear to be a distinct subspecies of C. dongirostris distinguished 
by the plain fore tarsi and also by the bright reddish third an- 
tennal joint, the first two joints being reddish grey with black 
hairs, by the shining reddish brown face, no grey or yellow tomen- 
tum covering it, and by the abdomen being usually more largely 
reddish yellow, the first three segments being almost wholly so, 
with only median blackish spots, the remaining segments blackish 
with reddish segmentations covered with short yellow hairs. 

Male specimens of this subspecies in Brit. Mus. coll. from 
Nepal (Hardwicke bequest); and from Darjiling (G. C. Champion, 
1895). 

In Indian Museum from Sikhim; Trivandrum, Travancore ; 
Sadiya, Assam; Margherita, Assam ; Himalayas. 


Corizoneura taprobanes, 2, Walker. 


List Dipt. v, Suppl. i, p. 324 (1854) [ Pangonia]. 

Ricardo, Ann. Mag. Nat. Hist. (7), v, p. 167 (1900) ; Pan 
goma rufa, @ , Macq., Dipt. exot. Suppl. iv, p. 322 (1850), nomen 
bis lectum. 


I believe the type of tapvobanes is the same as the species 
described by Macquart as P. rufa, but I have not seen this last 
type. The species belongs to the subgenus Corizoneura probably, 
though Macquart describes his type as having the first posterior cell 
closed ; in most of the specimens I have seen it is open, varying 
in width, but in two specimens it is so narrow that it might almost 
be called closed. Walker’s name must take precedence as Mac- 
quart. described another P. vufa from Peru in 1838. 

In Brit. Mus. coll. the Walker type 2 from Ceylon (Temple- 
ton); others from Karwar, N. Canara, India (Bell) ; Bangalore ; 
Nilghiri Hills, 6,000 feet (Hampson); Periyakulam, Kandy and 
Trincomalee (where it is stated to be very common), Ceylon 
(Yerbury). 

In Indian Museum specimens from Bangalore (common), 
Arnadi, Melghat, Berar, 1,800 feet (I. H. Burkill) (1908), 

Thorax black, with red pubescence. Abdomen red with dor- 
sal black spots. Eyes naked. Antennae and legs red. Wings 
with the second submarginal cell appendiculated. 

Length 6 lines @ , proboscis as long as the body. Palpi brown. 
Beard yellow. Face black, with some grey tomentum. Forehead 
black, sides with yellowish tomentum ; ocelli present. Antennae 
alight fawn colour. Thorax with indistinct bands. Abdomen 
fawn coloured ; the first four segments with transverse black spots ; 
the three last with black bands on the anterior borders ; segmenta- 
tions yellow ; under side wholly of a testaceous fawn colour. Legs 
fawn coloured ; coxae brown. Wings reddish, second submarginal 
cell with a long appendix ; the first posterior closed. 


368 Records of the Indian Museum. [Vor.TEV? 


From Bombay..:;M. Fontanier. Museum. Macq., Dipt. exot. 
Suppl. iv, p. 322. 

A species with a comparatively short proboscis, about the 
length of the whole insect, with a reddish yellow abdomen marked 
with median black spots and white or yellowish segmentations on 
the apical segments. Thorax blackish with two well-marked 
stripes. Antennae and legs reddish yellow. Palpi-with the second 
joint shorter than the first (female). Length (without proboscis) 
males 14—17 mm., females 13—18 mm. 

a. Face black, protuberant, covered more or less with greyish 
tomentum, cheeks black but densely covered with greyish yellow 
tomentum and with yellow hairs. Palp: brown, the second joint 
nearly as long as the first joint, slightly pointed, with short black 
hairs, first joint with some longer yellow hairs at its base. Beard 
pale yellowish or white. Antennae bright reddish yellow, the first 
two joints dull yellowish grey, with some long yellow hairs, the 
third joint bare. Forehead covered with yellowish grey tomentum 
and with pale pubescence. Eyes not quite contiguous. Tvorax 
blackish brown with yellowish sides and two yellowish grey tomen- 
tose narrow stripes, these and the whole dorsum with fairly thick 
yellow pubescence, sides and breast with yellowish pubescence. 
Scutellum blackish brown with a yellowish grey tomentose outer bor- 
der and with yellow pubescence. Abdomen pale reddish yellow, the 
first segment with a median black spot which extends anteriorly to 
the sides, the second and third with broad transverse black marks, 
the fourth with a narrower one, the last two almost wholly black, 
with whitish grey or yellowish segmentations, on the second and 
third these are slightly indicated on the centre, all segmentations 
with yellow hairs, which are white on the last two or three, some 
scanty black pubescence appears even on the yellow parts and also 
on the black parts ; under side paler, only blackish at extreme apex, 
segmentations with white hairs. Legs light reddish yellow, coxae 
black, posterior tibiae and all tarsi at apex brownish, pubescence 
on femora yellowish, elsewhere chiefly black. Wings hyaline, fore 
border pale yellowish, veins brown, first posterior cell open. 

2. Palpi brown, the first joint long, narrow, about one and a 
half times as long as the second joint, which is slightly incrassate 
and awl-shaped. Forehead almost twice as broad anteriorly as it 
is at vertex, covered with yellowish grey tomentum and with two 
distinct brown stripes which start from vertex and reach more 
than half-way towards .antennae, pubescence anteriorly black, 
posteriorly yellow. 

The species is at once distinguished from C. longirostris, 
Hardw., by the shorter proboscis which is rarely longer than the 
body. 


Pangonia amboinensis, Fabr. 
Syst. Antl., p. g1 (1805); Latr., Encyclop. Method., viii, p. 704 


(ré1n) Wied. Dipt:.exot.,1,"9:53 (182n);eteAuss:: zwetd diss aie 
p. 92 (1828) ; Ricardo, Ann. Mag. Nat. Hist. (7), p. 167 (1900). 


IQrI. | G. Rrcarpo: The Oriental Tabanidae. 369 


Thorax with indistinct stripes ; abdomen above brown; mar- 
gins of segments testaceous, white below. Head with ashy grey 
pubescence, antennae ferruginous and at apex brown. Thorax 
brown with white indistinct stripes. Abdomen brown above, the 
segmentations testaceous. Body below pale. Legs testaceous. 
Fabr., Syst. Antl., p. or. 

Head with ashy grey hairs, its vertex is blackish. Antennae 
are fawn coloured. ‘Thorax is blackish almost striped with white. 
Abdomen blackish above, the borders of segments of a yellowish 
fawn colour, under side of abdomen pale. Legs of a yellowish fawn 
colour. The insect was brought from Amboina by M. Labillardiere. 
Latr., Encyclop. Méthod., viii, p. 704. 

Brown ; thorax with white stripes; abdomen with side spots 
and segmentations ferruginous; under side pale red (or chamois 
leather coloured). Six lines 9. From Amboina. 

Antennae and palpi rusty yellow. Face chamois leather col- 
oured. Beard whitish. Thorax with yellow pubescence ; breast 
sides mouldy grey coloured. The side spots of abdomen are situa- 
ted on the first three segments ; on the others the side borders are 
also rusty brown; segmentations with yellow little hairs. Wings 
brownish grey. In Fabricius’s coll. Wied., Auss. zweifl. Ins., 1, 
p- 92. 

This species may possibly be the same as Corizoneura rufa or 
Corizoneura longivostris, subsp. varipes, but as no mention is made 
of the length of the proboscis nor of the first posterior cell of wing 
it is impossible to decide without seeing the type or specimens 
from further east ; I have seen no specimens of the genus beyond 
Malaysia. 


RHINOMYZA, Wied. 
Rhinomyza fusca, ? , Wiedemann. 


Nova Dipt. Gen., 8 (1820) ; 7d., Dipt. exot. 1, 59 (1821); St. 
Farg. et Serv., Encyclop. Méthod., x, 544 (1825); Wied., Auss. 
zweifl. Ins., i, 104, pl. i, fig. 3 (1828); Jensen, Bullet. Soc. Imp. 
Nat. Moscow, iv, 323 (1832) ; Macq., Suites a Buffon, 1, 196 (1834) ; 
Ricardo, Ann. Mag. Nat. Hist. (7), viii, 289 (1901). 

Brown; base and middle of abdomen rusty yellow; wings 
brown with two yellow spots, 6 lines @. From Java. Wiedemann, 
Nova Dipt. Gen., p.8. Antennae black; a tooth-like prolonga- 
tion at the base of the last joint ; face yellowish brown, concave. 
Thorax slightly shining, brown with brown down ; sides of breast 
with a yellow spot. First segment of abdomen rusty yellow, in 
the middle a little brownish, the second dull brown narrowly 
whitish at the base, third and fourth rusty yellow in the middle, 
brown at the sides and on posterior borders, the others brownish 
black. Wings at extreme base and a median and smaller apical 
spot yellowish. Pleurae blackish brown with brown fringes of 
hair. Halteres brownish with yellow club. Legs blackish brown. 
In Westermann’s collection. Wied., Auss. zweifl. Ins., 1, p. 104. 


370 Records of the Indian Museum. [VOL.? IV:, 


The other references are merely copies of the original descrip- 
tions, the insect apparently not being known to the authors. It 
is unknown to me. 


GASTROXIDES, Saunders. 


Trans. Entom. Soc. London, iii, p. 59 (1841). 

Ditylomyia, Bigot, Rev. et Mag. Zool. (2), xi, p. 305 (1859). 

This last genus was formed for one species from Ceylon, which, 
however, on examination proves to be a specimen of Gastvoxides, 
Saunders, a genus in the division Pangoninae : see Ricardo, Ann. 
Mag. Nat. Hist. (7), vill, pp-2280,; 207.1 (TOOT), .(G carcnn aee 
Saunders, /c., pl. v, fig.4; @, Saunders, /.c., etc., should read 
thus, not as printed on page 208.) 

Ricardo, Ann. Mag. Nat. Hist. (7), xiv, p. 372. 


Gastroxides ater, 7 2 , Saunders. 
(Plate xvi, fig. 9.) 


‘rans;Mnt., Soc. Loudon; iit, p,, 50.) spl. wv, fA pe See Losey 
et iv, p. 23, pl. xiv, fig. 3, # (1847); Loew, Dipt. Stidafrik., p. 15 
(1860); Ricardo, Ann. Mag. Nat. Hist. (7), viii, p. 298 (1901) ; 
et.(7); xiv; p. 372 (1904). 

Head transverse, with three ocelli placed in an equilateral 
triangle on vertex ; proboscis straight, inclined downwards, about 
as long as the head. Antennae rather longer than the head, three- 
jointed : first joint cylindrical, twice as long as the second, which 
is longer than broad; third joint a little longer than the two first, 
with four distinct rings dividing the joint into five divisions, the 
first of which is as long as the four following, and produced at the 
bases on the superior part into an acute spine pointing forwards, 
the four last divisions equal in length. Thorax oval, rather 
broader than the head. Abdomen seven-jointed, ovato-conical, 
terminating in an acute point. Legs long and slender. 

@. Coal-black, hairy; the wings of a black-brown, with a 
yellowish irregularly oval spot crossing the middle of the disk, 
and a smaller spot of the same nature towards the apex. Expan- 
sion I inch, length 4 inch. From Central India. In my own col- 
lection. 

Obs.—This genus approaches nearest to Tabanus but differs 
in having ocelli, in the antennae, and in the shape of abdomen. Its 
approach to other genera of the family ‘Tabanidae is also evident, 
but it differs from all the genera I am acquainted with, and there- 
fore I have made it the type of a new sub-genus under the name 
of Gastroxides, in allusion to the shape of the abdomen, which is 
unusual in the family to which the insect belongs. Saunders, Trans. 
Ent. Soc. London, iii, p. 59. 

@. Differs from the female in having the second joint of the 
abdomen rufous, and the eyes large and contiguous vertically. 


IQII. | G. Ricardo: The Oriental Tabantdae. 371 


Head broader than the thorax, semicircular. Face yellowish 
brown, with a large, elevated, shining chestnut-brown tubercle in 
the centre, extending from the mouth to the base of the antennae. 
Region of the mouth black, hairy. Pvroboscis about the length of 
the head. Eyes dark rufous brown, large and meeting on the 
vertex. Antennae black, rather more slender than in the female, 
and having the spine at the base of the terminal joint not so prom- 
inent. Thorax orbicular, black, hairy. Wings marked as in the 
female. Abdomen black, with the posterior margin of the first 
joint, the whole of the second joint, and the anterior margin of 
the third joint, bright rufous. Legs black. 

Length 4 inch, expansion of wings I inch. From Northern 
India. In the collection of Col. Hearsey. Saunders, /. c., iv, 
pe2ss. pls xiv; fig.3: 

One male from Barrackpore, Calcutta (Rothney), 82, 15; one 
male from India (Saunders coll.), 54, 13; one female from Bengal 
(Campbell), 42, 25 ; one male (Saunders coll.), 68, 4. 

Saunders described and figured both the sexes ; the male type 
he mentions as belonging to a Colonel Hearsey, so that it seems 
probable that the Museum does not possess the male type, and 
certainly not the female type. There are said to be three speci- 
mens labelled Saunders coll. in the Oxford Museum, which may 
perhaps include the types. Ricardo, Ann. Mag. Nat. Hist. (7), viii, 
p. 298. 

This species is easily distinguished by the presence of a pro- 
longation on the third joint of antennae at base. The male is 
black, with basal segments of abdomen bright rufous, the female 
wholly black, both with plackish brown wings in which two pale 
spots are very distinct, the lower one situated at base of second 
submarginal cell, extending across the apex of the first basal cell 
and base of first posterior cell, occupying the basal half of discal 
cell, and the apex of second basal cell and bases of fourth and 
fifth posterior cells; the upper one is on the second submarginal 
cell round the base of fork of third longitudinal vein extending 
into the first posterior cell. ‘There are two females in the Indian 
Museum from Calcutta and locality not specified. 


Gastroxides ornatus, 2, Bigot. 

Rev. et Mag. Zool. (2), xi, p. 305 (1859). [Dutylomyza.] 

Ricardo, Ann. Mag. Nat. Hist. (7), xiv, p. 372 (1904). 

The type, by the markings of the wings and the general shape 
of the abdomen, bears a general resemblance to Gastroxides ater 
(Saunders), the only heretofore known species of the genus. Bigot 
mentions the presence of ocelli, but did not note the spines on the 
hind tibiae, these two characters dividing the division Pangoninae 
from Tabaninae. ‘The antennae have only five rings on the third 
joint, not eight or nine as he supposes. In the shape of the head, 
the insertion of the antennae, and the presence of the tubercles on 
the forehead it resembles G. ater, but is larger and of different 
colouring. 


372 Records of the Indian Museum. [Vor. IV, 


Black, with lighter hind borders to the segments of. the 
abdomen. 

Face blackish, with grey tomentum, the tumid triangular pro- 
~ jection in the middle shining red-brown ; the palpi large, flat, as long 
as the proboscis, dark brown with black hairs. Forehead black, 
shining, with two prominent tubercles, one situated near the an- 
tennae, the other about in the middle of the forehead, neither 
reaching the sides ; vertex reddish ; the yellow ocelli are placed on 
a brownish, not very prominent tubercle bordered with grey which 
is continued along the sides of the forehead. Antennae longer 
than the head, red-brown, the long spine or tooth brighter red ; 
the first joint stout, long, the second only half as long, both with 
black pubescence ; the third joint twice as long as the first, its first 
ring as long as the remaining four together, stout, bearing the long 
spine at its base, the second ring is the smallest, the third and 
fourth about equal, the fifth as long as the last two together. The 
hairs below the head are blackish. Thorax reddish brown, with 
indistinct grey tomentum and two yellowish stripes; the scutellum 
yellowish red ; the breast reddish, with brown tomentum and black 
pubescence. Abdomen black, long and pointed; the second seg- 
ment widest, the first and third with yellow, the second and 
fourth with grey hind borders, the remaining segments black with 
some red on the sides; under side black. Legs reddish brown with 
black pubescence, the fore femora stout. Wangs dark brown with 
clear base; a clear band in the middle crossing the base of the 
discal cell and extending to the fifth posterior cell, another on the 
apex crossing the fork of the third longitudinal vein ; there is also 
a clear space in the anal cell, and the axillary part of the wing is 
clear. 

Length 18 mm. 

Through the kindness of Mr. Verrall this species is now placed 
in the British Museum collection with the species for which the 
genus Gastroxtdes was formed. Ricardo, Ann. Mag. Nat. Hist. (7), 
Live pa 372. 

a”. A male in the Indian Museum coll. from Colombo, and 
another in the Brit. Mus. coll. from Henaratgoda, Ceylon (Yerbury), 
are apparently males of this species, though the antennal third 
joint in both is wanting The colouring of abdomen is different 
from that of the female, being reddish with black spots, only black 
at the apex. Face with the same tumid red-brown shining middle, 
not so protuberant however and confined to middle of face; the 
palpi shorter than proboscis, club-shaped. Frontal triangle red- 
dish. yes with the large facettes reaching the base of antennae, 
bordered posteriorly by a narrow band of the small facettes. 
Thorax and scutellum reddish brown, the latter more red on its 
posterior border, sides of thorax with black hairs. Abdomen 
reddish, shining, on the first three segments, with a median black 
spot on each, the second segment narrowly black at sides, the 
third segment with a black spot at each side, the fourth segment 
black, dull yellow on its posterior border, the remaining ones 


IgII.] G. Ricardo: The Oriental Tabanidae. 373 


black, sides with black hairs; under side red on basal segments, 
then deep black, the abdomen has a flat appearance with a pointed 
apex. Legs reddish brown, the femora darker. Wings as in 
female. Tength 16 mm. 


MYCTEROMYIA, Philippi. 


Verh. zool.-bot. Ges. Wien, xv, p. 712 (1865). 


This genus was established by the author for Pangonia conica, 
Bigot, from Chili, and Bigot later added other species from Bra- 
zil, Cape of Good Hope and one from India. Philippi gives the 
chief characteristics as: Head narrower than thorax, elongated 
in front. Eyes naked. Ocelli present. Antennal joints rather 
longer and narrower than in Panegonia, Palpi biarticulate, the 
first joint short, the second elongated, compressed, narrow, the 
apex rounded. Legsslender. Wings with first posterior cell closed. 
Dr. Lutz in Zool. Jahrb., Suppl. x, 4, p. 62, 5 (1909), remarks 
that the genus can only be retained for the Chilian species with 
naked eyes, the four Brazilian species described by Bigot having 
eyes distinctly pubescent. The description of the Indian species 
is too short and inconplete to enable anyone to judge to what 
genus it belongs, and I have seen no specimen from India at all 
resembling the original description. 


Mycteromyia nigrifacies, ? , Bigot. 


Mem. Soc. Zool. France, v, p. 607 (1892); Ricardo, Ann. 
Mag. Nat. Hist. (7), v, p. 99 (1900). 

@. Length 20 mm. (without proboscis). 

Proboscis and antennae (incomplete) black ; palpi black; fore- 
head dull black, face shining black, beard greyish white: thorax 
and scutellum dull black; abdomen shining black; squamae and 
halteres black; legs black; wings blackish, no appendix to the 
fork of third vein. India. One specimen. Bigot, Mem. Soc. 
Zool. France, v, p. 607. 


SILVIUS, Meigen. 


Syst. Beschreib., ii, p. 27 (1820). 

Ectenopsis, Macq., Dipt. exot. i, p. 115 (1838). 

Mesomyia, Macq., Dipt. exot. Suppl. iv, p. 341 (1850). 

Veprius, Rondani, Archiv. per la Zool. Modena, iii, p. 83 
(1863). 


This genus, characterized by the short first and second joints 
of antennae, the second one shorter than the first, and by the 
usually hyaline wings, has not hitherto been represented in the 
Oriental Region, but a female specimen in the Indian Museum 
from Bhura, Naini Tal district, United Provinces, India, is no 
doubt a species of Szlvius not unlike in general appearance to the 
European species Silvius vituli, Fabr., but distinct. 


374 Records of the Indian Museum. [Vor 


Silvius indianus, 2 , n. sp. 


A small reddish yellow species, antennae and face, frontal 
callus, palpi and legs yellowish. Wings hyaline, with the fore 
border narrowly dark brown. Length 9 mm. 

Face greyish yellow, convex in the middle, the sides of this 
middle part are shining reddish yellow with a greyish yellow 
rather tomentose median stripe, foveae on cheeks distinct, lower 
part of face with short white pubescence, some black hairs near the 
eyes. Proboscis short. Palpfz reddish yellow, the first joint short, 
the second about four times as long, slightly curved, flat, ending 
in a point, with some very short black pubescence. Antennae red- 
dish yellow, the first two joints yellowish with black pubescence ,the 
first joint twice as long as the second one, the third long and slen- 
der, very slightly wider at base, blackish on the last four annu- 
lations. Forehead greyish yellow, the frontal callus shining red- 
dish yellow, heart-shaped, not reaching the eyes, ocelli at vertex 
placed on a blackish spot. Thorax blackish but densely covered 
with greyish yellow tomentum and pubescence, appearing yellowish. 
Scutellum identical. Abdomen more reddish yellow, with a 
median short black stripe on the first two segments reaching to 
the third segment, pubescence chiefly consists of very short black 
hairs ; under side the same. Legs reddish yellow, the apex of fore 
tibiae and all tarsi blackish. Wangs hyaline, the costal cell wholly 
yellowish brown, the marginal cell the same at its base with a slight 
shading at its apex on fore border, the stigma large, dark brown, 
the first longitudinal vein thickened, dark brown, the others pale 
yellow, all cells open except the apical cell, appendix hardly visible. 


CHRYSOPS, Meigen. 


Nouvelle Classification, p. 23 (1800); Illiger’s Magazine, ii, 
p. 267 (1803). 

The species of this genus of the subfamily Pangoninae are 
easily recognized by the wings being usually ornamented with a 
dark median cross-band, by the shining black tubercles on the face 
and forehead, with three ocelli on the vertex, and by the long 
antennae, the second joint usually as long or nearly as long as the 
first joint. ‘The species are small in size and are not very numer- 
ous; in Ann. Mag. Nat. Hist. (7), ix, p. 372 (1902), I brought 
the number up to 20, now reduced by discovery of further syno- 
nyms to 16 with 1 new species described, making the total 17. The 
following is a list of all the described species and synonyms :— 


Curyvysops alter, Rondani. 
bifasciata, Macquart = dispar, Fabr. 
cincta, Bigot. 
clavicrus, Thomson ? = cincta Bigot. 
designata, sp. n. 
dispar, Fabricius. 


IgII.| 


|S) 


On 


G. Rrcarbo: The Oriental Tabanidae. 375 


fasciata, Wiedemann. 
fixissima, Walker. 
flaviventris, Macquart. 
flavocincta, Ricardo. 
impar, Rondani= dispar, 
indiana, Ricardo. 


Fabricius. 


tranensts, Bigot = mlokosiewiczi, Bigot. 
“igata, Walker = dispar, Fabricius. 
lunata, Grey (Haematopota) = dispar, Fabricius. 


manilensis, Schiner. 

mlokosiewiczi, Bigot. 
pellucidus, Fabricius. 
rufitarsis, Macquart. 


semucirycula, Walker = dispar, 


signifer, Walker. 
sinensis, Walker. 
stimulans, Walker. 


Fabricius. 


striata, V. d. Wulp (nomen bis lectum) = mloko- 


siewiczi, Bigot. 


ternunalis, Walker = dispar, Fabricius. 


translucens, Macquart. 


unizonata, Rondani = fixissima, Walker. 


Table of spectes. 


Wings with a dark transverse 
band and an apical spot 

Wings with a dark transverse 
band but no apical spot 

Wings with a hyaline sinus on 
the posterior part of the band, 
in the fifth posterior cell 

Wings with no hyaline sinus on 
the posterior part of the band, 
in the fifth posterior cell 

Abdomen blackish, yellow at 
apex, with grey bands on the 
first two segments and a grey 
median stripe oe 

Abdomen yellowish with black 
stripes 

Abdomen black, w hitish or yellow: 
ish at base 

Discal cell hyaline. "Abdomen 
with four long black stripes 

Discal cell not hyaline 

Abdomen with a short black bifid 
stripe on the second segment, 


often extending to the third or 


fourth segment 


age 


W 


N 


mantlensis, Schiner. 


ue i 


mlokosiewic2, Bigot. 


dispar, Fabr. 


376 


Io. 


Records of the Indian Museum. 


Abdomen with a bifid stripe on 
the second segment and a net- 
work of black markings on the 
third segment, leaving only 
three yellow spots free 

Abdomen whitish on the first four 
segments, apex black 

Abdomen black, the first two 
segments largely yellow, the 
second and third segments 
with small median yellow 
spots. Face yellowish. Apical 
border of band of wing Tope 
concave 

Abdomen black, the second see 
ment yellowish on fore border. 
Face blackish. Apical border 
of band of wing straight : 

Small species, abdomen blackish, 
the first two segments yellow 
anteriorly. Face black 

Large robust species. Abdo- 
men blackish with paler me- 
dian spots, largely yellow on 
the basal segments. Face 
yellowish. Apical border of 
wing irregular 

Wings with a clear spot in the 
discal cell, first joint of an- 
tennae slightly incrassate. 
Abdomen blackish with pale 
spots 

Wings with no clear spot in dis- 
cal cell 

Band of wing broad throughout. 
Abdomen yellow with a black 
bifid stripe in the second seg- 
ment 

Abdomen yellowish yin a Black 
band in the middle ‘ 

Abdomen yellowish with ‘ee 
distinct black bands 

Abdomen with blackish legs and 
PACE i. 

Abdomen with yellowish roam 
legs and face By 

Abdomen with the black bands 
narrow : aie 

Abdomen darker, the black 
bands broader 


designata, n. sp. 


vufitarsis, Macq. 


tvanslucens, Macq. 


pellucida, Fabr. 


altey, Rondani,. 


sinensis, Walker. 


stimulans, Walker. 


indiana, Ricardo. 


fasciata, Wied. 
fixissima, Walker. 
signifer, Walker. 


ctncta, Bigot. 


[Vox: IV, 


(oa) 


10 


IQII.] G. Rrcarpo: The Oriental Tabanidae. 377 


11. Abdomen black with a yellowish 
band on the second segment. 
Small species ve ..  flavocincta, Ricardo. 


Norr.—Chrysops flaviventris, Macquart, is not included owing 
to the insufficient description. The only species of Chrysops recor- 
ded from New Guinea is Chrysops albicinctus, V.d, Wulp (Tijd. 
Ent. xi, p. 103,1868), which Osten Sacken in Ann. Mus. Civ. 
Genova. xvi, p. 418 (1882), remarks may be the same as C. pellu- 
cida, Fabr., but if V d. Wulp’s figure of the wing is correct it is 
quite distinct from any species of the Oriental Region, the whole 
apex of the wing being brownish grey, bearing between it and the 
transverse dark band a hyaline streak. 


Chrysops manilensis, ¢? , Schiner. 


Reise Novara Diptera, p. 104 (1868); Ricardo, Ann. Mag. 
Nat . Hist. (7), ix, p- 373 (1902). 

Thorax and scutellum black, with slaty grey tomentum, the 
former with two other indistinct contiguous pale grey stripes and 
on each side a rusty yellow spot; in front of the  scutellum 
appears a conspicuous band of golden yellow hairs; similar hairs 
appear on the rusty yellow spots (perhaps in well-preserved speci- 
mens they are spread over the whole thorax). Breast sides blackish, 
a yellow callus beneath the base of wings. Abdomen black with 
an oblique grey band at the bases of the first two segments, 
the second and third segments with a similar coloured median line, 
the following segments rusty yellow, the under side chiefly yellow on 
the middle, blackish at the base and apex (‘‘ mitte’’ is given in 
the original, probably an error). The abdomen is somewhat com- 
pressed in both specimens, it may therefore be somewhat different 
in design in well-preserved specimens. Head black, the face with 
golden yellow pubescence on each side and in the middle, so that 
the shining black tubercle takes the shape of a Latin V. Forehead 
with yellowish grey tomentum, with a large convex shining black 
callus above the antennae, impressed in the middle, and a shining 
black spot at vertex. Antennae unusually long and very slender, 
the joints almost equal in length, the two basal joints yellowish, 
the third brown. Palpi bright red-yellow. Legs red-yellow, the 
femora at their extreme bases, and the posterior ones almost 
wholly, brown. ‘Tarsi blackish on the last joints. Fore tibiae a 
little curved. Wings hyaline, the fore border and a median band 
pale brown, the outer border of the latter concave, posteriorly 
with a bluntly triangular hyaline incision. The lower margin of 
the brown fore border is paler on the other side of the median 
band. ‘Two females from Manila. Length 3 lines. 


The species could only be compared with Chrysops bifascratus , 


Maeq., but this species has a reddish yellow scutellum and the 
markings of the under side of abdomen are different; there can be 


378 Records of the Indian Museum. [Vor. IV, 


no question of its identity with Chrysops costatus, W., which Walker 
takes to be identical with Chrysops bifasciatus, Macq. Chrysobs 
costatus is distinguished from both species by the wholly grey 
apex of wing. ‘Till now no species of Chrysops has been known 
from Manila, and in general few from the continents lying to our 
east. Schiner, Reise Novara Diptera, p. 104. 


The allusion to Walker’s identification of Chrysops costatus 
with the Macquart species is to be found in List Dipt. Brit. Mus. 
v, suppl. i, p. 288, and may be disregarded; Chrysops costatus 
is a South American species. As I believe Chrysops bifasciatus, 
Macq., to be a synonym of Chrysops dispar, Fabr., this species is 
probably identical in the wing-markings with this latter, but can 
hardly be the same, judging from Schiner’s description of the 
abdomen. ‘The species is unknown to me. 


Chrysops mlokosiewiczi, 2 , Bigot. 


Ann. Soc. Entom. France (5), x, p._146 (1880); Ricardo, 
Ann. Mag, Nat. Hist. (7), ix, p. 378 (1902). 

Chrysops striatus, 2 , V. d. Wulp, Notes Leyden Museum, vii, 
p-.79 (1885). 

Chrysops ivanensis, 2, Bigot, Mem. Soc. Zool. France, v, 
p. 602 (1891). 

Thorax ashy grey with three brown stripes: abdomen 
ochraceous with four black stripes, head yellowish with a shining 
black frontal callus, the facial calli shining testaceous ; antennae, 
palpi and legs red; apex of antennae black ; wings hyaline; with 
the base, the costal border to the apex, and the median band 
which does not reach the posterior border, brown. @?. Length 
6 mm. 

Antennae one and a half as long as the head, almost bare, 
reddish yellow, black towards the end; second joint shorter than 
the first ; the third one about as long as the other two together. 
Front with yellowish grey dust ; the ocellar triangle darker ; above 
the antennae a large, shining black knob; face rather prominent 
on account of the two large knobs, which are testaceous, very 
shining and coherent ; cheeks with yellowish grey dust. Rostrum 
fuscous ; palpi reddish yellow. ‘Thorax yellowish cinereous, with 
three very distinct fuscous stripes ; scutellum coloured like the 
thorax. Abdomen ochraceous, with two black longitudinal bands, 
which are interrupted at the segmentations, and moreover with a 
less distinct lateral row of blackish spots; under surface of the 
abdomen uniform ochraceous, darker towards the end. Legs 
reddish yellow; posterior coxae cinereous; tarsi brown; the 
tibiae not dilated. Halteres yellowish brown. Wings hyaline, 
the base, the costal border and the usual cross-band brown; the 
latter extends tili half the breadth of the wing, surrounds the 
discal cell, the centre of which remains hyaline, and goes along 


TOTTI. | G. Ricardo: The Onental Tabanidae. 379 


the posterior cross-vein and the apical portion of the postical 
vein. A female specimen from Amoy, China (Buddingh). V. d. 
Wulp, Notes Leyden Museum, vii, p. 70. 

In Brit. Mus. coll a male and female from Tygosan, Chusan 
Archipelago, 1892 (Walker); and a series from Japan, 1909 
(Dr. Myajima). 

A well-marked species easily distinguished by the four narrow 
black stripes on the yellow abdomen and _ by the cross-band of 
wing not reaching beyond the fifth longitudinal vein, the posterior 
cross-vein and apical portion of the sixth longitudinal vein being 
shaded with brown only, the centre of the discal cell is hyaline. 
The fece is covered with yellowish tomentum, the sides with shin- 
ing testaceous tubercles, and the centre of face over the oral 
opening is the same colour; a transverse black shining spot is 
present on the lower part of each cheek, not attaining the oral 
opening. Forehead with greyish tomentum, darker at vertex, the 
frontal callus convex, black and shining, not reaching the eyes. 
Under side of abdomen often with medium black markings besides 
being darker at the apex. Length of specimens 8-10 mm. 


Among the types kindly lent me by Mr Verrall there is one 
(? female) specimen with the head wanting, labelled as above and 
from the Caucasus, but the specific name is crossed out and 
““ gvanensis’’ scribbled underneath ; there is another female speci- 
men only labelled ‘‘ N. Persia.’”’ Bigot described one female as 
C. mlokostewiczt, from N. Persia or Caucasus, and later two 
females from N. Persia as C. tvanensis; apparently he concluded 
finally that all were one species, which certainly seems likely 
judging from the descriptions: in this case the name mlokositewtczi 
would have priority ; it seems nearly related, if not identical, 
with a species described by Wulp from Amoy, China, and named by 
him C. striatus. On comparison of the two specimens of his species 
in the British Museum collection with the Bigot specimens, the 
only difference to be seen is the colour of the tubercle above the 
antennae, which is nearly wholly reddish in the specimen from 
N. Persia, not shining black, as Wulp describes; they are exactly 
similar in the wing; the headless specimen has the colouring of 
the abdomen more greyish than ochraceous, as in C. striatus. 
Considering the distance between the localities, it is perhaps pre- 
ferable to keep the species apart for the present, placing, however, 
the Bigot species among those of the Oriental Region. Ricardo, 
Ann. Mag. Nat. Hist. (7), ix, p. 378 (1902). 

There can now be little doubt that the two Bigot species are 
the same as Wulp’s species ‘‘ s/vzatus,’ as in the series lately 
received from Japan there is one female with the tubercle reddish 
yellow. As the name striatus is preoccupied by a species so named 
by Osten Sacken from N. America in 1876, the species must 
henceforth be known as Chrysops n lokostewiczt, and belongs to 
both the Palaearctic and Oriental Regions, ranging apparently 
from east to west of the Continent. 


380 Records of the Indian Museum. [Vor LV, 


Chrysops dispar, 7 @ , Fabricius. 
(Plate xv, fig. 3.) 


Ent. Syst. Suppl. 567 (Tabanus) (1798); cd., Syst. Antl., 
p. 112, 5. (§805);' Wied., Dipt. exot:, “—p. 102, 1 (1624) 7d., Auss: 
zweifl.” Ins:, 1, Dp. 106) (a828)%" Macq., Dipt. exot. 1 (rn) aepan los 
(1838) ; 7d., Suppl. iii, p. 174 (1848); V. d. Wulp, Sumatra Exped. 
Dipt., p. 19 (1881); Ost. Sack., Ann. Mus. Civ. Genova, xvi, 
p. 418 (1882); zd., Berlin. Ent. Zeitschr., xxvi, p. 97 (1882): 
Réder, Ent. Nachricht, xix, p. 234 (1893); Ricardo, Ann. Mag. 
Nat: Hist. (7), 1x; p: 374 (902): 

Haematopota lunata, 2, Gray, in Griffith’s Cuvier Anim. 
Kingdom, xv, p. 696, pl. exiv, fig. 4 (1832). 

Chrysops bifasciata, 2 , Macquart, Dipt. exot. (1), 1, p- 161 
(1838). 

Chrysops ligatus, @ , Walker, List Dipt. Brit. Mus., i, p. 195 
(1848). 

Chrysops semicirvculus, 2, Walker, List Dipt. Brit. Mus., i, 
p. 196 (1848); Ricardo, Ann. Mag: Nat. Hist. (7), ix, p. 375 
(1g02). 

Chrysops terminalis, 9 , Walker, List. Dipt. Brit. Mus., i, 
p- 195 (1848). 

Chrysops tmpar, 2 , Rondani, Ann. Mus. Civ Genova, vii, 
p. 460 (1875). 

Abdomen with the first segment pale; with a black hali- 
moon, wings white: with a black band. Antennae testaceous, 
the apex dark. Head below the antennae yellow with two large, 
raised, ovate, testaceous spots, above the antennae pale, with a 
large transverse raised black spot. Thorax ashy grey with sides 
yellow. Abdomen brown with the first segment pale, the middle 
half-moon-shaped spot brown. Wings white, the costal border 
and wide middle band black, the latter at the margin paler with 
an ovate white spot. Male with eyes joined, abdomen testaceous, 
the first segment however pale with a half-moon-shaped brown 
spot. 

Habitat.—-E. India. Fabr., Ent. Syst. Suppl. 567 (Tabanus). 

Abdomen with the second segment pale; with a black half- 
moon. Wings white with a black band. Habitat in East India. 
Babr .oyst; Ant! py rir2> 5, 

Antennae ochre brownish with a blackish brown apex. Face 
rusty brownish with a paler middle line; cheeks light rusty yellow 
with a smooth brown spot ; forehead yellowish ; in the female with 
a blackish brown transverse oval tubercle. ‘Thorax brown with two 
mouldy grey stripes only separated by a brown line, with golden 
yellow pubescence: breast sides brown with a golden stripe under 
the base of wings. Scutellum rusty yellow. Abdomen of the male 
rusty yellow, the first segment brown in the middle, the second 
segment pale, with two brown spots united towards the base of the 
segment in a somewhat rounded angle, not resting on the base 


IgII.| G. Ricarpo : The Oriental Tabanidae. 381 


itself, continued as far as the base of the third segment ; sides of 
abdomen brown, in the middle of the under side of abdomen an 
abbreviated brown band. Abdomen of the female more brownish, 
the first segment pale, the second paler still, almost whitish anteri- 
orly, the spots as in the male, but not continued to the third 
segment. All segmentations broadly paler, the first brown in the 
middle. Wings perfectly hyaline, on the fore border from the base 
to the apex, brown, below the cross-veins at the base is a some- 
what oblique small inconspicuous band; the larger band has a 
deep incision on the inner border. Legs bright rusty yellow. In 
Fabricius’s, Westermann’s and my own collection. Wiedemann, 
Auss. zweifl. Ins., i, p. 196. 

A female specimen from Java in the collection of M. Serville 
differs from Wiedemanu’s description by the shining blackness of 
the face, with a grey spot on each side; by the grey forehead 
with two black calli, by the posterior border of thorax with 
golden yellow hairs, by the black scutellum, by the fawn-coloured 
legs with black coxae, and also the posterior femora. Macquart, 
Dipt: exot. i (1), p. 163: 

According to Wiedemann the spot or rather the brown bifid 
stripe of the abdomen extends over the second and third segment 
of the male, it does not extend beyond the second one in the 
female. We have observed five females and two males in which 
the stripe is equally present on the second and third segments. 
From Java. M. Payen’s collection. Macquart, Dipt. exot. Suppl. 
ili, p. 174. 

Seven females from Buitenzorg, Java (1875) (Ferrari). The 
identification is certain, although there are discrepancies, especi- 
ally in the description of the face. Macquart notices the same 
differences in Dipt. exot. i, p. 163. Osten Sacken, Ann. Mus. 
Civ. Genova, xvi, p. 418. Rd6der, Ent. Nachricht, p. 234, records 
the species from Ceylon. 

Six females from different places (in Sumatra). The above 
specimens differ from the description and from most specimens 
from Java by their darker colour, especially by the black 
scutellum and the black- brown posterior femora. ‘The posterior 
tibiae in this species are broader in their whole length, and each 
side with a short fine fringe of hairs. The face is very shining. 
V.d. Wulp, Dipt. Sumatra, p. 19. 

The figure of Haematopota lunata is evidently that of a 
specimen of Chrysops dispar. No description is given. Chrysops 
ligatus, Wik., from Bengal is only a rather pale-coloured specimen 
with some lighter spaces in the wing-cells. Chrysops semicirculus, 
a female in very bad condition, is, I now believe, only a poor 
specimen of this species with the black markings of the abdomen 
obsolete or almost so. Chrysops terminalis, W1k., is a pale-coloured 
specimen with the black bifid stripe not reaching beyond the 
second segment, as in Wiedemann’s original description of 
Chrysops dispar. Walker identified several specimens as Wiede- 
mann’s species and then described his two new species, placing 


382 Records of the Indian Museum. [VoL. IV, 


them directly after Chrysops dispar in his Catalogue, but I can 
see no differences sufficiently marked to justify them being made 
distinct species. Chrysops impar, Osten Sacken believed to be 
the same as the Wiedemann species, after seeing the type (see 
Berlin. Ent. Zeit., xxvi, p. 97), an opinion with which I agree, having 
also seen the type, which has the abdomen however entirely black 
after the third segment, and the frontal callus is deeply incised 
posteriorly a peculiarity I have not seen in any other specimens, 
but there are very few specimens in the collections I have had 
access to, from Borneo, whence Rondani’s came. 

The type of Chrysops bifasciata, Macquart, I have seen in the 
Paris Museum and believe it to be a specimen of Chrysops dispar 
though the black stripes on abdomen are of unusual length, ex- 
tending to the end of the abdomen, which I have not observed in 
any other specimens from India or elsewhere. Macquart’s type 
is from Bengal. His description of the antennae is incorrect, 
they are almost wholly yellowish, a little darker on the second 
and third joints, not ‘‘ black, red at the base.” 

The species varies a little, chiefly in the colour of the scutellum 
which ranges from reddish to reddish brown or nearly black. 
The /egs are usually testaceous or yellowish but the posterior 
femora are sometimes brownish or blackish, the coxae are also 
darker in some specimens ‘The forehead is greyish or yellowish. 
The abdominal stripe beginning on the second segment is continued 
to the base of the segment or to the base of the third and occa- 
sionally reaches the fourth segment. It is an easily distinguished 
species, with a yellowish abdomen, the bifid stripe always distinct, 
and a dark narrow band on the posterior border of the firstsegment. 
The thorax in well-preserved specimens has golden yellow pubes 
cence at the sides and base. The face and cheeks yellowish 
brownish in the centre with the exception of a narrow yellow line 
and with two black stripe-like spots on the lower part of cheeks. 
Wings with a brown fore border and transverse band, chiefly 
incised on its outer border. Length from 8—11r mm. 

In the British Museum are specimens from Ceylon (Yerbury, 
Green) ; Nepal (Hardwicke bequest); Bangalore (Watson) ; Moul- 
mein, K. India (Clark); Singapore and Penang (Ridley); Ran- 
goon; Annam; Hongkong (Bowring) ; Java; Sumatra. 

In Indian Museum coll. from Balighai, near Puri, Orissa; 
Kulattupuzha, W. base of W. Ghats, Travancore ; Nedumangad, 
to miles N. E. of Trivandrum, Travancore; Morapai, S. Bengal ; 
Bukit Besar, Patani States, Siam; I have also identified speci- 
mens from Formosa. On one of the specimens from Ceylon Col. 
Yerbury has the following note: ‘‘ Common and generally distri- 
buted. ‘Tormentscattle. The scutellum and pilose stripe on the 
pleurae in life bright gamboge yellow.” 

[Flies of this common species conceal themselves during the 
day on the lower surface of the leaves of shrubs, becoming active 
towards evening. I found thespecies exceedingly common at Bali- 
shai in October.—N. ANNANDALE. | 


IQII. | G. Ricarpo: The Oriental Tabanidae. 383 


Chrysops designata, @ o,sp.n. 
(Plate xv slice Te) 


Type female from Sarah, Nepal, 24-xi-o8, in Indian Museum 
coll. and another from Yunnan, W. China; type male from Jaulasal, 
Naini Tal district. 

@. Abdomen yellowish, marked with black on the first 
three segments, bearing only three yellow spots on the third 
segment. Wings with cross-band and apical spot, the former 
with a well-marked sinus on posterior border, its outer border 
irregular. Antennae long, cylindrical. Length 9 mm. 

Face yellowish, the facial tubercles shining, blackish brown, 
reaching the oral opening but bearing a large yellow stripe in the 
centre of face, lower part of cheeks black, with yellow hairs. 
Palpi reddish yellow with scanty yellowish pubescence. Antennac 
long, cylindrical, the first joint a little longer than the second one, 
reddish yellow with black pubescence, the second and_ third 
blackish, the second with black pubescence, the third joint the 
longest of the three but barely as long as the first two joints 
together. Forehead same colour as face with yellow pubes- 
cence, the frontal callus shining, blackish brown, transverse, not 
reaching the eyes, with an impressed line in the centre and slightly 
produced toward the antennae, ocelli visible at vertex which 
is not darker in colour. Thorax blackish brown, with a very 
broad yellowish grey tomentose median stripe divided in two by 
a narrow brown line, sides yellowish clothed with bright yellow 
pubescence, which is also present on the dorsum as short scanty 
pubescence. Scutellum yellowish grey, blackish in the centre but 
probably this is due to denudation, with darker pubescence. 
Abdomen yellowish, paler yellow on the first two segments, on the 
following segments more reddish yellow, with the apical slightly 
darker, pubescence yellowish, thick on the apical segments; the 
black markings consist of a narrow band on the posterior border 
of first segment widening in the centre into a broad median spot 
reaching the anterior border, with concave sides, on the second 
segment of the ordinary chevron-like black spots meeting on the 
anterior border, with likewise a narrow band on the posterior bor- 
der, on the third segment the black chevron-like spots are present 
but broader, not meeting on the anterior border, the sides and pos- 
terior border have black bands, so that the yellow colour remains 
as three large spots, the middle one irregularly triangular, the side 
ones oblong, large, on the fourth segment two small black triangular 
spots with their apices pointing downwards are present on the 
anterior border which is also blackish at the sides; under side 
wholly yellow. Legs reddish yellow, the coxae, femora and apical 
joints of tarsi blackish. Wangs hyaline with brown fore border 
reaching the apex, at base extending slightly into the basal cells, 
and with a brown cross-band which reaches the anal cell; the 
fifth posterior cell is largely hyaline, the lower border of band 
almost straight not extending into the basal cells, its upper border 


384 Records of the Indian Museum. [Vor IV, 


irregular, produced towards the base of fork of third longitudinal 
vein and again at the fourth posterior cell; the apical spot is 
narrow, the same width throughout, reaching exactly to the centre 
of apex of wing. 

». Smaller than the female, measuring 8 mm. 

Palpt blackish. First joint of antennae nearly the same 
length as the second, both darker in colour. Eyes join between 
the frontal triangle and the ocelligerous tubercle. Thorax with a 
wide median black stripe bordered by a_ yellow stripe, black 
bevond; sides yellow with a black stripe, pubescence yellow, 
golden yellow on the sides. Scutellum black. Abdomen similar 
to that of female with the exception of the first segment which is 
wholly black, only pale yellow on the sides, the black spots on 
second segment are larger and leave only a small yellow dot in 
their centre, the middle yellow spot on the two following segments 
is smaller ; under side yellow, black at apex. Legs black, only the 
middle femora and first joint of middle and posterior tarsi pale 
vellow. 


Chrysops rufitarsis, ~, Macquart. 


Dipt. exot Suppl. 111, p. 174 (1847) ; Ricardo, Ann. Mag. Nat. 
Hist. (7),-1x, p. 373 (1902). 

Thorax black. Abdomen white, at the base and apex black. 
Wings at the base, costal border and band brown. Length 4 lines, ~. 

Palpi brown. Face yellowish white with two shining black 
callosities. Forehead black. Antennae slender ; first joint hairy, 
dark testaceous ; the two others black. Thorax with blue reflec- 
tions, a spot of yellowish white hairs in front of the wings ; scutel- 
lum black. Abdomen: first segment black, yellowish white on its 
posterior border, second and third the same white colour ; the 
second with a small triangular spot resting on the anterior border ; 
the fourth on the anterior part white, sinuous, and posteriorly 
black, the fifth, sixth and seventh black ; under side as the dorsum. 
Legs black ; tibiae dilated, a little rounded on the outside, the 
posterior ones lightly ciliated; tarsi fawn coloured, the anterior 
ones black ; the two last joints of the intermediate and posterior 
pair black. Wings hyaline ; the cross-band hollowed out on its in- 
terior border ; veins pale, normal. From Java. Monsieur Payen’s 
collection. Macquart, Dipt. exot. Suppl. iii, p. 174. 

This type is unknown to me, but there is a male from Burma 
in the Brit. Mus. coll. which may perhaps belong to this species 
though there is no sign of a black spot on the second segment, the 
abdomen being almost silvery white or very pale yellow, black at 
base and apex. ‘The colouring of the large facial tubercles is red- 
dish brown, of the antennae chiefly reddish yellow, of the /egs 
more reddish brown than black ; the tibiae are all incrassate. Wings 
with a faint large hyaline sinus on posterior border, apical spot 
reaching apex uniform in width, apical border of band slightly 
concave. 


IgII.| G. Ricardo: The Oriental Tabanidae. 385 


Chrysops translucens, 2 , Macquart. 


Dipt. exot. (I), i, p. 162 (1838); Ricardo, Ann. Mag. Nat. 
Hist. (7), ix, p. 375 (1902). 

Black. Abdomen with the first and second segment white 
pellucid at the base. Femora brown, red below; tarsi red. Wings 
with a black band, excised on the interior border. Length 4 lines. 

Palpi brownish. Face pale fawn coloured, shining ; sides and 
cheeks shining black, with a spot of grey tomentum at the border 
ofthe eyes. Forehead denuded, entirely shining black. Antennae: 
first joint bare, pale fawn coloured, of ordinary length ; the second 
bare, same length as the first, brownish fawn coloured ; the third 
wanting Thorax denuded, shining black; a triangular spot of 
golden yellow hairs in front of the wings. Abdomen: first seg- 
ment transparent white, black on its posterior border, a little nar- 
rowed in the middle, reduced to nothing on the outer borders ; 
second silvery white with a large triangular black spot on the 
posterior border in the middle of which is a small dorsal white 
spot, transparent and triangular; the third shining black with a 
small dorsal white band, following that of the second segment, the 
others shining black ; under side : the first two segments transparent 
white, the second black on its outer borders. Femora brown- 
ish black above, posterior ones black ; tibiae brown, rather stout ; 
anterior tarsi brown; base of the first joint fawn coloured, inter- 
mediate and posterior ones fawn coloured ; the last joints brown. 
Halteres black. Wings hyaline; the outer border narrow, reach- 
ing the apex ; a little spot near the base and a cross-band brown; 
this latter hollowed out in the middle of the fifth posterior cell. 
From Java. Monsieur Serville’s collection. ‘This species is nearly 
related to Chrysops pellucidus, Fabr. and Wied., of which perhaps 
itis only a variety, but the latter differs in particular by the 
antennae being 1} lines long (though the third joint is wanting in 
C. translucens, they could not attain this length), by the abdomen 
and by the black legs. Macquart, Dipt. exot. (1), i, p. 162. 

One female from Kuala Lumpur (H. C. Pratt). Presented to 
Brit. Mus. coll. by London School Tropical Medicine. 

From a comparison of this specimen with the male and 
female specimens of Chrysops pellucida, Fabr.,in Brit. Mus. coll. the 
distinctions between the two species ¢ranslucens, Macq., and pellu- 
cida, Fabr., appear well marked. The face in Chrysops translucens is 
yellowish with a black spot on each side below antennae, the sides 
of face bordering the median line yellowish brown shining, becom- 
ing biackish over the oral opening, base of cheeks black. Forehead 
black, with broad band of grey tomentum below the frontal callus, 
which appears transverse, its anterior border sinuous, reaching 
the eyes. Antennae yellowish, not bare, the second one browner 
and the dark pubescence thicker. Abdomen distinguished at once 
from the Fabrician species by the white base of the first segment, 
and by the presence of the whitish or yellowish spots on second 
and third segment, as described by Macquart, conspicuous in the 


386 Records of the Indian Museum: [VonsIV, 


middle of the black colouring. Legs brownish, the anterior femora 
yellowish, the anterior tibiae curved and slightly incrassate, the 
posterior ones broad, all with short fringes of black pubescence, 
most conspicuous on the hind pair, tarsi pale yellowish on first 
joint and at base of other joints. Wangs similar to those of Chry- 
sops pellucida. Wength about 8 mm. 


Chrysops pellucida, Fabr. . 
(Plate xv, fig:-2:) 


Syst. Antl., 113, 11 (1805) ; Wied., Dipt. exot., 107, 8 (1821) ; 
id.,, Auss. zweifl. Ins., i, p. 206 (1828) ; Ricardo, Ann. Mag. Nat. 
Hist. (7)5,1x;-p.373) 902). 

Black, with the second segment of the abdomen at its base 
white, pellucid. Wings white: band black. Habitat Tranquebar 
(S: Eo India): 

‘In size and general appearance related to Chrysops dispar, of 
which itis perhaps only a variety. Antennae cylindrical, black, 
almost as long as the body. Head and thorax black, unspotted. 
Abdomen black, with the second segment at the base white, pel- 
lucid, which colour extends to the sides. Wings white; the wide 
middle cross-band black, paler at the border with a white spot. 
Legs black, tibiae compressed. Fabr., Syst. Antl., p. 113. 

Black, the second segment of the abdomen white pellucid ; 
wings with a black band, excised on the interior margin. 3 
Himes <9 + 

Undoubtedly a distinct species from Chrysops dispar. An- 
tennae yellowish at the base; their length hardly more than 14 
lines, and therefore the Fabrician text is hardly to be understood, 
unless he intended not to include the head. Head grey haired. 
Calli black. Thorax black, with golden yellow haired sides; 
breast sides blackish, with a golden yellow stripe under the root 
of wings. The transparent white part of the second segment of 
abdomen slopes downwards on both sides almost to the hind bor- 
der, so that a black triangular spot remains. The outer border of 
the wings black to the apex. In the legs the knees and base of tarsi 
are yellowish. In the Royal Museum and also a mutilated speci- 
men in the Fabrician collection. Wied., Auss. zweifl. Ins., i, p. 206. 

In the Indian Museum coll. there is a male from Trichinopoly, 
S. E. India, which exactly fits the above description. The length 
of antennae given by Fabricius as nearly equal to that of the body 
must be an error. 

@. Eyes brown with some blackish markings. Ocelligerous 
tubercle very prominent, blackish. Frontal triangle biack covered 
with grey tomentum. Jace black covered with dense grey tomen- 
tum, in the middle below the antennae is a large shining black 
tubercle and a small black spot on each side, the lower part of 
face below also black; pubescence in lower part of face whitish. 
Palpi small, blackish, with black hairs. Antennae long, slender, 


IQII. | G. Rrcarpo: The Oriental Tabanidae. 387 


about as long as the head and thorax together, first joint yellowish 
with pale hairs, its extreme apex and the second joint are obscure 
reddish with darker hairs, both joints about the same length, the 
third joint a little longer and blackish. Thorax black with some 
golden yellow hairs on its posterior borders, and a stripe of the 
same coloured hairs running across the root of wings at sides ; 
breast black with some grey tomentum. Scutellum black, shining. 
Abdomen black, the second segment pellucid, whitish, on its an- 
terior border, the black posterior border produced in the middle 
as a triangular spot which attains the posterior border of first seg- 
ment; under side yellowish on first two segments, then black. 
Legs black, the first two joints and part of the third one of 
middle and posterior tarsi pale yellow, all the tibiae slightly in- 
crassate, pubescence black. Wings hyaline, brown at base, and 
along fore border to the apex, the transverse brown band deeply 
incised in posterior margin, so that the fifth posterior cell is 
almost hyaline. Length 8 mm. 

There is a female in the Brit. Mus. coll. from unknown local- 
ity very much damaged which however is no doubt a specimen 
of this species. 


Chrysops alter, 2 , Rondani. 


Ann. Mus. Civ. Genova, vii, p. 44 (1875); Ricardo, Ann. Mag. 
IN Peet ists (7),.1% D. 27 3( LG02): 

Female length 5 mm. From Sarawak, Borneo. 

Antennae with the first joint reddish, the two following 
dusky black. Forehead in the middle narrowly greyish, with a 
very large anterior callus which is subrotund, shining black, 
taking up almost the whole width of forehead. Palpi reddish. 
Face shining black, at the sides narrowly and under the antennae 
pale pollinose. Thorax with scutellum black, the sides and the 
pleurae under the root of the wings yellow haired. Calyptra and 
halteres black. Wings with the base luteous; the costal border 
brown, the apex with a paler expanded brown colour; the black- 
ish band is beyond the middle of wing, posteriorly narrower. 
Legs: the anterior legs blackish, with the exception of the base of 
femora which are pale testaceous, the intermediate femora, the 
extreme apex of tibiae and the tarsi pale testaceous, the tibiae 
blackish; the posterior legs black, with a subapical ring of the 
femora, the apex of the tibiae and the tarsi luteous testaceous. 
Rondani, Ann. Mus. Civ. Genova, vii, p. 44. 

Type 2 in Genoa Museum was seen by me. It is a small 
species, the face wholly black, shining, convex, the frontal callus 
black, large, reaching the eyes, a narrow grey band divides it from 
the vertex. Antennae yellow. Abdomen which is in a bad state 
appears black, the first segment yellow anteriorly, and the second 
segment has a yellow band on its anterior border. Wings with 
one band, and the apex narrowly dark. 

Length 54 mm. 


388 Records of the Indian Museum. [VoL. IV, 


This species must be nearly alliea to Chrysops pellucida, with 
the face black, and to Chrysops translucens, with the first segment 
yellow anteriorly. Rondani omitted to describe the abdomen, 
which is now hard to distinguish as to colouring. 


Chrysops sinensis, 9 7, Walker. 
(Plate xv, fig. 6.) 


Dipt. Saund., i, p. 453 (1856); Ricardo, Ann. Mag. Nat. 
Hist (7) ..15 P1772 902)- 

In Brit. Mus. coll. 

Type (female) labelled ‘‘China.’’ Male from Haining, 
Chekiang, China, 1893 (Walker). 

In Indian Museum coll. females from Hankow, China. 

Specimens from Formosa were sent me some time ago by 
Dr. Kertesz for identification and proved to be this species, which 
is distinct from any species belonging to the Palaearctic Region 
and so far has not been found north of the Oriental Region 
boundary. 

A large robust-looking blackish species with the first two 
segments of abdomen largely yellow, and sometimes the remaining 
segments appear more yellow than black, with black median 
oblique spots, or the abdomen appears blackish with median grey 
triangular spots. Thorax black with distinct yellowish stripes. 
Wings with a cross-band and apical spot, the former irregular 
on its outer border becoming narrower as it approaches the fifth 
posterior cell. Face, legs and antennae yellowish. Length type 
and others 10o—11 mm. 

2. ace yellowish, the facial callosities reaching the oral 
opening are shining, tawny, cheeks with a small black spot on 
lower part near eyes and below blackish. Palpi tawny, rather 
bare and shining. Antennae tawny, the third joint blackish, the 
first two joints about equal in length, with very few yellowish 
hairs, the first joint slightly incrassate, the third as long as the 
first two joints together. Forehead covered with greyish yellow 
tomentum, the frontal callus large, protuberant, transverse, not 
reaching the eyes, vertex with the three ocelli blackish. Thorax 
black, somewhat shining, with two well-marked median grey 
tomentose stripes, sides with greyish yellow tomentum, breast 
black with some grey tomentum. Scutellum black. Abdomen in 
this species appears variable in colouring, in the type blackish, the 
first two segments largely yellow, the second segment with only 
two chevron-like black spots not quite joining above, their bases 
resting on the posterior border, the remaining segments blackish 
with narrow yellowish or yellowish grey borders expanding in the 
middle to triangular spots, and with traces of a yellow spot on 
each side of the third segment; in another specimen the yellow 
colour is more greyish and on the posterior segments is more 
extended leaving only black spots very similar to those in the 


IQII.] G. Ricarpo: The Oriental Tabanidae. 389 


second segment but not converging so much towards each other, 
the scattered pubescence on all the specimens yellowish. Legs 
reddish yellow, knees darker, base of hind femora blackish. 
Wings with the usual band and apical spot; the shape of the 
former is somewhat peculiar, projecting on its outer border towards, 
but not attaining, the fork of the third vein; in the first posterior 
cellit becomes concave, then slightly convex tillit reaches the fifth 
posterior cell, where the sinus is not very marked, and follows the 
fifth vein; it is narrow in width, only filling the discal cell, not 
encroaching on the basal cells at all, and filling the fourth 
posterior about half-way up; the basal cells have only dark shad- 
ing at their extreme base; the apical spot is the same width 
throughout, only crossing the anterior branch of the third vein at 
its apex. 

@. Similar. Eyes with the upper haif composed of larger 
facets. Abdomen in one specimen has the first segment almost 
entirely black, and the rest of abdomen reddish yellow with black 
oblique spots on each side, surrounding the grey triangular spots. 
Wings similar, centre of discal cell sometimes paler. 


Chrysops stimulans, o , Walker. 
(Plate xv, fig. 5.) 


Dipt. Saund., i, p. 73 (1850); ¢d., List. Dipt. v, Suppl. i, 
p. 265 (1854); Ricardo, Ann. Mag. Nat. Hist. (7), ix, p. 376 
(1902). 

Type (male) from East India, Saunders coll.; two males and 
one female from Pusa, Bengal, in Mr. Howlett’s coll 

A small blackish species, the female lighter in colouring, 
easily distinguished by the markings of the wings, having a round 
clear spot in the discal cell in the middle of the dark cross-band 
and a narrow apical spot in the inner border of which is a clear 
small oblong spot reaching the anterior branch of the third vein. 
Male with a black abdomen, marked with three spots on each of 
the anterior segments, the female with the abdomen greyish and 
two median united spots on the second and third segments. 
Length of males 6—8 mm., female 8 mm. 

@. Face covered with grey tomentum, the facial tubercles 
black shining, reaching the oral opening leaving the centre of the 
face with a broad grey stripe, a black, narrow spot on each lower 
cheek reaching from the eyes to the oral opening, pubescence 
below pale yellow. Palpfi yellowish brown with black pubes- 
cence. Antennae blackish brown, the first joint slightly incrassate 
with black pubescence, the second and third joints cylindrical, with 
less pubescence, the first two joints are about equal in length, the 
third a little longer than either. Frontal triangle grey, sub- 
triangular. Eyes only meeting at one point beyond, vertex large 
with the ocelli distinct. Thorax blackish with grey tomentum 
Sometimes appearing as stripes and with black pubescence. 


a 


390 Records of the Indian Museum. [VoL. IV, 


Scutellum black. Abdomen black, on the second segment appears 
a large pale vellowish almost square spot and a minute triangular 
greyish spot in centre on posterior border, on third and fourth 
segments appear three small triangular greyish spots situated on 
the posterior borders of segments; under side black with yellow 
spots. Legs black, the middle tibiae yellow, black at extreme apex, 
pubescence of legs black. Wungs hyaline with a dark brown 
base, costal border reaching to the apex, and a median band, the 
brown colouring at base extends beyond the middle of both basal 
cells, leaving a narrow hyaline band between them and the cross- 
band which is broad extending into the apical cell, becoming 
paler in the fifth posterior cell, its upper border irregular, at the 
base of branches of third vein it is produced, reaching towards the 
apical spot and thus giving rise to the clear spot or incision men- 
tioned above: the apical spot extends over the anterior branch of 
third vein. 

2. Facial black tubercles are smaller. Forehead covered 
with grey tomentum, the frontal tubercle black, transverse, but 
small, not reaching eyes. Thorax grey with three very distinct 
black stripes. Scutellum identical. Abdomen greyish, the two 
black spots uniting in centre on first and second segments well 
marked, on third segment appears a zigzag black band on its 
anterior border, and on following segments a small ill-defined 
black spot on each side of median line; the abdomen of the only 
female specimen is not in very good preservation; under side 
greyish with black markings. Legs yellow, knees and tarsi black- 
ish. Wangs similar to those of male. 

The male from Persian Gulf mentioned in Ann. Mag. Nat. 
Hist. (7), ix, p. 376, appears a different species, the abdomen 
having only median grey spots ; it is probably a new species. 

This Walker species bears a strong resemblance to the 
Palaearctic species Chrysops punctifera, Loew, récorded from Syria, 
in the wing-markings and in the abdominal markings, especially 
of the male, but it is distinguished by the slightly incrassate first 
joint of antennae, and larger frontal triangle in the male, and also 
in the female by the more greyish colour of the abdomen. 


Chrysops indiana, o 2 , Ricardo. 


Ann. Mag. Nat. Hist. (7), ix, p. 379 (1902). 


One male from Nilghiri Hills, 88, 112 (Hampson). 

One female from Khasi Hills district, India, 96, 135 
(Chennell, 1878). 

A species allied to C. dispar, Fabr., but in the wing approach- 
ing C. fasciatus, Wied.; it is larger and more robust than 
C. dispar, Fabr. 

Yellow, with a black bifid stripe on the second segment, 
enclosing a small, almost round, yellow spot, but the black does 
not join on the posterior border. 


IQIt. | G. Rrcarpo: The Oriental Tabanidae. 391 


@ @ (types). Face yellow, with yellow hairs on the cen- 
tral stripe and at the sides. Antennae long and slender, nearly 
equal in length to the head and thorax, all the joints about equal 
in length; yellow, the third joint darker, the first clothed with 
long black hairs, the second with shorter ones, the third bare. 
Palpi yellow. Thorax and scutellum brownish, with yellow pubes- 
cence, thicker at the sides, the breast the same colour. Abdomen 
light yellow, the second segment paler; the first segment with a 
natrow black band on the posterior border, the second with 
a black stripe which begins in the centre and then divides, send- 
ing out a branch on each side reaching to the hind border, and 
there extending outwards till it ends in a point; a small, yellow, 
oblong spot is thus left in the middle, surrounded by the black, 
excepting on its posterior border ; the black stripes are continued 
very faintly on the third segment; the sides of abdomen have 
short black pubescence; the under side yellow, with three faint 
brown stripes on the third segment. Legs yellowish brown, with 
short black pubescence; the tibiae dilated, especially the anterior 
and middle pairs. Wings clear, with brown colouring at the 
extreme base only filling the basal cells one-third of their length, 
continued along the fore border to the apex, and as a transverse 
band which attains the posterior border just contiguous to the 
anal cell; it is a little fainter in colouring in the fifth posterior 
cell; its apical border is straight, also the inner one as far as the 
anal cell. 

Length 113 mm. 

The female is identical. 


Chrysops fasciata, ¢, Wiedemann. 
(Plate xv, fig. 7.) 


WMipi -eXOt 1. “Oa £03 (1S2L) +04... uss. zwelll.. Pns:, 1 p.-198 
(1828); Doles., Natuurkund. Tijd. Nederl. Ind., xvii, p. 84 (1858) ; 
Ricardo, Ann. Mag. Nat. Hist. (7), ix, p. 375 (1902). 

Thorax black; abdomen at the base white, in the middle 
black, with the apex ochraceous; wings at the base, costal border 
and band brown. 4} lines, @ , from Java. 

Antennae brown, the last joint deep black at apex, face 
golden yellow with a brownish black triangular tubercle. Thorax 
with golden yellow tomentum; breast sides with thick golden 
yellow silky pubescence. Abdomen with the first two segments 
white, the last one black on its posterior border, the third black 
with ochre-brown median and side spots, which do not attain the 
side border, the fourth and fifth ochre-brown with black side 
borders, the sixth and seventh wholly ochre-brown. Wings very 
hyaline, the usual band hardly incised in the inuer border, Legs 
brown, femora darker, the tarsal joints at base very pale, the fore 
tibiae and tarsal joints at base blackish; the posterior tibiae fringed 
with black hairs. In Westermann’s collection. Wiedemann, Auss. 
zweifl. ‘Ins., i, p. 198. 


392 Records of the Indian Museum. [Vou. IV, 


Doleschall records this species from Amboina. In Brit. Mus. 
coll. a female from Malay States (H. C. Pratt), presented by 
London School of Tropical Medicine; and two females from Ross 
Island, Andaman Islands, May 1904 (Col. Bingham). In Indian 
Museum coll. a female from Assam and another from Sibsagar, 
Assam. 

In London School of Tropical Medicine coll. a female from 
Sarawak, Borneo (A. R. Wellington). 

A species with the abdomen pale yellow at base, black in the 
middle and reddish yellow at apex. Legs and face blackish. 
Wings with the dark cross-band hardly incised on inner border 
though sometimes paler. 

Length 8}—12 mm. 

Face yellowish or golden yellow, with the middle of face 
shining black having only a short yellow median stripe below the 
antennae, base of cheeks shining black. Beard yellow. Palpi 
yellowish red, smooth, long and pointed with some black hairs. 
Antennae yellowish, cylindrical, the second joint browner with 
thicker pubescence, the third joint black except at base, longer. 
Forehead black with a broad band of yellowor grey tomentum, the 
frontal callus large, reaching eyes, protuberant. Thorax black, with 
golden yellow pubescence at sides and on posterior border, and 
with pale yellow appressed pubescence on dorsum. Scutellum 
blackish. 

Abdomen with the first two segments pale yellowish, the first 
segment black beneath the scutellum, the second segment with a 
narrow posterior black border, the third segment blackish some- 
times with obscure yellowish or reddish markings, the remaining 
segments reddish yellow. Legs blackish brown, the fore t biae 
black, swollen, the middle and posterior tibiae almost wholly pale 
yellow, the posterior femora incrassate with fringes of black hairs, 
pubescence of legs wholly black. Wangs with a large broad trans- 
verse band, broadly brown on costal border to apex and at base 
brown, the band with almost straight borders, not incised on 
posterior border though sometimes paler in the filth posterior cell, 
reaching to the anal cell, the apex of which it entirely fills. 


Chrysops fixissima, @ , Walker. 


Proc. Jinnt Sockvsi -p: 112 (1850). Ricarco =, AnaseMetes 
Nat. Hast: (7); -15 0: 37 0u(rae2): 

Chrysops unizonatus, Rondani, Ann. Mus. Civ. Genova, vii, 
Pp. 459 (1873); Ost. Sacken, Berlin. Ent. Zeit., xxvi, p. 97 (1882). 


The type is not to be found in the British Museum collection, 
but only the variety. Cclonel Yerbury has a note on one of the 
specimens to the effect that the species is ‘‘ Rare.”’ 

Type (var., female) from Sarawak, 68, 4 (Saunders). 

Two trom Sandakan, Brit. N. Borneo, 98, 38 (D. Cator), and 
one, from same locality, 95, 134; one from Singapore, 96, 114 


IgII.] G. Ricarpo: The Oriental Tabanidae. 393 


(Flower); one from Sarawak, 56, 14 (Saunders); two from ‘Trin- 
comalee, 2, 10 g0O—-10, 9, 91 (Yerbury). 

Walker describes the var. ‘‘ with two bands, one black, the 
other brown.” I should rather describe it as having one black 
band only, on the posterior border of the second segment. It is 
related to C. fasciata, but distinguished from it by the facial 
tubercle being yellowish brown, becoming black only at its upper 
corners ; thelegsare also paler. The species described by Rondani 
as C. unizonatus is a synonym of this, and not of C. signifer, 
Walker, as suggested by Osten Sacken, who states that the face in 
the Rondani species is yellow, which agrees with the colouring 
of the face of this species (see Osten Sacken, in Berlin. Ent. Zeit., 
XXvi, p. 97). Ricardo, Ann. Mag. Nat. Hist. (7), ix, p. 376. 

Length of type 11 mm., of others g—11} mm. 

When further material is available, this species may prove to 
be only a subform or identical with Chrysofs fasciata, Wied., 
the abdomen with only one black band on the posterior border of 
second segment and the more largely yellow colouring of the face 
appearing the only differences. 


Chrysops signifer, ~, Walker. 


Proc. Linn. Soc. London, v, p. 276, 1861; Ricardo, Ann. 
Mag. Nat. Hist. (7), ix, p. 376 (1902). 

There is only the type, a male, in Brit. Mus. coll. from 
Batchian, but Osten Sacken records males and females from the 
Philippines and states they vary very much in size. Ost. Sacken, 
Berlin. Ent. Zeit., xxvi, p-'97, 1882. 

The species is distinguished from Chrysops fixissima and 
fasciata by the shape of the second band on the third segment, 
which is not present in Chrysops fixissima and Chrysops fasciata, 
in this species it is concave in the middle, neither of the bands 
reach the sides, but the third and fourth segments are bordered 
with a narrow black stripe. Legs in the type are yellowish brown, 
the hind pair darker, tibiae slightly incrassate. Wings as in 
Chrysops fasciata. 

Length of type 9} mm. 


Chrysops cincta, Bigot. 


Mem. Soc. Zool. France, v, p. 602 (1892); Ricardo, Ann. 
Mag. Nat. Hist. (7), ix, p. 378 (1902). 

? Chrysops clavicrus, Thomson, Eugen. Resa, Dipt., p. 452 
(1868); Ricardo, Ann. Mag. Nat. Hist. (7), ix, p. 373 (1902). 

Antennae as long as the head and thorax together, black, the 
first segment (joint) pale yellow, the second the same length, 
palpi whitish yellow; face and forehead covered with a yellowish 
tomentum, under the antennae a heart-shaped spot, an oval callus 
at base of forehead, another transverse one at vertex, all shining 
black; thorax black, a little yellowish down at the sides: 


394 Records of the Indian Museum. [Vor. IV, 


scutellum black; abdomen black, the segments very narrowly 
bordered with yellowish white, a large band of the same colour at 
base of second segment; coxae black, femora reddish, the posterior 
ones black exteriorly, tibiae dilated, reddish, blackish exteriorly, 
tarsi reddish with the apex blackish; squamae greyish, club of 
halteres brown; wings whitish, blackish exteriorly, from the base to 
the apex included, with a large median cross-band of the same 
colour. PhilippineIslands. One specimen. Bigot, Mem.Soc. Zool. 
France, v, p. 602. 

I have examined the type (female) from the Philippines lent 
me by Mr. Verrall; it is darker than the three other species with 
black bands, and the yellow colour of the ahdomen is not 
diaphanous; it is nearly allied to C. fascratus, Wied., and 
C. stgnifer, Wik., but the abdomen is darker and the black bands 
broader. 

I believe it to be the same as C. clavicrus, Thomson, from 
Malacca; and if this proves to be correct, Bigot’s name must 
be sunk. 

The original description may be amended thus :— 

Face black, yellow at the sides, and with a central short 
yellow stripe. Abdomen with the first segment pale yellow anda 
narrow black posterior border; the second is pale yellow on its 
anterior border; the black band posteriorly is nearly equal in 
width to the yellow colour; the third segment is almost wholly 
black, with only a narrow yellow border anteriorly; the two black 
bands join at the sides, but the second band is entirely straight 
on its posterior border, thus differing from C. szgnifer, WIk., 
which is deeply indented in the centre; the fourth, fifth, sixth 
and seventh segments are reddish yellow, with obscure black 
square spots in the centre of the fourth and fifth. The wings are 
clear, dark at the base and along the fore border to the apex, 
and with the usual transverse band which extends to the posterior 
border. Ricardo, Ann. Mag. Nat. Hist. (7), ix, p. 378. 

Black. Antennae, head and palpi yellow, legs brown, anterior 
femora and tarsi testaceous, anterior tibiae club-shaped, fusiform ; 
wings hyaline, with the marginal border and an oblique median 
band brown. Abdomen at apex brownish, at base honey-yellow, 
pellucid, with black median band. 

9. Length 10mm. Malacca. 

Related to Chrysops rufitarsis, Macquart, Dipt. exot. Suppl. 
iii, p. 174, with the anterior tibiae club-shaped, but in the 
colouring of the abdomen and with the median band of wing not 
incised, distinct. Head almost hemispherical, the width of thorax, 
the large eyes bare, ocelli placed on a triangular callus, forehead 
golden haired, with a large shining callus above the antennae; face 
golden haired, the facial tubercle sub-protuberant, heart-shaped, 
shining, the cheeks golden haired, proboscis deflexed, length of 
the head brown-black; palpi yellowish, deflexed, conical, awl- 
shaped. Antennae as long as the head and thorax, porrect, 
filiform, with very short black hairs, separated at the base, black, 


IQII.] G. Ricardo: The Oriental Tabanidae. 395 


the first joint yellow, the second equal in length, lineal, the third 
one and a half longer than the second, 5 ringed. ‘Thorax sub- 
glabrous, golden haired at sides and beneath wings; black, at the 
base above scutellum golden haired, scutellum large, transverse- 
triangular. Halteres dull yellow, the large club black. Wings 
with the fore border widely brown, and the band brown, narrower 
before the base, elsewhere wide, transverse, passing behind the 
middle of discal cell. 

Abdomen with fine yellow pubescence, subopaque, the first 
segment pellucid yellow with a narrow band before the apex, the 
second yellow, denuded, posteriorly black, the third with the black 
part wider, with a narrow yellow basal part, fourth, fifth and 
sixth brown, the fourth with an obsolete discai brown spot; 
under side brown, at the base yellow, the fourth segment with a 
biown median spot. Breast black, a little shining, with fine 
scanty pubescence. Legs with fine short brown pubescence, 
anterior coxae dull testaceous, brown at apex, the posterior ones 
black, femora brown, the anterior pair dull testaceous, anterior 
tibiae stout, club-shaped, brown, the intermediate ones less stout, 
dull testaceous, brown in the middle, posterior femora brown, 
tarsi yellow with the apex brown. Thomson, Eugen. Resa, Dip- 
tera prt452: 

This latter description is given here with a few unnecessary 
particulars omitted, as I believe it to be the same species as 
Bigot described. 


Chrysops flavocincta, @ , Ricardo. 
(Plate sy; tise A.) 


Ann. Mag. Nat. Hist. (7), ix, p. 380 (1902). 


Type (female) and another female from Khasi Hills, Assam, 
97, 82 (Heyne) ; two females from North Khasi Hills, lower ran- 
ges, 96, 135 (1878, Chennell); one female from Sarawak, 56, 44 
(Wallace) ; one female from Trincomalee, 54 (Yerbury). 

Black. Abdomen with a yellow band on the anterior half of 
the second segment. 

Face black, shining, with a grey tomentose stripe just below 
the antennae and an obscure yellowish spot where the usual stripe 
begins; a line of grey tomentum divides the cheeks from the 
upper part of the face; forehead with the usual black tubercles, 
which are large, a narrow band of grey tomentum divides them. 
Antennae rather long, yellow, the second and third joints darker ; 
the first two joints pubescent, the third bare, Palpfi yellow. 
Thorax black and shining, with traces of a broad grey stripe, some 
white pubescence on the dorsum and on the posterior border, and 
yellow hairs at the sides. Scutellum black. Abdomen black, with 
some white pubescence, with a yellow band on the anterior half 
of the first two segments; that on the second is widest and 


396 Records of the Indian Museum. [Vou. IV, 


concave in the middle, becoming broader at the sides; the under 
side yellow at the base and black on the apex. Legs black, the an- 
terior and middle femora and the middle and posterior tarsi yellow. 
Wings clear, the dark colouring hardly perceptible at the extreme 
base ; along the fore border it is narrow and only reaches the band, 
the apex being quite clear ; the band hardly attains the hind bor- 
der and becomes paler in the fourth posterior cell, only just con- 
tinuing into the fifth; in some of the specimens it does not reach 
beyond the fourth and never attains the anal cell ; it is straight on 
both its borders. 

Length 8} mm. 

One of the specimens from Borneo does not measure more 
than 64 mm. Ricardo, Ann. Mag. Nat. Hist. (7), ix, p. 380. 

In Indian Museum coll. are females of this pretty little species 
from Pallode, 20 miles N. E. of Trivandrum, Maddathorai, west 
base of W. Ghats, all in Travancore. The greyish stripe or band 
below the antennae and a narrow border between the eyes and the 
facial tubercles are in some otf these specimens almost golden yel- 
low. The species is easily recognized by the long cylindrical 
antennae, all the joints about equal in length, the black shining 
face, and the attenuated band of wings usually not attaining the 
fifth posterior cell, its outer border is rather concave in these 
specimens. 

[This species, which is very common on the west side of the 
W. Ghats in Travancore in November, resembles C. dispar (p. 382) 
in habits —N. ANNANDALE. | 


Chrysops flaviventris, ? , Macquart. 


Dipt. exot. Suppl. i, p. 172 (1845); Ricardo, Ann. Mag. Nat. 
Hist. (7), ix, Pp. 373 (1902). 

Thorax black. Abdomen yellow with a spot and the apex 
black. Legsred. Wings with margin and band brown. Length 
33 lines, 2. 

1, Related to C. tvanslucens. Face shining black, the upper part 
fawn coloured. Forehead brownish black, shining. Antennae 
wanting. Thorax shining black, with a large spot of fawn-coloured 
tomeutum in front of the wings and prolonged below; scutellum 
black, bordered with testaceous colour. Abdomen pale yellow, 
the second segment with a blackish chevron, the last one black. 
Posterior femora brown. Wings on posterior border and the 
cross incised band brownish. From India. M. Bigot’s collection, 
Macquart, Dipt. exot. i, p. 172. 

This type is unknown to me. From the description it is not 
clear whether an apical spot is present on the wing, but as it is 
said to be related to C. tvanslucens, Fabr., which possesses one, it 
may be assumed to have one. From the description it might 
possibly be a specimen of Chrysops dispar, Fabr., with the black 
stripe almost obsolete. It is not included in the table. 


IQgIt. | G. Ricarpo: The Oriental Tabanidae. 397 


Chrysops flavipes, Meig. 


Klass., i, p. 159, pl. ix, fig. 13 (1804); Villen, Ann. Soc. En- 
tom. France, Ixxiv, p. 310 (1905). 

Chrysops perspicillaris, Loew, Neue Beitr., iv, p. 25 (1856) ; 7d., 
Verh. zool-bot. Ges. Wien, viii p. 633 (1858); Schiner, Fauna 
Aust. Dipt., i, p. 41 (1862) ; Gobert, Mem. Soc. Linn. Nord France, 
1881, p. 48 (1881); Pand., Revue d’Entom., ii, p. 226 (1883); 
Griff., Boll. Mus. Zool. et Anat. comp. Torino, x, No. 218 (1895) ; 
Ricardo, Ann. Mag. Nat. Hist. (7), ix, pp. 426, 430 (1902). 

One female from Baluchistan in Indian Museum coll. rather 
pale in colouring but identical with the species described by Loew, 
now stated by Villen to be identical with the Meigen type of C. 
flavipes seen by him in the Paris Museum. ‘The species is widely 
distributed, being recorded from Central and South Europe, N. 
Africa and Asia Minor. 


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Chrysops, Meig. Sc 


Corizoneura, Rond. 


Diachlorus, Macq. 


Diatomineura, Rond. 
Gastroxides, Saund. 


Page 
365, 374 
alter, Rond. et 8 O7aA 
cincta, Bigot 393 
designata, Ric. 383 | 


Haematopota, Meig. 


Page 

Haematopota cilipes, Bigot 360 
cingalensis, Ric. 351 

cingulata, Wied. 326 

dissimiliss Rice f.- 331 

fasciata, Ric. 358 


dispar, Fabr. es OOn| 
fasciata, Wied. 56 Bu 
fixissima, Wlk. 392 | 
flavipes, Meig. 397 
flaviventris, Macq. 396 | 
flavocincta, Ric. Jo. -Sho)3 | 
indiana, Ric. S06. Shex0) 
manilensis, Schiner .. 377 
mlokosiewiczi, Bigot 378 | 
pellucida, Fabr. 386 
rufitarsis, Macq. 384. 
signifer, Wlk. eS OS 
sinensis, WIk. 388 
stimulans, W1k. 389 | 
translucens, Macq. 385 | 
364, 365 | 
longirostris, Hardw. 365 | 
taprobanes, Wlk. .. 367 | 
varipes subsp., Ric. 367 
321, 364 | 
flavipennis, Macq. .. 364 
364, 365 
365, 370 
ater, Saund. Se) SV) 
ornatus, Bigot 3714 
So0 ee 
annandalei, Ric. 335 
assamensis, Ric. 343 
atomaria, WIk... 332 | 
bilineata, Ric. .. 350 
borneana, Rond, 329 
brevis, Ric. 349 
cana, WIk. 330 


fuscifrons, Austen 357 

immaculata, Ric. 359 

inconspicua, Ric. 358 

indiana, Bigot .. 328 

irrorata, Macq... 352 

javana, Wied. 340 

lata Rict 344 

latifascia, Ric. .. 356 

limbata, Bigot .. 325 

lunulata, Macq. 354 

marginata, Ric. 347 

pachycera, Bigot 336 

punctifera, Bigot 327 

roralis, Fabr. 323 

rubida, Ric. 338 

sinensis, Ric. 345 

singularis, Ric... 339 

tessellata, Ric.. 348 

unizonata, Ric. . 332 

validicornis, Ric. 333 

| Mycteromyia, Phil. 365, 373 
nigrifacies, Bigot 373 

Neotabanus, Ric. .. 321, 363 
ceylonicus, Ric. .. 363 
| Pangonia, Latr. : a. 28g 05 
amboinensis, Fabr. .. 368 
Rhinomyza, Wied. .. 365, 369 
fusca, Wied. OO 

Silvius, Meig. ae 365, 373 
indianus, Ric. re sy ie! 
Udenocera, Ric. 321, 361 


brunnea, Ric. 


aa 


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Peco eR res ULrLONS TO THE BRAUN A: OF 
VUNNAN BASED ON COLLECTIONS 
MAD BV: |. COCGCIN BROWN. 
Bioc:, .r909—19O27G.! 


PART VII.—TABANIDAE. 
By GERTRUDE RICARDO. 


The Tabanidae sent me for identification by Dr. Annandale 
are few in number, comprising the following species :— 


TABANUS, Linné 


Tabanus orientis, Walker. Two females from Man Hsien. 


Tabanus monotaeniatus, Bigot. Two females from the 
same locality, rather smaller in size than usual, the abdomen 
being narrower, and reddish in colour. 


HAEMATOPOTA, Meigen. 


Haematopota lata, Ricardo. Four females from Man Hsien. 


CHRYSOPS, Meigen. 


Chrysops mlokosiewiczi, Bigot (Chrysops striatus, Wulp). 
A series of females from between Teagvueh and Tali Fu (J. Coggin 
Brown). The frontal callus is reddish yellow, not black. This 
species has a wide distribution in the Palaearctic and Oriental 
Regions, being recorded from China, Japan and Persia. 


Chrysops designata, Ricardo (p. 383, anfea). One female. 


For references to these species see Rec. Ind. Mus., vol. iv, 
No. vi, IQII, containing a revision of the species of Tabanus 
from the Oriental Region, and the same publication, vol. iv, No. 
viii, for species of Chrysops and other genera of Tabanidae. 


1 The previous papers of this series appeared in Rec. Ind. Mus., v, p. 193 
(1910), and vi, pp. 13 and 215 (1911). 


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EXPLANATION OF PLATE XV. 


1.—Chrysops designata. 


Zi 9 
Star 2) 
ieee oa) 
ear a) 
o— 29 


icra + BS) 


pellucida. 
dispar. 
flavocincta. 
stimulans. 
sinensis. 
fasciata. 


EK eC 


+a Ve 


mec id= Mus. Vol. IV, rorr. Piste 


CHRYSOPS, 


H.G. Herring. Bemrose.L*4, Derby. 


EXPLANATION OF PLATE XVI. 


8.—Udenocera brunnea. 
9.—Gastroxtdes ater. 
10.—Haematopota latifascia. 


II.— Br atomaria. 
2 BS unmizonata, o. 
13.— i unizonata, @ « 
14.— » tessellata. 
15.— i. cingalensts, 


to. ae brevis. 


ZV 
Rec. Ind. Mus. Vol. IV, fort. Plate 


UDENOCERA , GASTROXIDES, HAEMATOPOTA. 


H.G Herring. Bemrose.L4, Derby. 


EXPLANATION OF PLATE XVII. 


. 17.—Haematopota fuscifrons. 
18.— i rubida. 
19.— re indiana. 
20.— ; punctifera. 
21.— a annandalet. 
22.— Me dissimilis. 
23.— validicornts. 
24.— voralis. 
25.— : javana. 

6.— lata. 


“<3 Vain a 


Rec. Ind. Mus. Vol. IV, rorr. Plate #. 


HABRMATOPOTA. 
H.G. Herring. Bemrose.Lt4, Derby. 


ny 


" 1 ‘ ny } yi " 
ef Hy sm " 
} we avs ie} i 


i 


ca 


ngs we = 
wht 
ay 
< 


7.328029 


EXPLANATION OF PLATE XVIII. 


27 
5 B= e sinensts. 
See: coal <5 assamensis. 
ae OO sess * tvrorata. 
eet Siac inconspicua. 
iA oe en >» marginata. 
ie. 33-5 ss tmmaculata. 
BS ABA the aa sy bilineata. 


Rec. Ind. Mus. Vol. IV, Iori. ; Plate +. 


own, I ee seo ee 


HAEMATOPOTA. 


H.G. Herring. Bemrose.L4,Derb 


- Culicidae and Corethridae in the Indian Museum. Mdscellanea :—Measurements 
of the skeletons of two large Indian elephants in the Indian Museum. The 
young of Aclurus fulgens.. Some Batrachia recently added to the collection of 
the Indian Museum. Breeding habits of Tylototriton verrucosus. The occurrence 
of Rhinodon tvpicus at the head of the Bay of Bengal. Note on Ephydatia 
meyent (Carter). 


Part IV.—Aquatic animals from Tibet, I. Aculeate Hymenoptera in the Indian 
Museum, I. Indian Psychodidae. A new species of mouse from the Madura 
District, Madras. Some Cleridae of the Indian Museum. The Fauna of Brackish 
Ponds at Port Canning, Lower Bengal, XII. A new species of Saw-Fish captured 
off the Burma Coast. A new Sting Ray of the genus Tvygon from the Bay of 
Bengal. New Microlepidoptera from India and Burma. Chrysomelid Beetles in 
the Indian Museum. Six new Cicindelinae from the Oriental Region. A new 
Slug from Tibet. 


Part V.—Revision of the Oriental Leptidae. Revised and annotated Catalogue of 
Oriental Bombylidae. 


Vol. III, 1909. 


Part I.—The races of Indian rats. 


Part II.—¥reshwater Sponges, X. Aquatic animals from Tibet, II. Some amphibious 
Cockroaches. Quelques nouvelles Cécidomyies des Indes. New land and marine 
_ shells from Ceylon and §. India. Two new species of Cavanx from the Bay of 
Bengal. Some little known Indian Ophidia. Some forms of Dipsadomorphus. 

A pelagic Sea-Anemone without tentacles. Rhynchota Malayana, II. 


Part III.—The Neuroptera in the Indian Museum. New Indian Leptidae and Bom- 
bylidae. The Trichoptera in the Indian Museum. New species and varieties 
of Freshwater Crabs, I—3. Lizards from Travancore. Three new Cicindelinae 
from Borneo. The relation between fertility and normality in Rats. A 
Barnacle of the genus Scalpellum from Malaysia. The Hemipterous family 
Polyctenidae: Freshwater Sponges, XI. Two new shells from S. India. A 
new genus of Phylactolaematous Polyzoa. Miscellanea :—Major Wall on some 
forms of Dipsadomorphus. Notes on Indian Batrachia. Notes on Indian Fresh- 
water Fish. Field notes on Indian Insects. The habits of Indian King-Crabs. 
The rate of growth in Conchoderma and Lepas. Large colonies of Hislopia lacus- 
tvis. Byrvanchiocerianthus imperatory von der Kiiste von Oman und Baluchistan, 


Pavt IV.—A_ minute Hymenopterous insect from Calcutta. The Insect Fauna of 

Tirhut, No. 1. New species of Botia and Nemachilus. New Oriental Sepsinae. 

A new species of Hvedericella from Indian lakes. New-species and varieties of 

freshwater crabs, 4. Some new or little known Mygalomorph spiders from the 
Oriental region and Australasia. : 


Vol. IV, 1910-1911. 


No. I.—Second report on the collection of Culicidae in the Indian Museum. 


Nos. II and III.—The Indian species of Papataci Fly (Phlebotomus). Taxonomic 
values in Culicidae. 


No. IV.—Revision of the Oriental blood-sucking Muscidae. 
No. V.—A new arrangement of the Indian Anophelinae. 


"No. VI.—A revision of the species of Tabanus from the Oriental Region, including 


notes on species from surrounding countries. 


No. VII.—New Oriental Nemocera. Miscellanea :—Synonymy in Corethrinae. 
Indian Phlebotomt. 


Vol. V, 1910. 


Part I.—The Hydroids of the Indian Museum, I. Freshwater Sponges, XII. New 
Shells in the Indian Museum from Burma, Siam and the Bay of Bengal. Mate 
rials for a revision of the Phylactolaematous Polyzoa of India. Studies on the 
aquatic Oligochaeta of the Punjab. An undescribed Burmese Frog allied to Rana 
tigrina. Miscellanea :—The occurrence of Vultur monachus in Calcutta. An 
albino Owl. ‘‘ Matla bengalensis’’ ; a correction. 


Pavt II.—Description d’Ophiures nouvelles provenant des derniéres campagnes de 
‘‘l’Investigator’’ dans 1l’Océan Indien. Description d’Holothuries nouvelles 
appartenant au Musée Indien. The races of Indian rats, II, A new species of 
Scalpellum from the Andaman sea. Five new species of marine shells from the 
Bay of Bengal. Fish from India and Persia. 


t a r te 
= ihe Ay Spry 


+ 


Paints (se: 


ieee rie A ae 


1 


At 
“ 


Ss 


4 
4 
wf 


a 


Pavt IIJ.—A new genus of Psychodid Diptera from the Himalayas and Travancore. 
The Indian barnacles of the subgenus Smilium, with remarks on the classification 
of the genus Scalpellum. A subspecies of Scutigerella unguiculata, Hansen, found 
in Calcutta. The distribution of the Oriental Scolopendridae. Decapoda in 
the Indian Museum, I. A new species of Nemachilus from Northern India. 
The larvae of Tovorhynchites immisericors, Wik. A South Indian frog allied 
to Rana corrugata of Ceylon. Contributions to the fauna of Yunnan, 
Introduction and Part I. Miscellanea :—The Darjiling skink (Lygosoma sikkim- 
ense). Cockroaches as predatory insects. Note on Aedeomyia squammipenna, 
Arribalzaga. Named specimens of Chrysomelidae in the Indian Museum, Two 
barnacles of the genus Dichelaspis new to Indian seas. Slugs from the Eastern 
Himalayas. 


Pavt IV.—Indian Microlepidoptera. Some aquatic oligochaete worms commensal in 
Spongilla carteri. Bothrioneurum tris, Beddard. Nudibranchs from the Indian 
Museum. The classification of the Potamonidae (Telphusidae). Catalogue of the 
pheasants, peafowl, jungle fowl and spur fowl in the Indian Museum. Species of 
Palaemon from South India. Alluaudella himalayensis, a new species of degener- 
ate (d') cockroach, with an account of the venation found in the genera Cardax 
and Alluaudella. Rhynchota Malayana, III. 


Vol.oVi) 1919. 


Pavt I.—A Rhizocephalous Crustacean from fresh water and on some specimens of 
the order from Indian seas. Decapoda in the Indian Museum, I]. Contri- 
butions to the fauna of Yunnan, Parts II to V. Pedipalpi in the Indian 
Museum, I and II. Six new species of shells from Bengal and Madras. Mis- 
cellanea :—Fleas from India and China. Flies found associated with cattle in 
the neighbourhood of Calcutta. Mosquito sucked by a midge. Large egg laid 
by a beetle. 


Pavt II.—Some sponges associated with gregarious molluscs of the family Vermetidae. 
Aquatic animals from Tibet, III. Cyprinidae from Tibet and the Chumbi Valley, 
with a description of a new species of Gymnocypris. New species and varieties 
of Crustacea Stomatopoda in the Indian Museum. The development of some 
Indian Ascalaphidae and Myrmeleonidae. Miscellanea :—The occurrence of 
Dactylopius cityi, Risso, in the Himalayas, Note on Aquatic Rhynchota. 


Pavt III,—Nouveaux Chironomides de |’ Indian Museum de Calcutta. 


Pavi IV.—Indian Isopods. Systematic notes on the Ctenostomatous Polyzoa of 
fresh water. Some aquatic Oligochaeta in the Indian Museum. Contributions 
to the fauna of Yunnan, Part VI. Asiatic species of Crustacea Anostraca in the 
Indian Museum. Freshwater sponges, XIII. Méscellanea:—Synonymy in 
Corethrinae. The distribution of the different forms of the genus Jb/a. 


Other Publications edited and sold by the Superintendent of the India 
Museum (also obtainable from Messrs. Friedlander & Sohn) issued 
by the Director of the Royal Indian Marine, 


Illustrations of the Zoology of the R.I.M.S. ‘‘ Investigator ’’ 1892. Fishes, Plates I to VIE 

_ Crustacea, Plates I to V, 1894. Fishes, Plates VII to XIII. Crustacea, Plates VI to VIII. Ech 

-  inoderma, Plates I to ida 1895. Echinoderma, Plates IV and V. Fishes, Plates XIV to XVI, 

ag Crustacea, Plates IX to XV, 1896. Crustacea, Plates XVI to XXVII, 1897. Fishes, Plate XVII, 

_ Crustacea, Plates XXVIII to XXXII. Mollusca, Plates I to VI, 1898. Fishes, Plates XVIII to 

XXIV. Crustacea, Plates XXXIII to XXXV. Mollusca, Plates VII and VIII, 1899. Fishes, 

_. Plates XXV and XXVI. Crustacea, Plates XXXVI to XLV, 1900. Fishes, Plates XXVII to 

p XXXV. Crustacea, Plates XLVI to XLVIII. Index, Part I, 1901. Crustacea, Plates XLIX 

to LV. Mollusca, Plates IX to XIII, 1902. Crustacea, Plates LVI to LXVII. -Crustacea, Plates 

_LXVIII to LXXVIL. Fishes, Plates XXXVI to XXXVIII, 1905. Crustacea (Malacostraca), Plates 

BeEXXVII to LXXIX. Crustacea (Entomostraca), Plates I and II. Mollusca, Plates XIV to 

XVIII, 1907. Fishes, Plates XXXIX to XLIII. Crustacea (Entomostraca), Plates III to V. 

_ Mollusca, Plates XTX and XX, 1908.—Re. 1 per plate. Mollusca, Plates XXI to XXIII, 1909.— 
As. 8 Be ‘plate. 


“3 
“y 


RECORDS 
of the 


INDIAN MUSEUM | 


Vol. I, 1907. 


Part I1.—Contributions to the Fauna of the Arabian Sea. Heimiptera and Hymenup- 


tera from the Himalayas. Indian Freshwater Entomostraca. The Fauna oi 
Brackish Ponds at Port Canning, Lower Bengal, I—III. A Sporozoon from the 
heart of a Cow. Miscellanea :—The appendicular skeleton of the Dugong. An 
egg laid in captivity by a Goshawk. Melanic specimens of Barbus ticto. Two 
barnacles new to Indian seas. Mosquitoes of the genus Anopheles from Port 
Canning. Anopheles larvae in brackish water. Mosquitoes from Kumaon. 
Peculiar habit of an earthworm. : 


Part II.—Revision of the Oriental Stratiomyidae. An Oligochaete Worm allied to 


Chaetogaster. The Fauna of Brackish Ponds at Port Canning, Lower Bengal, IV. 
A Polyzoon from the Himalayas. Batrachia, Reptiles and Fish from Nepal and 
the Western Himalayas. The Fauna of Brackish Ponds at Port Canning, Lower 
Bengal, V. Oriental Diptera, I and II. Miscellanea :—Gecko verticillatus in 
Calcutta. The distribution of Kachuga sylhetensis. ‘The distribution of Bufo 
andersont. Note on Rutilia nitens. Some Indian Cerambycidae. Some Indian 
Hemiptera. A preoccupied specific name in Macrothyixv. An enemy of certain 
Pearl Oysters in the Persian Gulf. The distribution in India of the African snail, 
Achatina fulica. Statoblasts from the surface of a Himalayan pond. Notes on 
Hislopia lacustris. 


Part III.—Marine Polyzoa in the Indian Museum. ‘The Fauna of Brackish Ponds at 


Port Canning, Lower Bengal, VI. Earwigs (Dermaptera) in the Indian Museum. 
Oriental Diptera, III. A new snake from Nepal. Marketable fish from Akyab. 
Freshwater Oligochaete Worms from the Punjab. Phosphorescence in Marine 
Animals. The rats of Dacca, Eastern Bengal. Freshwater Sponges, I—V. 
Miscellanea :—The original home of Mus decumanus. Colour change in Hylo- 
bates hoolock. Eggs of Tylototriton verrucosus. The hosts of Tachaea spongil- 
licola. A second species of Dichelaspis from Bathynomus giganteus. 


Parvt IV.—Nudiclava monocanthi, the type of a new genus of Hydroids parasitic on 


Fish. Three new Nycteribiidae from India. Annotated Catalogue of Oriental 
Culicidae. Oriental Diptera, IV. Freshwater Sponges, VI, VII. A new Cyprinid 
Fish of the genus Danio from Upper Burma. Miscellanea :—A colour variety of 
Typhlops braminus. Reptiles and a Batrachian from an island in the Chilka 
Lake, Orissa. ; ; 


Vol. II, 1908-1909. 


Part I.—Retirement of Lieut.-Col. Aleock. The Fauna of Brackish Ponds at Port 


Canning, Lower Bengal, VII. A new Dictyonine Sponge from the Indian Ocean. 
Freshwater Sponges, VIII. Remarkable cases of variation, I. A new species of 
Lizard of the genus Salea, from Assam. ‘The Fauna of Brackish Ponds at Port 
Canning, Lower Bengal, ‘VII. A new Cavernicolous Phasgonurid from Lower 
Siam. New species of Marine and Freshwater Shells in the Indian Museum 
Oriental Syrphidae, I. A new variety of Spongilla loricata. Oriental Diptera, V. 
Miscellanea : ~Remarks on Simotes splendidus. Corrections to No. IV of ‘‘ Notes 
on Oriental Diptera.’’ The Isopod genus Tachaea. The habits of the Amphipod. 
Quadrivisio bengalensis. New varieties of Nanina berlangeyi and Corbicula 
fiuminalis. Recent additions to the Entozoa in the Indian Museum. A sub- 
fossil polyzoon from Calcutta. Corrections as to the identity of Indian Phylac- 
tolaemata. A peculiar form of Euglena. 


Part II,—Gordiens du Musée Indien. The Fauna of Brackish Ponds at Port Canning, 


Lower Bengal, IX. A new species of Danio from Lower Burma. Rhynchota 
Malayana, I. Cimex rvotundatus, Signoret. Freshwater Sponges, IX. Fruit 
Bats inhabiting the Andaman and Nicobar Archipelagos. A new species of Sun- 
Bird obtained near Darjiling. Three Indian Phylactolaemata. Two new species 
of Hagle-Rays (Myliobatidae). A new species of the genus Sesayma, Say., from 
the Andaman Islands. New species of and, Marine, and Freshwater Shells from 
the Andaman Islands. 


Part III.—The Fauna of Brackish Ponds at Port Canning, Leiter Bengal, X, XI. 


Oriental Solifugae. The difference between the Takin (Budorcas) from the Mishmi 
Hills and that from Tibet. Cavridina nilotica (Roux) and its varieties. A new 
species of Chavaxves !rom the Bhutan Frontier. First report on the collection of 


X. ANNOTATED CATALOGUE OF ORIENTAL 
CULICIDAE—SUPPLEMENT. 


By E. BRUNETTI. 


Almost immediately after the publication of my ‘‘ Annotated 
Catalogue of Oriental Culicidae ’’ ' I obtained access to a copy of 
the fourth volume of Mr. Theobald’s Monograph on this family, 
and to Mons. Blanchard’s copious work, ‘‘ Les Moustiques.”” Sub- 
sequent to these is a very lengthy and valuable paper on the 
Culicidae of the Malay States by Dr. G. F. Leicester (with a preface 
by Mr. C. W Daniels, Director of the Institute for Medical Re 
search) published by that Institution.? Also a long paper by Mr. 
‘Theobald (‘‘ 2nd Revort on the Indian Museum Culicidae ”’).? 

I have also received a copy from Miss Ludlow of her thesis on 
the mosquitoes of the Philippines. The very consilerable list of 
additions and corrections of importance relating to Or ental species 
gleaned even from these five works alone render a supplement to 
my catalogue imperative, and the information contained in the 
present paper is mainly derived from these sources with the inclu- 
sion of the 5th volume of Theobald’s Monograph recently issued. 

The splitting of genera and species still continues to such 
extremes as to invoke the severe deprecation of more than one 
systematic dipterologist, more e pecially in th> case of groups 
higher than genera, of which, none of those recentlv erected in this 
family approach in zoological value groups of similar rank in the 
other families of Diptera. 

I have dealt elsewhere* with the quest’on of taxonomic 
values in Culicidae, and therefore need not recanitulate here any 
observations that apply only to nomenclature. In that paper was 
mentioned that in addition to the vast accessions in generic rank 
proposed by the new school of culicid students, new methods were 
adopted in presenting to others the results of their labours, and 
though I feel ill fitted to condemn or criticize, it does not seem 
entively out of place to sugzest that, in as many particulars as 
possible the generally accepted rules of zoological literature should 
be adhered to. 

Mr. Theobald’s methol in his Mo 1egravh of placing the 
author’s name after the quotation insteal of hefore it, renters it 
rather awkward to follow the data presented in this unusual 


1 Rec Ind. Mus , i, 297—377 (1907). 
or aie from the Institute for Medical Research, Federated Malay States, 
vol. iii. 
3 Rec Ind Mus., iv, 1—33 (1910). 
* ** Taxonomic values in Culicidae.’’ Rec. Ind. Mus., iv, §3 (1910). 


404 Records of the Indian Museum. [ Wor. iV 


manner. Another serious inconvenience in referring to this author’s 
work is the index, where the genera themselves are alphabetically 
arranged, all the species of each genus forming a separate alpha- 
betical list under each generic name. ‘This plan is still retained 
even in the fifth volume. So, unless the student knows to which 
genus this author refers each species, he has to search the whole 
index. A single alphabetical list of specific names as is given in 
the valuable Catalogue of Diptera by Kertesz would have much 
facilitated reference, and the genus of each could have been entered 
in a second column. 

In Blanchard’s otherwise admirable work he adopts a very 
laborious method of quoting merely a reference date and letter 
(1g01a, 1902a, 1902b, etc.) for each paper of each author, thus 
necessitating an examination of his list of papers (given at the end 
of the book) every time a quotation is desired. The amount of 
additional and unnecessary labour entailed by this double reference 
is enormous. 

The habit of authors on Culicidae of allowing the female to 
take precedence of the male is in absolute defiance of zoological 
rule, and it is to be regretted that Dr. Leicester in his great paper 
on the Malay species, has continued this practice, even though he 
had before him males of the greater number of the species dealt 
with. 

In connection with this question it is well to mertion the case 
of the Anopheline mosquito known as cut/licifacies, Giles. 

Giles originally described under that name, what afterwards 
proved to be two distinct species (namely, his ~ is now known as 
turkhudz, Liston, and his 9 as culicifacies, Giles). 

Now, in accordance with the strict rules of zoological litera- 
ture, as I have always understood them, in such a case the name 
of the species is invariably retained for the ~, and a new name 
provided for the 2. This being so, the name culicifacies should 
have been retained for the male (now called turkhudi), and the 
female (still known as culicifacies) renamed. However, to avoid 
further confusion the synonymy was not altered in my catalogue 
nor in the present supplement, but it seems advisable to call 
attention to the fact and to protest against the 2 taking prece- 
dence of the ~ in such cases. 

A brief review of the recently published works on Culicidae 
may now be made. 

Mr. Theobald’s 4th volume (Monog. Culicidae World) contains 
notes (p. I) on the growing of Lemna minor, L. arrhiza and other 
duckweeds, on the surface of all unavoidable collections of water 
as a preventive against the breeding of mosquitoes. Mr. Green 
notes that they breed freely in the flowers of Heliconia brasiliensis. 

On p. 3 Mr. Theobald gives a list of the species that are 
known to be agents of infection. On p. 6 he gives Prof. Felt’s 
table for the identification of culicid larvae ; on p. 11, Dr. Dyar’s 
grouping and formation of genera by @ genitalia. As a criticism 
on the classification by larvae, Mr. Theobald remarks (referring to 


1912. | E. BrunET?I1: Catalogue of Oriental Culicidae. 405 


Messrs. Dyar and Knab’s paper, in which only 3 sub-families are 
recognized ,—Anophelinae, Culicinae and Sabethinae), ‘‘ all other 
genera of Anophelinae are sunk as synonyms of Anopheles, but 
the authors raise one species—barbert of Coquillett, a species so 
close to bifurcatus that it is hardly separable—to generic rank, 
calling it Coelodiazests.”’ 

Mr. Theobald mentions their plates of portions of the larvae 
as being valuable for future work. On p. 14 Theobald mentions 
Coquillett’s classification of the family on adult characters, on 
p- 15 the proposed separation of Corethra and Mochlonyx from 
Culicidae, to form a separate family. Dr. Lutz’s classification is 
given on p. 15 followed by a modification of it by Theobald on 
p. 17; the latter writer being in favour of the separation of 
Corethva. The further notes of interest in Mr. Theobald’s work 
are mentioned under the genera and species to which they apply. 

In the 5th volume of his Monograph of the Culicidae Mr. 
Theobald reviews all the species included in the previous volumes. 
It is a huge work of over 600 pages, illustrated by 261 text-figures 
and six plates of wings, and contains descriptions of 2I new 
genera and 392 new species. 

It contains apparently lists of all known species in each genus 
(except those purposely excluded for given reasons) and presents 
them in tabular form. 

It is satisfactory to see the author deploring the brief nature 
of some authors’ description of their species, ‘‘ wholly inadequate 
for correct diagnosis,’’ and as certainly leading to much confusion 
and increased synonymy. He also, rightly enough, objects to new 
genera and species being created on larvae of which the adult 
forms are unknown, and he emphasizes this objection by ignoring 
the species thus erected by Messrs. Dyar and Knab on American 
and West Indian forms. 

It is curious that Theobald makes no reference either to the 
voluminous monograph on the Malayan Culicidae published by Dr. 
Leicester, nor does he apparently notice any of that author’s very 
numerous new species, mostly described from bred specimens. 

In one or two cases he quotes verbatim descriptions of species 
by other authors without notifying from which region of the globe 
they come. 

Blanchard in his ‘‘ Moustiques”’ (1905) devotes chap. 1 to the 
position of the Culicidae, chap. ii to the morphology and anatomy 
of the family, and chap. iii to their metamorphoses and habits. 
Notes on mosquito parasites occur on pp 132-135. A long chapter 
of nearly 300 pages, illustrated by 120 figures, is devoted to the 
systematic description of genera and species. Chapter v treats of 
the medical aspect, chap. vi of methods of prevention of attack 
and of extermination, and chap. vii of their collection, preserva- 
tion, breeding and mounting. An appendix giving recently des- 
cribed species a very complete bibliographical catalogue, and a 
copious index to the whole work completes the volume, which 
totals 673 pages. On p. 390 he givesa key to the new genera 


406 Records of the Indian Museum. [Vou. IV, 


contained in the 3rd volume of Theobald’s * Monograph which 
appeared whilst Blanchard’s work was in the press. 

Miss Ludlow’s paper on the connection between malaria and 
the occutrence of various species of mosquitoes in the Philippines 
contains very extensive information respecting their distribution. 

Many of the species appear widely distributed, Myzomyia 
ludlowit being reported from no less than 42 different localities 
in these Islands, M. indefinita from 26, M rossi, vanus, pseudo- 
harbirostris, funestus and barbirostris from ten or more localities 
each, besides other species from a lesser number of localities each. 

She notes that it is probable that some species may pass 
through the dry season as adults, hibernating amongst the dry 
vegetation, and also notes that in localities where the rainy season 
advances gradually, the Anophelinae are more numerous and exist 
in considerable numbers throughout a good part of the dry season, 
whereas in localities where the rainy season is introduced by very 
excessive and constant deluges they are markedly less in numbers, 
presumably by the breeding places of the insects in their earlier 
stages being washed away. 

“Four Anophelinae, funesta, barbivostris, fuliginosus, and 
ludloww . . . seem likely to be acting as hosts for the malarial 
parasite in the Philippines, and concerning Stegomyia calopus Mg.(= 
S. fasciata F.), Culex fatigans W., and Mansonia uniformis Theob.., 
there are too few data to judge if they be carriers of disease’’ 
(Ludlow). Regarding Stegomyia fasciata, the acknowledged sole 
carrier of yellow fever, this author significantly remarks: ‘‘ Yellow 
fever has so far never been present in the Philippines. The wide 
distribution of S. calopus (= S. fasciata F.) is, however, very 
suggestive taken in connection with the building of the Panama 
Canal, as to the result likely to follow, should vellow-fever-infected 
mosquitoes or patients in the proper stage of the disease reach the 
Islands.” 

Mr. G. F. Leicester in his important and extensive paper on 
* The Culicidae of Malaya”’ devotes over 250 pages to fully redes- 
cribing the mosquitoes of this region, including nearly a hundred 
new species. In his preface he notes that the 3rd volume of 
Theobald’s Monograph appeared just before the publication of his 
own work and that an appendix will be necessary, involving some 
changes of nomenclature, and that a further paper on the larval 
characters may eventually follow. 

In this paper he devotes 14 pages to the breeding grounds of 
mosquitoes with some notes on collecting and preserving them, 
but although he seems to have bred a great number of the species 
and fully described numbers of them from long series of fresh 
specimens he gives no definite dates of appearance. 

A further report by Theobald on the Indian Museum Culicidae 
(the 2nd) has recently been issued! in which four new genera 
and twenty-one new species are described. It has appeared 


1 Rec, Ind. Mus., iv, 1—33 (1910). 


1912.] E. BrRuNETTI: Catalogue of Oriental Culicidae. 407 


unnecessary to copy the whole of the data supplied in this author’s 
paper (which is easily accessible) in cases of very common or 
widely distributed species, especially if the localities therein given 
are already in my Catalogue. 

In such cases the species are reported in this paper as ** com- 
mon throughout India,’’ etc., as easy reference can be made to 
exact details. 

Major S. P. James in ‘‘ A new arrangement of the Indian 
Anophelinae”’ (Rec. Ind. Mus., iv, 95—109) criticizes the evanescent 
nature of Anopheline genera, noting that if maculipennis, Mg., be 
the type of Anophcles, s. s., there is no Indian species of the genus. 
He divides the Indian Anophelina into two groups, those with. 
and those without abdominal scales. Of the Ist group he admits 4 
genera—WNeostethopheles, gen. nov. (pl. i), with attkenz, James, as 
type; Myzomyia, Blanch. (pl. i), with culicifacies, Giles, as type ; 
Patagiamyia, gen. nov. (pl. i), type gzgas, Giles ; and Pyretophorus, 
Blanch. (pl. i), with palestinensis, Theob., as ‘‘ type example.”’! 

In the 2nd group he gives these genera: Nyssorhynchus, Blanch. 
(pl. ii), type maculatus, ‘Theob.; Myzorhynchus, Blanch. (pl iv), 
type barbirostris, V.Wulp; Cellia, Theob (pl. iii), type pzlcherrima. 
Theob.; Neocellia, Theob. (pl. ii1), type endica, Theob.; Aldrichia, 
Theob., type error, Theob.; Nyssomvzomyia, gen. nov. (pl. ii), type 
vOSS11, Giles - Christophersia, gen. nov. (pl. iv), type hallit. 

The two new genera proposed, Neostethopheles and Pata- 
giamyia, are not admitted in the present Catalogue, as my manu- 
script was practically completed when these genera were set up, and 
also because it is quite evident that the workers in mosquitoes are 
further off than ever from any definite agreement amongst them- 
selves as to either the number or the limits of the genera to be 
recognized.” The third genus proposed by James—Christophersia— 
is acknowledzed here as it comprises one species only, which has 
not previously been located in any other genus. 

In the 2nd edition of Messrs. James and Liston’s ‘‘ A Mono- 
graph of the Anopheline Mosquitoes of India’’ a good deal of 
additional matter is introduced. Their classification into Mega- 
rhinae, Limatinae, Anophelinae, Aedinae and Culicinae need not 
be criticised here. Their suggestion (p. 15) to make use of the 
botanical terms to describe the different shapes of the scales seems 
an excellent one, asthe terms now used are ambiguous and have 
not the same meaning for every author. Collecting and mounting 
are detailed, but it is certainly time that the method of using 


} The true generic type is costalis but has not been seen by James, and, as 
he remarks, the theracic scales mav be different. In any case hewever castalés 
must still rema‘n the tvpe of the genus, 

2 In the ‘‘ Bulletin of Entomological Research” for Mav tori Mr J. W. W. 
Stephens calmly announces that ‘ A caref] examination with a pocket 'ens (1!) 
should enabl: you to state almost with certainty whether or no all the Anophe- 
lines you have caught are of the same species.’’ Yet those who have studied 
them for years are, as stated, still very much at variance as to specific limits. 


408 Records of the Indian Museum. [ VOL. Ven 


cardboard discs should be abolished and neatly cut blocks of pith 
be substituted.! + 

The authors’ suggestion (p. 17) to ignore the legitimate claims 
of priority, cannot, of course, be sustained. One feature of this 
work is that the descriptions are drawn up without reference to 
sex, applying presumably to both, except where, here and there, 
a character is noted as present in one sex only. 

It is extraordinary how specialists in mosquitoes continue to 
wrongly name the veins, and especially what they call ‘‘ cross- 
veins.” 

In James and Liston’s work, the longitudinal veins are cor- 
rectly designated, but they speak of the short basal section of the 
2nd longitudinal vein (before it takes its longitudinal course) as a 
cross-vein (the ‘‘ marginal’’). It is nothing of the sort, the mar- 
ginal cross-vein not being present in the Culicidae at all: and in 
those families in which it does occur it is always in the distal half 
of the wing. 

Again, James and Liston’s ‘‘ supernumerary cross vein’’ is 
merely the basal section of the 3rd longitudinal vein, and cer- 
tainly not a cross-vein at all. 

Their ‘‘ mid cross-vein’’ may be thus called though ‘‘ anterior ” 
cross-vein is the more correct term; and the posterior cross-vein 
they have happily correctly recognized. 

The subcostal cross-vein of James and Liston is not this 
vein at all, but the humeral cross-vein, the subcostal cross-vein not 
being present in the Culicidae, and in those families in which it 
does occur it joins the auxiliary and Ist longitudinal veins. There 
are only three cross-veins in Culicidae—the humeral, anterior 
and posterior. 

In speaking of the cells, James and Liston say that the ‘‘ areas 
enclosed between these branches ” (7.e., of the forked longitudinal 
veins, the 2nd and 4th) “‘ have received names ”’ (mentioning only 
the Ist submarginal, 2nd posterior and ‘‘ anal ’’ cells) but continue 
by cheerfully ignoring all the other cells as ‘‘ for our present pur- 
pose they need not be mentioned ’’! 

Moreover, they are wrong again in their ‘‘ anal’’ cell, which 
is really the ‘‘ 4th posterior’ cell. The anal cell is always behind 
or posterior to the 5th longitudinal vein, or the hinder branch of 
it when this vein is forked. 

Perhaps Theobald is most to blame for these errors, as being 
the pioneer of a false terminology. ‘This author’s ‘‘ supernumerary 
cross-vein’’ is merely the basal section of the 3rd longitudinal 
vein. He also figures the somewhat similar section of the 2nd 
longitudinal vein as a cross-vein, but gives it no title, either in 


c 


; : The method I adopt for mounting all very small Diptera is to thrust the 
minute pin through the right side of the thorax, immediately below the dorsum, 
at such an angle that the point emerges from the left side immediately above or 
between the legs. It is then possible to view the greater part of both the dorsal 
and side surfaces without removing the specimen from the cabinet. 


19gt2.] EE. Brunetti: Catalogue of Oriental Culicidae. 409 


the explanation of the figure (Monog., i, p. 18) or in the accom-. 
panying letter-press. 7 

In the hope that it may be of service to those who have not 
sufficiently mastered the terminology of the veins in Diptera the 
present figure is offered, with the correct names of the veins 
appended. 

Colonel A. Alcock has recently published a paper on the 
classification of the Culicidae,! in which he deplores the undue 
rank apportioned to mere groups of species in this family, and 
recognizes Corethrinae as certainly belonging here, but follows 


Terminology of venation in Culicidac. 


C costa. a_ costal cell. 

S.C sub-costal or auxiliary vein. b_ sub-costal cell. 

1 Ist longitudinal vein. c marginal cell. 

2 2nd do. d_ ist sub-marginal cell. 
The two branches are termed the e 2nd do. 
anterior (or upper) and posterior f 1st posterior cell. 
(or lower) branch respectively. g 2nd do. 

3. 3rd longitudinal vein, h 3rd do. 

4 4th do. {oe 4th do, 

(The branches named as in the k anal cell. 
2nd vein.) 1 axillary cell. 
5 5th longitudinal vein. m_ st basal cell. 
(The branches named as in the n 2nd do. 


2nd and 4th veins.) 

6th longitudinal vein. 

humeral cross-vein. 

anterior (or ‘‘ mid,’’ or ‘‘ small ’’) 
cross-vein.? 

posterior (or ‘* hinder ’’ or ‘‘ large’’) 
cross-vein. ? 


SHO 


N 


the practice of other authors in instituting new terms by propos- 
ing the establishment of four sections: (1) Megalorhini (=the Mega- 
rhinae of Theobald), (2) Epialurgi (evolved from “ ague fever ’’ 
and ‘‘ work,”’ this group representing the Anophelinae of authors), 
(3) Culicales (— Culicinae, Heptaphlebomyinae, Dinoceratinae, 
Aedinae and Uranotaeniinae of Theobald), (4) Metanototricha 


1 «* Remarks on the classification of the Culicidae, with particular reference 
to the constitution of the genus Anopheles,’’ Ann. Mag, Nat. Hist. (8), viii, 
No. 44, p. 240 (August I9I!1). 

2 All three sets of terms have been used by good dipterologists but anterior 
and posterior are eminently the best fitted for permanent adoption since these - 
relative positions are constant in all wings in which both cross-veins are present, 
whereas the other terms are sometimes inappropriate. 


410 Records of the Indian Museum. [VOL IV; 


(= Trichoprosoponinae, Dendromyinae, Limatinae ot Theobald), 
‘ this group being entirely conventional.”’ ! 

One of the most valuable points in the paper is the recognti- 
tion of most of the so-called ‘‘ genera ”’ as sub-genera only. 

I have endeavoured to retain the sequence of genera as 
near as possible to that followed in my Catalogue, for there is as 
yet, apparently no uniform agreement as to the disposition of 
many of the genera, even into the so called sub-families. 

Several genera admittedly hold intermediate positions, which 
clearly supports the contention that the less the number of genera 
in Culicidae, the more zoologically correct the classification. Res- 
pecting the value of the so-called species the present writer offers 
no opinion, but as authors are already speaking of ‘‘ Culex so-and- 
so, and its allies,’’ it is reasonable to conclude that considerable 
doubt exists still as to specific limits, and that the opinion ex- 
pressed four years ago in the introduction to my Catalogue 
that ‘‘a few more years careful study of the family is more likely 
to result in the reduction than otherwise of the total number of 
what today are regarded as distinct species’’ seems within posst- 
bility of realization. 

Four new ‘‘ generic’’ names are proposed in the present paper 
for names already preoccupied, and it is significant as showing how 
little culicidologists concern themselves with dipterological litera- 
ture. that three out of these five names should have been pre- 
viously used in the order Diptera itself ! ? 

Other names are so similar to long previously established 
ones that confusion is at least probable. Such are Popfea, Ludlow, 
closely resembling Poppfea, Stal. (1567), in Hemiptera; whilst two 
other recently established genera (non-Oriental) bear names re- 
matkably sim lar to others long established in other divisions of 
the animal kingdom. ‘These are Carrollia, Lutz, practically pre- 
occupied by Carvollia, Gray (1838), in Mammalia, and by Carolia, 
Cantr (1837), in Mollusca; also Ankvlorhynchus, Lutz, preoccu- 
pied by Ancylorhynchus, Schonh. (1836), in Coleoptera. 

Miss Ludlow has emended her generic name Calvertia to 
Calvertina (Can. Ent., xli, 234), it being preoccupied by Warren 
in Lepidoptera. 

There also exist two other very similarly named genera Calver- 
tius, Sharp, in Coleoptera and Calveria, Carp., in Echinodermata. 

During two tours made by me, one round the Punjab and 
north-western part of India in tg05 and one round the far east 
in 1906, I collectela certain number of Culicidae but paid no 
espe-ial attention to their capture or preservation, with the result 
that the condition of the specimens renders them practically 


b) 


1 Col. Alcock now accepts for these groups the more appropriate names 
“* Megalorhinina,’’ ** Anophelina,’’ ‘‘ Culicina’’ and *‘ Metanototrichina ’’ (Bull 
Ent. Res., ii, p. 241, 1911). 

2 A ff h instance was included :n the original MS of this paper — Aldvichsa, 
Theob. (preoccupied in Bombylid ie by Coquillett)—but in his last volume Theobald 
alters it 10 Aldvichinella This genus (Aldyichia) made another instance of 
ignored preoccupation in Diptera ! 


¥91z.} EE. BRUNETTI: Catalogue o} Oriental Culicidae. 411 


valueless for the cabinet. Mr. Theobald, however, most generously 
‘ooked tnrough them and suggested the identity of a certain num- 
ber of the more easily recognized species, and the data referring 
to these are included in the present paper, mainly for the sake of 
recording the localities. These species are Myzomyza rossi, Giles ; 
Myzorhynchus sinensis, Wied. ; Desvotdya obturbans, Wk. ; Theo- 
baldiomyia (nom. nov. for Leucomyra) gelidus, Theob.; Culex 
fatigans, Wied. ; concolor, R. Des. ; tigripes, Grandpré ; impellens, 
Wlk. ; microannulatus, Theob. ; sericeus, Theob.; Mansonia annu- 
lipes, Wik. ; and Stegomyta fasctata, Fab. 


ADDLTIONS| COLE LIST OF “LITERATURE .ON 
ORIENTAL CULICIDAE. 


Adie, Major, 1904. Indian Medical Gazette, xxxix, June, No. 6. 

Aitken, E. H., ryo2 ‘‘ Notes ona Tour in the North Canara 
District of India in search of Mosquitoes,’’ Journ. Trop. 
Med., v, 325—327 ; 341—343- 

Alcock, Col. A., 1911 (August). ‘‘ Remarks on the classification 
of the Culicidae, with particular reference to the genus 
Anopheles,” Ann. Mag. Nat. Hist. (8), viii, No. 44, p. 240. 

Annandale, Dr. N., 1g11. ‘‘A new genus of short- beaked Gnats 
from Ceylon,’’ Spol. Zeyl., vii, 197. 

Blanchard, R.,1905. ‘‘ Les Moustiques; historie naturelle et medi- 
cale ’’’ (673 pp.). (Containsa very complete bibliography.) 

Brunetti, E., 1909 ‘‘ Oriental Culicidae’’ (reply to criticism of 
my catalogue), Can. Ent., xli 121. 

Id., 1910. ‘‘ Taxonomic values in Culicidae,’’ Rec. Ind. Mus., 
IV, 53- 

Id., 1911. ‘‘Synonymy in Corethrinae,’’ loc. ctt., iv, 317. 

Id., 1911. ‘‘ Further notes on Synonymy in Corethrinae,’’ doc. 
Cit NA 227 

Dyar, Dr., 1905. A paper on the grouping of genera by the o 
genitalia, in Proc. Ent. Soc. Wash., vii, No 1. 

Dyar and Knab, 1906. ‘‘ The larvae of Culicidae classified as 
independent organisms,’’ Journ. New York Ent. Soc., 
Xiv, 169—2_0. 

Id., 1907. ‘‘ On the classification of the Mosquitoes,’’? Can. Ent.., 
MAKI, 47: 

Id., 1908. ‘‘ Notes on Mosquito work,’’ Can. Ent., xl, 309. (This 
is a short critical article on classification.) 

Eysell, Dr. Adolf, 1905. ‘‘ Sind die Culiciden eine Familie ? ’’ 
Archiv. fur Schiffs und Tropen Hygiene, ix 51—55. 

Felt, Prof., 1905. Bull. 97, Entom. 24, Divis. Entom. New York 
State Mus., 445. 

Ficalbi, 1896. The ‘‘ Revisione,’’ etc., quoted in my catalogue is 
from the Boll. d. Soc. Ent. Ital., vol. xxi, e¢ seg. (1888 
et seq.), 300 pp., 4 plates. 

Giles, G. M., 1900. ‘‘ Species of Anopheles in Shanghai and 
Java,’’ Brit. Med. Journ., 1, 485. 


412 Records of the Indian Museum. [Vor. IV, 


Giles, G. M., Igor. *‘* Notes on Indian Mosquitoes,’’ Journ. Trop. 
Med., iv, 159. 

Id., tgo1. **A plea for the collective investigation of Indian 
Culicidae,’’ Journ. Bom. Nat. His. Soc., xiii; 592. 

Id., 1903. ** Note on Mansonia anopheloides,’’ Journ. Trop. Med. , 
Vi 5320: 

Id.. 1904. ‘* A revision of the Anophelinae,’’ being the rst Supp. 


to the 2nd Ed. of “ A Handbook of Gnats or Mosquitoes.” 
London, 47 pp., 8vo. 

[d., 1904 (Dec.). ** Notes on some collections of mosquitoes, etc., 
received from the Philippine Islands and Angola, with 
some incidental remarks upon classification.”’ 


Id., 1904 (Dec.). ** Notes on some collections of mosquitoes re- 
ceived from abroad,’’ Journ. Trop. Med., vii, 365—3609. 

Grinberg, K.. 1907. ‘‘ Die Blutsaugenden Dipteren.’’ Jena. 

James, S. P., Igt0 (Nov. 18th). ‘‘ A new arrangement of the 


Indian Anophelinae,’’ Rec. Ind. Mus., iv, 95. 
James and Liston, r911. ‘‘ A Monograph of the Anopheline Mos- 
quitoes of India.” Calcutta. . 
Knab, F., 1907. ‘‘ Culicid characters,’’ Can. Ent., xxxix, 349. 


Laveran, 1901. ‘‘ Sur les culicides provenant de Hanoi’’ (Tonkin), 
Comp. Rend., liii, ggr. 

Id., tg01. “* Sur les culicides provenant de Haut-Tonkin,’’ loc. 
cit., 993. 

Lebredo, M., 1904. ‘‘Some observations on the anatomy of mos- 
quitoes,’’ Revista de medicine tropicale.. Havana. 


Leicester, G. F., 1908. ‘‘ Notes on the Culicidae of Malaya,’’ pub- 
lished in Studies from the Institute of Medical Research, 
Kuala Lumpur, vol. iii, with prefatory notes by C. W. 
Daniels, and followed by (in the same volume, which 
contains papers by other authors bearing on the malarial 
aspect) another paper by Daniels containing notes on the 
mosquitoes on the eastern side of the Malay Peninsula. 

Liston, W. G., rgo01. ‘‘ The distribution of Anopheles in Ellichpur 
Cantonment, State of Berar,’ Ind. Med. Gaz., xxxvi, 
I29—132, and Journ. Trop. Med., iv, 164. 

Id., 1901. ‘* A year’s experience of the habits of Anopheles in El 
lichpur,’’ Ind. Med. Gaz., xxxvi, 361—366 and 441—443. 

Id., 1902. ‘‘ Classification of Anopheles in India,” Journ. Trop. 
Med., v, 146. 

Ludlow, 1902 (Aug. 23rd). ‘‘ Description of a new Anopheles,”’ 
Journ. Am. Med. Assoc. 

Id., 1902 (Sept.). ‘*‘ Two Philippine mosquitoes,’ Journ. New 
York Ent Sock, x. 127: 

Id., 1902 (Sept.). ‘‘ Notes on Culex annulatus,’’ Journ. N.Y. Ent. Soc. 

Id., 1905. ‘‘ Mosquito Notes,’’ No. 3, Can Ent., xxxvii, 94 and 
129; No. 4, loc. cit., 385, and (1906) xxxvili, 132 (con- 
cluded). : 

Id., 1906. Id., No.5, .loe: (cit xxxvan, (367, and (1907) meexxixg 
129 (continued), and 413 (concluded). 


1912. ] E. BRUNETTI: Catalogue of Onental Culicrdae. 413 


Ludlow, 1908. ‘‘ Mosquito Notes,’’ No. 6, loc. cit., xl, 32, 50, 331. 
Id., 1908 (Nov.). ‘‘ The Mosquitoes of the Philippine Islands.”’ 
Washington University. 

Id., 1909. *‘ Mosquito comment,’’ Can. Ent., xli, 21. 

Id., id. ‘*‘ New Philippine Mosquitoes,” loc. cit., 97. 

Id., id. ‘‘ Mosquito observations,”’ loc. ctt., 233. 

Id., id. ‘* Anopheles perplexens,’’ loc. cit., 293. 

Lutz, 1904. ‘‘ Mosquitos do Brazil.” 

Mitchell, Miss Evelyn Groesbeeck, 1907. ‘* The classification of 
the Culicidae,’’ Can. Ent., xxxix, 198. 

Page, H., 1906. ‘‘ Malaria and Mosquitoes at Lucena Barracks, 
Philippine Is,’’ Journ. Assoc. Milit. Surg., xix, 65—76. 

Peryassa, Dr. Antonio Goncalves. 1908. ‘‘Os Culicideos do 
Brazil.’’ Rio de Janeiro, 400 pp., 26 plates. 

Ross. Major Ronald, 1899. ‘‘ Life history of the parasites of 
malaria,’ Nature, lx, 322—324. 

Id., 1900 (Mar. 29). ‘‘ Malaria and Mosquitoes,”’ Joc. cit., 1xi 
522—527. 

Theobald, F. V., 1905. ‘* New Culicidae from India, Africa, Bri- 
tish Guiana and Australia,’’ Journ. Econ. Biol., i, pl. i. 

Id., 1907. ** Monograph of the Culicidae,”’ iv, Brit. Mus., London. 

Td., 1908. ‘‘ First report on the collection of Culicidae and Core- 
thridae in the Indian Museum, Calcutta, with descriptions 
of new genera and species,’’ Rec. Ind. Mus., ii, 287—302. 

Id., 1909. ‘‘ Second report, zd. id. id.,”’ Rec. Ind. Mus., iv, I—33. 

Id., 1910. ‘‘ Monograph of the Culicidae,’’ vol. v, 646 pp., 5 plates, 
British Museum, London. 

Williston, S.W.,1906. ‘‘ The classification of the Culicidae,”’ Can. 
Ent., xxxviii, 384.1 

N.6b.—To my catalogue, the following addition should be 
made on p. 302, line 13, after the word ‘‘ sufficient ”’ :— 

‘“ Coquillett in 1906 (Tech. Sc., ii, Bureau of Entom, U. S. 
Dept. Agric.) sinks Aedeomyinae and Haemagoginae in Culicinae, 
retaining Anophelinae, Megarhininae, Psorophorinae, Culicinae, 
Deinoceratinae, Uranotaeniinae and Trichoprosoponinae.”’ 


5) 


ADDITIONS TO LIST OF LOCALITIES. 


Berars .. Central India. 

Bukit Kutu .. 3,400 ft., Federated Malay States. 

Chittagong .. Eastern Bengal. 

Cochin State .. south India. 

Deccan 7. Ania. 

Deesa .. West Central India. 

Dehra Dun .. Foot of Mussoorie Hills (W. Himalayas). 

Jugra .. Onthe Langat River, on a solitary hill 
(1,000 ft.), Federated Malay States. 

Kangra Valley .. 4,500—4,800ft., Punjab (W. Himalayas). 


1 Not relating to Oriental species, but a severe criticism of present-day 
taxonomy in this family. 


414 


Records of the Indian Museum. 


Kawkareik 
Klang 
Kumaon 


Laguna 
Lake Chilka 
Luzon 


Maddathorai 
Mandalay 
Manipur 
Mindanao 
Moulmein 
Pallode 

Pangkor Laut 
Phagu of 
Port Swettenham 
Puri 

Raub 

Rizal +55 
Shasthancotta .. 
Shencotta 
Soerabaya 
Sukwani 
Tayabas 
Thaumaspur 
‘*The Gap” 


Theog 
Travancore 
Trivandrum 
Ukhrul 

Ulu Gombak 
Ulu Klang 


[VOL Iv: 


Base of Dawna Hills, Tenasserim, Lower 
Burma. 

On the Klang River, Federated Malay 
States. 

In Western Himalayas, embracing Naini 
Tal and Bhim Tal. 

One of the Philippines. 

Orissa, east coast India. 

The principal island in the Philippines, 
containing Manila. 

Travancore State, South India. 

Upper Burma. 

6,400 ft., Assam. 

Oue of the Philippines. 

Tenasserim, Lower Burma. 

Travancore State, South India. 

Malay Peninsula. 

9,000 ft., near Simla. 

Selangor, Federated Malay States. 

Coast town in Orissa. 

Federated Malay States (Pahang). 

One of the Philippines. 

Travancore State, South India. 

Travancore State, South India. 

City at eastern extremity of Java. 

Nepal near Bengal frontier. 

One of the Philippines. 

Nepal (Terai, base of Himalayas). 

Pass between Selangor and Pahang, Fede- 

rated Malay States. 

Near Simla. 

State in Scuth India. 

Capital of Travancore State, South India. 

6,400 ft., Manipur State, Assam. 

13 miles from Kuala Lumpur, Selangor. 

Near Kuala Lumpur. 


CATALOGUE. 
CULICIDAE. 


ADDITIONAL REFERENCES.—Blanch., Moust., r5r (tab. gen. 
in Culicidae). 


Sub-Family ANOPHELINAE. 


App. ReF.—Blanch., Moust., 157 (sub-fam. chars.). 
Theob., Monog. Culic., iv, 22 (tab. genera), 
24 (list of known spp.—16+1 uncertain), 25 
(tab. known spp.). 
Leices., Culic. Malaya, 18 (tab. genera). 


1Q12. | E. BRUNETTI: Catalogue of Oriental Culicidae. 415 


ANOPHELES, Meig. 


App. Rer.—Leices., Culic. Malaya, Ig. 
James and Liston, Monog. Anoph. Mosq. Ind., 
2nd Ed., 4o. 


N.B.—Col. Alcock recognizes seven groups—which he regards 
as sub-genera only—covering all the species of the Anophelinae, 
which according to him should all be comprised in a single genus. 
These sub-genera are, Christya, Arribalzagia, Myzomyia, Ano- 
pheles, Myzorhynchus, Nyssorhynchus and Chagasia. ‘Two quota- 
tions verbatinu appear advisable: ‘‘ For the sake of convenience 
the species that compose the genus may be grouped in sub-genera 
according to the following table; but the groups, though they 
can be defined with sufficient precision, grade into one another.”’ 

Atl these considerations justify the conclusion that 
the so- -called ‘ genera’ of the proposed ‘ sub-family ’ Anophelinae 
cannot be separately focussed as distinct generic conceptions, but 
must all be merged in one generalization.’’ In the sub-genus 
Anopheles, Alcock includes Stethomyta, Theob., Neostethopheles, 
James, Patagiamyia, James, and Cvclolepidopteron grabhami, 
Theob. 


A. aconita, Don. (aconittus). 


Removed here trom Mvzomvia by Theobold (Monog., v) saying 
that Donitz says (Zeits. fur Hygiene, xliti, 233) that A. jormosaensis, 
Tsuzuki, from North Formosa is only a variety of aconita, and 
proposes to change the name (t!nnecessarily) to cohaesa. It trans- 
mits malaria. James and Liston put it in their new genus Neostetho- 
pheles. 


A. aitkenii, James in Theob. 
ADDITIONAL LOcALiIty.—Meenglas, Dooars, Jalpaiguri, 9-viti- 
07 (Wallich], a » and @ in Indian Museu n the only specimens 
seen by Theobald since describing the species. 
A. barianensis, James, IQII. 
Monog. Anoph. Mosq. Ind., 2nd Ed., 70. 
Loc.-- Murree Hills, Punjab (7,000 ft.). Taken by Assistant 
Surgeon J. L. Wredden. 
A. dthali, Patton. 
Now referred by Theobald to Myzomyta. 


A. formosaensis, Jsuzuki, 1902. 


Archiv. f. Schiffs u. Tropen Hygiene, vi, 289. 


I am still in doubt as to what is meant by this specific name, 
not being able to consult the original descriptions. Apparently 


416 Records of the Indian Museum. [Vox. IV, 


there are two species of the same name by the same author from 
North and South Formosa respectively, the former being synony- 
mous with aconita, Don. (according to Donitz himself, who suggests 
renaming it var. cohaesa), and the latter being apparently specifi- 
cally distinct, as Theobald (Monog., v, 84) quotes it as ‘* formosa- 
ensis II,’’ though he is unable to place it generically. 


A. farmosus, Ludl., 1909. 
Can Ent ssxli.22- 9); 


Theob., Monog., Culic., v, 8, 2. 


Miss Ludlow notes (Mosq. Phil. Is., 10) the above species as 
shortly to be described, the description appearing in due course as 
quoted above. She says it is the only species in the Philippines 
belonging to Anopheles (s. strv.). Theobald has not seen it and 
suggests it may not be Anopheles. 

From Benguet, Ph. Is., March 1908. 


A. gigas, Giles. 


App. REer.—Blanch., Moust., 184 (Myzomyra 1d.). 
App. Loc.—Ceylon [E. E. Green]; Deesa, W. Centr. India 
[Maj. Nurse]. 


N.B. —This is made the type of James and Liston’s new genus 
Patagiamyta. 


A. immaculatus, Theob. 


Type in British Museum. 


A. lindsayi, Giles. 


App. REF.—Blanch., Moust., i169. 

App. Loc.—Dehra Dun (Mussoorie Hills, foot) [Thomson]; 
Ferozepore (Punjab) [Maj. Nurse}. 

Type in British Museum. 


var. maculata, Theob., I1gto. 
Rec. Ind. Mus., iv, I. 


‘‘ A very distinct variety.” Kurseong (5,000 ft.), 5-vii-o8, a 
perfect 2 [Annandale]. Type in Indian Museum. 


A, simlensis, James and Liston, IgIt. 
Monog. Anoph. Mosq. India, 2nd Ed., 41. 


This species, with gigas, Giles, and lindesayi, Giles, is placed 
by the authors in their new genus Patagiamyia, but as this genus 
is not admitted in this Catalogue, simlensis is referred to Anopheles. 


1912. | E. BRUNETTI: Catalogue of Orienta! Culicidae. 417 


A. treacherii, Leices., 1908. 


Culic. Malaya, 19, 7 9. 

N.B.—Described from a series bred from larvae taken in hill- 
side streams in jungle. Amongst the notes the author says: 
‘* This species is widely distributed in the Peninsula. . . . ° It 
is a blood sucker; it will not breed in captivity. and the larvae, 
unless mature, usually die.’’ 


A. wellcomei, Theob. 


App. REF.—Theob., Monog. Culic., iv, 33 2. 
Described from several 2 2, all slightly damaged. 
Type in British Museum. 


PATAGIAMYIA, James, roto. 
Rec. Ind. Mus.; iv, 98. 


James and Liston, Monog. Anoph. Mosq. India, 2nd Ed., 41. 

This genus is not adopted in the present Catalogue. James 
and Liston make gigas, Giles, the type species, the other Indian 
species being lindsayi, Giles, and szmlensis, James. all herein 
referred to Anopheles. 


NEOMYZOMYIA, ‘Theob., 1910. 
Monog. Culic., v, 29. 
N. elegans, James in Theob. 


Anopheles elegans, James and Liston, Anoph. Mosq. Ind., 82. 

Myzomyia id., Theob., Monog. Culic., iii, 51. 

Pyretophorus id., id., op. cit., iv, 77. 

Theobald redescribes the species in vol. v, 30, 7 2 with two 
figs. of the 2 wing, giving other details. 

Locs.—Karwar (Bombay Presid.) [Coghzl/]; Andaman Is. | Ray 
White|, Meenglas, Jalpaiguri [Wallich}. 


N. leucophyrus, Don. 


Nyssorhynchus id., Blanch., Moust., 213, 2 . 

Myzomyta td., Leices., Culic. Malaya, 28, 7 @. 

N.B.—The latter author describes both sexes (the @ for the 
first time) from a large series bred from larvae from water in open 
bamboos in jungle. ‘The species is wholly sylvan, and is removed 
to this genus by Theobald (Monog., v, 44). 


MYZOMYIA, Blanch. 


App. ReErs.—Theob., Monog. Culic., iv, 41 (list of known 
species—20), 42 (table of spp.). 


418 Records of the Indian Museum. [VorL. IV, 


Leices., Culic. Malaya, 23, o &. 
James and Liston, Monog. Anoph. Mosq. India, 
2nd Ed., 40. 
N.B.—Col. Alcock sinks Neomyzomvia, Theob., Pyretophorus, 
Blanch., and Nvssomyzomyia, James, in Myzomyza. 


M. albirostris, Theob. 


App. REF.—Leices., Culic. Malaya, 23, 7 2. 

Type in British Museum. 

N.B.—Leicester savs that the larva occurs in small running 
streams with grassy edges. It is a blood sucker, and is easily 
identified when fresh by the parti-coloured proboscis. 

In bungalows, Kuala Lumpur. 


M. annularis, Wulp. 


I can glean no further information about this species, the type 
of which, described from Java, should be in the Leyden Museu.n. 
Theobald does not mention it in his 5th volume. 


M. azriki, Patton. 


App. Rrer.—Theob., Monog. Culic., v, 27, # 2, figs.o 2 palpi 
and palmate hair. 


M. christophersi, Theob. 


Admitted as a good species by this author (Monog., v) who 
adds the following localities : Sylhet, 2-v-5 [Ha//]; Kangra Valley ; 
Meenglas, Jalpaiguri, 13-vii-o7 [Wallich|; Calcutta, 2 2 2 
| Alcock}. 

M. culicifacies, Giles. 


App. Rrer.—Blanch., Moust., 182, 2. 

Theob., Monog. Culic., ii, 309 ; ili, 39 ; iv, 51; v, 
25. 

App. Locs—Mandalay, ~ 2 in bathroom, 13—14-ili-08 
| Annandale|; Lucknow, 21-i-08 [Hdgart] ; Deesa| Nurse]; Enuur, 
Goa. Bombay, Secunderabad, Aurangibad (Hyderabad State), 
Deccan (all in India and all ¢. Theobald). 

Type in British Museum. 

N.B—This varies greatly in size. I do not know what 
Blanchard’s @ of this species is. In the introduction to this paper 
some remarks are made as to the name of this species. 


M. deceptor, Don. 


Theobald removes it here from its vague position in ‘‘ A no- 
pheles’’ (Monog. Culic., v) and adds Trincomalee, Ceylon [Green], 
as a locality. 


1912.] EE. Brunerti: Catalogue of Oriental Culicidae. 419 


M. dthali, Patton. 
Also referred here from ‘‘ Anopheles ’’’ by Theobald (Monog., v). 


M. elegans, James. 


Referred to Neomyzomyta. 


M. funesta, Giles. 


App. REF.—Blanch., Moust., 180,7 2. 


N.B.—In his 4th volume Mr. Theobald definitely confirms 
kumasti, Chalmers, as synonymous with M. funesta, Giles, and 
remarks that it has not yet been found in India. 

Miss Ludlow records this species from Samar, Laguna, Pan- 
gasinan, Pampanga, Tayabas, Mindanao, Sciassi, Rizal, and other 
localities in the Philippine Islands, where it appears to occur all 
the year round except during March and April. 

““ A proven host of the malarial parasite in Africa, with a 
moderate distribution in the Philippines ; is always taken where 
malaria is present or prevalent’’ (Ludlow). 

The types of the varieties uwmbrosa and sub-umbrosa are in 
the British Museum. 


M. indefinita, Ludl., 1904. 
M. rossii var. indefinita, Ludl., Can. Ent., xxxvi, 299. 


App. Rrer.—Theob., Monog. Culic., iv, 47. 


N.B.—Now accepted as a good species, although at one time 
considered intermediate between vossii and Judlowtt, Miss 
Ludlow records it from all parts (26 different localities) of the 
Philippines, where it occurs all the year round. This author says, 
“‘ Never the subject of experiment, though widely distributed, 
taken in large numbers, and present during malarial outbreaks ; 
does not occur alone in a sufficient number of stations to be indi- 
cative, and its ability as host must be left in doubt.” 

Type in Army Medical Museum, Washington. 


M. jehafi, Patton. 


App. RrF.—Theob., Monog. Culic., v, 22, figs. of w ? palpi and 
palmate hair. 


M. leptomeres, Theob. 


Giles, in his ‘‘ Revision of the Anophelina ” (1904) thinks this 
= Anopheles pictus, Lw. (1845), but Theobald (Monog., iv, 124, 
and v, 29) says that his (Theobald’s) species is nothing like Loew's 
description. 

Type in British Museum, 


420 Records of the Indian Museum. [Vior,SiVE 


M. leucophyrus, Don. 


Removed to Neomyzomyia. 


M. listoni, Liston. 


App. Re¥F.—Myzomyta christopherst, Theob. Blanch., Moust., 
183. 
App. Loc.—Kangra Valley (Punjab, 4,800 ft.), May, June, 
July [Dudgeon]; Berars (Centr. India) ; Ceylon [Green]. 

Under the name christophersit, Theob., Mr. Theobald (Rec. 
Ind. Mus., iv, 2) gives the localities Sylhet, 2-v-05 [Hall]; and 
Jalpaiguri, 13-viii-o7 [Wallich]. 

N.B.—Mr. Theobald (Monog., iv, 51) sinks this name for his 
own chyristopherst, and remarks, ‘‘ The name /istont was used by 
Giles for a large variety of culicifacies, so Liston’s name cannot 
stand.” 

Even if the variety referred to is sufficiently distinct to be 
considered as such, and constant enough to be accorded a name 
(and Mr. Theobald does not rank it as such), there is no reason 
why the name should not be given elsewhere to a distinct species. 

Liston’s description holds priority. In describing the species, 
I take it that he considered he was describing the species named 
after him by Giles (U/stonz, Giles, 1901, Ent. Month. Mag., xxxvii, 
197); therefore now that the error is discovered, and it is known 
to be a different species, the name should surely be retained, 
Capt. Liston’s mistake absolving him from being considered to 
have purposely named the species after himself. 


M. ludlowi, Theob. 


Apparently occurs all the year round all over the Philippines 
from the long list of dates and localities afforded by Miss Ludlow, 
who records it from forty different places in these Islands. Also 
found in the Malay States. ‘‘ Never a subject of experiment, 
appears coincident with malaria in the Philippines, and seems 
likely to be connected with its transmission ’’ (Ludlow). 

However, in a footnote (Mosq. Phil. Is., p. 30) she adds that 
since writing her paper, two articles have appeared, by Capt. 
Ashburn and Lieut. Craig, and by C. S. Banks, the results of the 
investigation of the first authors being against the probability of 
malaria being actually spread by this insect; whilst the last 
author proves at least its capability by actual experiments of 
transmitting the disease. 

James and Liston desire to place this species in their new 
genus Nyssomyzomyta. 

Type in British Museum. 


M. punctulata, Don. 
Removed by Theobald to Cellia. 


IgI2. | E. BRUNETTI: Catalogue of Oriental Culicidae. 421 


M. punctulata, James and List. 


This is not claimed to be a new species, and the only infer- 
ence is that it is Donitz’s species redescribed, though why these 
authors rele@gate the name to themselves is certainly not obvious. 


MI. rosisi, Giles. 


App. REF.—Blanch., Moust., 178, 7 @, fig. 162, wing; 163, 
transverse veins. 
Leices., Culic. Malaya, 25, 7 2. 

N.B.—In Monog., iv, errata, Mr. Theobald says “‘ This species 
has been found on recent microscopic examination to belong to a 
distinct genus from Myzomyia, owing to the peculiar squamose 
characters of the thorax. The genus is being described by Mr. 
Rothnell as Pseudomyzomyia.” 

I have seen no description anywhere of this proposed genus 
Pseudomyzomyia and Theobald does not mention it in his Monog., 
vol. v. James and Liston (Monog. Anoph. Mosq. India, 2nd 
Ed., 44) propose to erect the name Nyssomyzomyza in its place, 
for the reception of vossii, ludlowi, Theob., and punctulata, 
‘“‘ James and List.,’’ but for reasons previously stated the genus is 
not recognized here. Anyway, should Pseudomyzomyia actually 
have been published anywhere and cover the same set of species 
it is impossible to throw out the genus at James and Liston’s 
desire for the purpose of adopting a new name of their own. 

Theobald, in his “‘rst Rep. Ind. Mus. Coll. Culic.’’ (Rec. Ind. 
Mus., ii, 287), retains the species in Myzomyia without comment 
and in his 2nd Report does so also. 

Leicester (Culic. Malaya) says the larvae occur in any small 
collections of stagnant water near houses, giving as localities, 
Kuala Lumpur, Singapore, Penang, Klang, and notes that its 
only affinity in Malaya is albirostiris. 

Miss Ludlow records it from the Philippines nearly all the 
year round, saying ‘‘ Doubted as a host in India; has a moderate 
distribution, is taken infrequently and in small numbers in the 
Philippines, and its connection with malaria is not indicated.” 

App. Locs.—Laguna, Mindanao, Albay, Pangasinan, Tayabas, 
Pampanga, Batangas, Bulacan, Bataan, Cavite, Rizal [all Phil. 
Is. ¢. Ludlow]. Calcutta apparently all the year round; I have 
taken it there in April, June, July and August. 

In the Indian Museum collection [¢. Theob. | from Purt, 
Ganjam lake district, Travancore State (several localities, taken by 
Dr. Annandale, 5—25-xi-o8) ; Ferozepore, Chittagong [Had/], and 
on board ship ten miles off Coconada, Madras coast, 17-iv-08 
[Paiva]; Lucknow, 4-ix-05 [Brunettt]. 

Theobald adds ‘‘ one example from Calcutta, quite typical. 
was labelled stating that it was determined by Giles as Anopheles 
costalis, he does not mention this in his handbook.”’ 

Mr. Green says this species is probably the malaria carrier in 
parts of Ceylon, especially the Batticaloa district. 


422 Records of the Indian Museum. [ VOE.-EY¥- 


In his last volume Mr. Theobald gives a long list of localities 
from Indian Museum specimens, showing it to be found in India 
practically all the year round, rarest in February and March, 
occurring frequently in houses and public conveyances. 

He adds as localities: Trincomalee, Ceylon, 14—28-i-07 ; 
T-ii-o7 ; 1-x-07 [all Green] ; Phrapatoon, Siam, 18—29-i-07 ; Ig—29- 
iii-o7 ; viii and ix-o6 [all Dv. P. G. Woolley]; Chittagong, 19-ix-08 ; 
21-ix-08 ; 5-vii ; 8-vili [all Hall). 

Type in British Museum. 


M. tessellata, Theob. 
Tvpe in British Museum. 


M. thorntoni, Ludlow. 


App. Loc.—Philippines (August, ¢. Theob.); Mindanao, Pangas- 
inan, Tayabas, Pampanga, Samar, Rizal (all Phil. Is., Feb. and 
May to November, ¢. Ludlow). 


M. turkhudi, Liston. 


App. Rer.—Blanch., Moust., 183. 

App. Loc.—Aurangabad (Hyderabad State, India). 

N.B.—Tvpe in British Museum, 

Mr. Theobald’s description of the ~ was drawn up from Giles’s 
type, which at that time was considered to be the @ of culicifactes, 
Giles, and which was first described as such by both Giles and by 
Theobald. 

Vide note in introduction on Myzomyia culicifacies. 


NYSSOMYZOMYIA, James. 
Rec. Ind. Mus., iv, ror. 


James and Liston, Anoph. Mosq. India, 2nd Ed., 43. 

Proposed by the above authors for the three species rossiz, 
Giles, /udlow:, Theob., and punctulata, ‘‘ James and List. ,” and their 
suggestion is that it takes the place of the genus Pseudomyzomyia, 
a genus spoken of by Mr. Theobald, but apparently never 
described. 

The three species in question are retained here under Myzo- 
myia (I presume their punctulata is identical with Donitz’s 
species). 

. NEOSTETHOPHELES, James, rgro. 

Rec. Ind. Mus., iv, 98. 


James and Liston, Monog. Anoph. Mosq. India, 2nd Ed., 4o. 

I have not adopted this genus, with others erected in the 
above work, but these authors place two species only in it, aitkeni, 
James, and culiciformis, James and Liston. 


1912.] EE. BRUNETTI: Catalogue of Oriental Culicidae. 423 


STETHOMYIA, Theob. 


App. REF.—Theob., Monog. Culic , iv, 59, and v, 35 (tab. of 4 
known spp.). 


N.B.—Messts. James and Liston in their latest work suggest 
the abolition of this genus.! 


S. culiciformis, James and Liston. 


App. REF.—Theob., Monog. Culic., iv, 62 (J. and L.’s descr. 
copied). 

N.6.—He doubttfully retains (even in vol. v) the species in 
this genus. The ~ is mentioned once or twice in the description, 
but it is not definitely stated that this description applies to both 
sexes. 

Placed in their new genus Neostethopheles by James and 
Liston. 

S. fragilis, Theob. 


Type in British Museum. 


S. pallida, Ludlow. 


Recorded by this author from Pampanga (Phil. Is.), Sept. 
1905. 
PYRETOPHORUS, Blanch. 


App. REeF.—Theob., Monog. Culic., iv, 63 (list of species), 64 
(tab. of spp.). 
Leices., Culic. Malaya, 37. 
James and Liston, Monog. Anoph. Mosq. India, 
end Wd: -Ar- 
N.B.—In vol. v Theobald quotes Howardina, not Howardia, 
as his reference in Journ. Trop. Med., v, 181. I cannot say which 
is correct. 


P. elegans, James. 


Removed to Neomyzomyta. 


P. freerae, Banks. 
Theobald (Monog., v, 43) thinks from the description that 


this may be a Nyssorhynchus. Only the imperfect type specimen 
seems known. 


P. minimus, Theob. 
App. Rrer.—Blanch., Moust., 188, 2 , fig. 169 (wing and trans 


verse veins). 


1 Monog. Anoph. Mosq. India, 2nd Ed., p. 39. 


424 Records of the Indian Museum. _ [Vor. IV, 


N.B.—Its place in this genus is uncertain, owing to the bad 
condition of the type. Giles places it here; Blanchard says, 
“near Myzomyia funesta.”’ 


P. nigrifasciatus, Theob., 1907. 
Monog. Culic., iv, 65,2. 


From India, taken by Major Nurse at Peshin in April. 
Resembles Myzomyia turkhudi, Liston. Also occurs in Cyprus. 
Type in British Museum. 


P. nursei, Theob., 1907. 
Monog. Culic., iv, 66, 2. 


Taken by Major Nurse at Quetta in November, a unique 
specimen resembling nigrifasctatus. 
Type in British Museum. 


P. watsonii, Leices., 1908. 
Culic. Malaya, 38,2. 


A single 2 taken by Dr. Watson in jungle a few miles from 
Klang. 


MY ZORHYNCHUS, Blanch. 


App. Rer.—Theob., Monog. Culic., iv, 81 (list and table of 
the 16 known species). 


Leices., Culic. Malaya, 29. 


N.B.—Theobald thinks it probable that malaria contracted 
in the open is generally accountable to species of this genus. 

One Japanese species (s7wensts) has been shown by Tsuzuki 
to carry malarial parasites in Japan. Col. Alcock embodies 
Lophoscelomyia, Theob., in Myzorhynchus. 


M. albotaeniatus, Theob. 


App. REF.—alboannulatus, James and Liston, Anoph. Mosq. 
Ind: SE: 

ADD. Syn.—Leices., Culic. Malaya, 34,2. 

N.B.—\Warvae in pools near Kuala Iumpur, and in pools at 
Port Swettenham within tidal influence, and with a salinity of 
2‘8 per 1,000: 

Type in British Museum. 


M. barbirostris, V. Wulp. 


App. RreF.—Blanch., Moust., 197. 


1912.] E. BRUNETTI: Catalogue.of Oriental Culicidae. 425 


Leices., Culic. Malaya, 33, 72. 


App. Locs.—Calcutta, December, 1 2 [Ind. Mus. coll.]; Pam- 
pasinan, Daraga, Tayabas, Mindanao, Bataan, Nueva Kcija, 
Rizal, Siassi, Pampanga (all Phil,-Is.,;, Feb.;’ March, June to 
December) [¢. Ludlow]; Kuala Iumpur (in bungalows), Klang 
(in jungle) [¢. Letcester] ; Phrapatoon, Siam, 24-iii-07 [Dr. Woolley] ; 
Nedumangad, 10 miles N.E. of Trivandrum, S. India, 14-xi-08 
[Annandale]; Assam, 15-i-07 [Hall]; Chittagong, 15-viii-o8 [| Hall]; 
Andaman Is. (1908) [Ray White] ; Digoel (in Amsterdam Museum), 
and taken on the New Guinea Expedition [all ¢. Theob.]. 

Type in Leyden Museum. 

N.B.—Dr. Leicester says the larva can be found in any 
large open water, and adds, ‘‘ The larva described by Theobald 
in vol. iii is, in my opinion, the larva sinensis and not barbirostris.” 
Theobald says malarial parasites can develop in this species but 
rarely do so in nature. The larvae of the true form live in dark 
pools of all depths, with or without vegetation. 


M. minutus, Theob. 


App. REF.—Theob., Monog. Culic., iv, 87. 

App. Syn.—Anoph. nigerrimus, Giles, in James and Liston’s 
(1900) Anoph. Mosq. India, 79, 2 , col. pl. iii (full insect). 

App. Loc.—Kuala Lumpur [Durham]. 

Type in British Museum according to Theobald’s 5th volume, 
but he previously informed me that it was lost. 

N.B.—‘ The larvae are usually found in deep shady pools, 
containing grass and water-weed, at some distance from habita- 
tions, and the adults are seldom met with in houses’’ (James and 
Liston); 

The larval characters given by these authors (loc. czt., p. 81) 
refer to the present species and not to the true nzgerrimus, Giles, a 
species which is quite distinct, but which has been confused with 
minutus. 

The embryos of Filaria bancrofti can develop in this species. 


M. nigerrimus, Giles. 
Abb. REF.—Blanch., Moust., 197. 
Type in British Museum. 
M. peditaeniatus, Leices., 1908. 
Culic. Malaya, 31,07 . 


Described from a large series bred from larvae, which occur 
in any large collection of water, if not very stagnant, in the 
Malay Peninsula. 


420 Records of the Indian Museum. [Vous TV, 


M. philippinensis, Ludl. 


Removed to Nyssorhynchus. 


M. pseudobarbirostris, Ludlow. 


App. Rrer.—Theob., Monog. Culic., iv, 83, 2. 

N.B.—Type in Army Medical Museum, Washington. 

Closely allied to barbivostris, Wulp. 

Miss Ludlow records it from Marinduque, Mindanao, Ambos 
Camarines, Pangasinan, Bulacan, Tayabas, Cavite and Rizal, 
all in the Philippines, appearing apparently from May to October. 


M. separatus, Leices., 1908. 
Culic. Malaya, 36 (sex not mentioned). 


Bred by the author from larvae from large collections of 
water in the open near Kuala Lumpur. ‘‘ It may be a variety of 
sinensis.’ 


WI. sinensis, W. 


App. REF.—Blanch., Moust., 190, 2 , fig. 170 (wing scales). 
Leices., Culic. Malaya, 30,0 2. 


ADD. Syn.—Anoph. plumiger, Don., Ins. Borse, Jan. rgor. 

Anoph, jesoensis, ‘Tsuzuki, 1902, Centr. fiir Bakteriol., xxxi, 
703. 
App. Locs.—Hong-Kong, Malacca, Java, Sumatra [all ¢. 
Blanchard] ; Calcutta, vi, vii, viii, 1908, in bathrooms at Museum 
[Annandale]; at light on board steamers at Damukdia Ghat, 
R. Ganges [Annandale] ; Sylhet, i, ii, v, vi, vii [Hall] ; Manipur 
(6,400 ft.), vili-o8 [Pettigrew]; Ferozepore [Adie]; Maddathorai, 
r¢-ix-0g [Annandale] (all in Ind. Mus. coll., identified by Theobald) ; 
Jolo (September) and Rizal (August), Phil. Is. [¢. Ludlow]; 
Shanghai, 8—10-v-06, in ditch [Brunett?]. 

Theobald adds the following localities in his vol. v : Phrapa- 
toon, Siam, vili, ix [Dv. Woolley] ; Pampanga, Angeles, Ph. Is. 
|Whitmore]; West Lake, Hankow [Cornford]; Ukhrul, Manipur 
(6,400 it.), vilt-o8, 20—26-1-03 , 23-ii-05, 27-v-00, 4——7-vi-05 [ Petti- 
grew]; Sylhet, 13-1-03, 13-1-04, 24-vii-o8 [Hal/] ; Ferozepore[A die] ; 
Calcutta, 9-vi-08, vili-o8, 28-vii and g-vi-o8 [Annandale]; Madda- 
thorai, S. India, 18-ix-08 [Annandale]. 

N.B.—Dr. Leicester says, ‘‘larvae taken in ponds and road- 
side ditches, both in town and country.”’ 

Miss Ludlow reports, ‘“‘ a proven host in India; has been 
taken at too few stations to show that it affects markedly the 
malarial conditions of the Islands’’ (Philippines). Dr. Leicester 
includes under this specific name, vanus, Wlk. 

Respecting pseudopictus, Grassi (a European species), Theo- 
bald still (Monog., iv, 87) considered it distinct from sinensis, W.., 


1912.] E. BRUNETTI: Catalogue of Oriental Culicidae. 427 


with pictus, Ficalbi, for a synonym, but in vol. v he suppresses 
this synonym. Amongst the uncertain species ranked under ‘‘ Ano- 
pheles ” he places ‘‘pictus, Lw.’’ He gives no further information 
about his Myzomyta indiensts. 

M. sinensis has been studied in Japan, where it has been 
found to transmit malaria. It is very variable. 


M. umbrosus, Theob. 


App. REF.—Leices., Culic. Malaya, 35,2. 

He notes only one example, from jungle near Kuala Lumpur, 
*< very like barbirostris.” 

Type in British Museum. 


M. vanus, Wk. 


App. REF.—Blanch., Moust., 196,77 2 . 

App. Locs.—China [#¢. Theobald] ; Celebes [¢. Walker]; Java, 
Malacca [¢. Blanchard]; Calcutta, Oct., Nov., Dec., common in 
last two months; Port Canning [¢. Theob., Ind. Mus. coll.j;; Albay, 
Laguna, Pangasinan, Bulacan, Tayabas, Manila, Cavite, Bataan, 
Nueva Ecija, Rizal (all Phil. Is., April, and from June to Janu- 
ary) [¢. Ludlow]; Dondra, Ceylon, 4-xii-o7 [Green]; Galle, Ceylon, 
8 xii-07 [Green]. 

Type in British Museum. 

N.B.—Dr. Leicester considers vanus synonymous with sinen- 
sts, W., but Theobald in his latest volume keeps them distinct. 


LOPHOSCELOMYIA, Theob. 


App. RreF.—Giles, I9g04, Journ. Trop. Med., vii, 366 (Lopho- 
myta). 
Blanch., 1905, Moust., 635. 
Theob., 1907, Monog., iv, 91,07 ?. 
Leices., 1908, Culic. Malaya, 21. 

N.b6.—In his monograph (iv, 92) Mr. Theobald redescribes, 
and says that the name was spelt correctly at the erection of the 
genus (Entom., Jan., 1904). In the Genera Insectorum (Fasc. 26) 
he spells it Lophocelomyia. 


L. asiatica, Leices. 


App. RrEF.—Theob., Monog. Culic., iv, 92,7 2. 


Leices., Culic. Malaya, 21,7 9. 


‘‘Lophomyia asiatica Theob. MS.,’’ in Giles (1904), Jour. 
Trop. Med., vii, 366. 

App. Locs.—Malaysia, according to Leicester, who says it 
breeds exclusively in pools of water in bamboo, adding that those 
bred in captivity will not bite. 


428 Records of the Indian Museum. [Vou. IV, 


NYSSORHYNCHUS, Blanch. 


App. REF.—Blanch., Moust., 202. 
Leices., Culic. Malaya, 39. 
James and List., Monog. Anoph. Mosq. India, 


and Ed., 43. 
Laverania, Theob., 1902 Journ. Trop. Med., ii, 
181. 


N.B.—In this genus Col. Alcock sinks Neocellia, Theob., Cellia, 
Theob., Calvertina, Ludl., and Christophersia, James. 


N. fuliginosus, Giles. 


App. REF.—Blanch., Moust., 205, fig. 179a, normal wing, 
179), wing of var. pallida. 

App. Locs.—Calcutta, 15-vii-o8, 12-viii-o8, at light in house 
[Annandale]; Balighai, near Puri, 23—24-x-08, ‘‘ numerous in old 
wells, resting by day’ [Annandale]; Ferozepore, numerous [ Adie] ; 
Deesa [Nurse]; Ceylon [Green, Major Manders, Chalmers]; Ellich- 
pur in April, Calcutta, December, at light; Bombay, Java, Sumatra 
[all ¢. Blanchard]; Pangasinan, Tayabas, Guimaris Is., Albay, 
Rizal, Cavite, all Phil. Is. [¢. Ludlow]. 


N.B.—Capt. James has found that tertian, quartan and malig- 
nant tertian parasites will develop in this species artificially, but 
says it has not yet been found naturally infected (v. James, Sci. 
Mem. Ind., new. ser., No. 2, p. 39). 

Major Adie has found the sporozoits in wild fuliginosus (Ind. 
Med. Gaz., xxxvili, July 7, 1903). 

Theobald (Monog., iv, 99) notes a variety from Chingelput 
(S. India). 

Miss Ludlow says, ‘‘ Questioned as a host in India; hasa 
moderate distribution taken infrequently in small numbers in the 
Philippines, and its connection with malaria is not indicated.” 

This species appears to have several varieties, a form occur- 
ring in the Punjab all the winter (Adie, Ind. Med. Gaz., xxxviii, 
July 7, 1903 and Jan. 4, 1905); this variety flourishing from the 
middle of November till the end of April, whereas in that part of 
India, the typical form flourishes from the middle of March to 
the beginning of June. 

Theobald’s variety pallida has been considered a good species 
by Giles, but the former author states that it is not so (Monog., 
iv, 100). 


N. indiensis, Theob., 1903. 
Monog. Culic., iii, 99. 


Anoph. id., James and Liston (non Giles), Anoph. Mosq. Ind., 
95, plate 2, figs. 


1912. | E. BRUNETTI: Catalogue of Oriental Culicidae. 429 


This form, previously regarded as a variety of maculipalpis, 
Giles, is raised by Theobald (Monog. Culic., iv, 98) to the rank of 
a species (vide N. maculipalpis, post.). 

Type in British Museum. 

N.B.—Theobald (Monog., v) restricts the localities of this form 
to the Central Provinces of India, Nagpur, Goa, Karwar, Travan- 
core. 


N. jamesii, Theob. 


App. REeF.—Blanch., Moust., 206. 

App. Locs.—Calcutta, 5-vili-o8 [Annandale]; Shamnagar, 
Bengal, 3-viii-05 [Gourlay]. 

Type in British Museum. 


N. karwari, James in Theob. 


ApD. REF.—Leices., Culic. Malaya, 39,7 2. 
N.B.—\atvae taken at Jugra and near Kuala Lumpur, in 


the grassy edges of slowly flowing streams. 
Type in British Museum. 


N. maculatus, Theob. 


App. REer.—Blanch., Moust., 207, 7 @ , fig. 180 (wing). 
Leices., Culic. Malaya, 41, ~ 2 (copies Theobald’s 
description, not having seen the species). 
App. Locs.—Nara Ghat, Nepal, near Bengal frontier, 25— 
26-ii-08 ; Thamaspur, Bengal frontier, 18 and 20-11-08, 7 2. 
N.B.—This species is the type of the genus and the type was 
described as in Dr. Rees’s collection, but Mr. Theobald informs 
me that it is in the British Museum. 


N. maculipalpis, Theob. 


N.B.—The variety indiensis is now raised to the rank of a 
species (vide supra). In my catalogue, therefore, delete the locali- 
ties Nagpur, Karwar, Goa and Travancore, all of which relate to 
indiensts. 


N. nivipes, Theob. 


ADD. REF.—Theob., Monog. Culic., iv, tor, 2 . 
Leices., Culic. Malaya, 44, 7 @. 

N B.—Bred from larvae in small collections of water in the 
open near Kuala Lumpur, Klang, etc. Dr. Leicester describes the 
species from a long series of both sexes. 

Type in British Museum. 


430 Records of the Indian Museum. [VOESnV, 


N. philippinensis, Lud. 


App. ReF.—Theob., Monog. Culic., iv, 103, ?. 

App. Locs.—Pangasinan, Tayabas, Abra, Rizal, Pampanga 
(all Phil. Is.), where it occurs all the year round except from Feb- 
ruary to May. 

Type in Army Medical Museum, Washington. 

N.B.—Theobald says (Monog., iv, 104), allied to mivipes, 
Theob., may be a variety of it. The species has been referred by 
Giles to Pyretophorus but Theobald (loc. cit., v, 63) retains it here. 


N. pseudowillmori, Theob., rgro. 
Monog. Culic., v, 65,9. 


From Meenglas, Jalpaiguri, 13-vii-o7 [Wallich]. 
Type in Indian Museum. Perhaps a var. of w7llmori. 


N. stephensi, Liston. 


App. Re¥F.—Blanch., Moust., 2ro. 

App. Locs.—Calcutta (Museum gardens), 15-vii-o8 [Annan- 
dale]; Phil. Is. (¢. Miss Ludlow in Can. Ent , xli, 234). 

This was removed to Neocellia but Theobald replaces it here 
in his vol. v, where see p. 20 for notes. 


N. theobaldi, Giles. 


Erratum.—Correct p. 300 to 299 in my reference to Giles’ 
Handb., 2nd Ed. 

App. REF.—Blanch., Moust., 210. 

App. Locs.—Benguet (Phil. Is.), March, April, November. 

N.B.—Miss Ludlow says ‘‘ a proven host in India; has been 
taken at only one station (referring to the Philippines), and can- 
not be held responsible for much of the transmission of malaria.” 


N. willmori, James. 


App. REF.—Leices., Culic. Malaya, 42,7 2. 

Has been placed in Neocellia but Theobald (Monog., v) again 
refers it here, and adds Ceylon, Pundaluoya, July—‘‘ known as 
the instep-biting mosquito’’ [Green]; Malay States and Meenglas, 
Jalpaiguri, 13-vii-o7 [Wallich], to the localities. 

N.B.—Dr. Leicester says the larvae live in hill streams in 
jungle near Kuala Lumpur, Jugra and elsewhere, and only records 
the adult from a hut in Ula Gombak. 

Type in the Central Research Institute, Kasauli, India, 


1gI2. | E. BRUNETTI: Catalogue of Oriental Culicidae. 431 


KERTESZIA, Theob., 1905. 
Ann. Mus. Hung., iii, 66. 
Monog. Culic., iv, 117. 
Intermediate between Nyssorhynchus and Cellia. 
N.B.—Col. Alcock sinks Kerteszia in Arribalzagia , Theob. 


K. megregori, Banks, 1910. 
Phil. Journ. Sci., iv, 548, 0. 


Loc.—Basilan, Ph. Is. (Type No. 6666), Entomological Collec- 
tion, Bureau of Science, Manila. 


CHRISTOPHERSIA, James. 


Paludism, vol. i, 33 (July 1910) (nom. nud.). Rec. Ind. 
Mus!,’iv, 103 (descr.). 


C. halli, James, rgro. 


Paludism, vol. i, 33. Plate: figs. of palpi, thorax (dorsal and 
lateral), abdomen (dorsal, lateral and ventral) and hind leg. 

James and List., Monog. Anoph. Mosq. India, 2nd Ed., 123. 

Loc.—Sylhet (Assam), February, June, July and December 
[Li-Col. Hall}. 

Type in Indian Museum. 


CELLIA, Theob. 


ADD. REF.—Theob., Monog. Culic., iv, 104, 105 (list and tab. 
known spp.). 
Blanch., Moust., 214. 
Leices., Culic. Malaya, 46. 


C. flava, Ludlow, 1908. 
Can, Ent... <l, 32%.a7 


Described from four examples from Tayabas (Phil. Is.), Sep- 
tember 1907 ; incorrectly quoted Tayubar in original description, 
corrected by author, Joc. cit., 52. 

Type in Army Medical Museum, Washington. 


C, kochi, Don 


App. REF.—Blanch., Moust. 208 (Nvssorhynchus). 
Leices., Culic. Malaya, 46,7 @. 
App. Locs.—Kuala Lumpur, xi, 1902 [Durham]; Singapore. 
N.b.—Widely distributed in the Malay Peninsula, more 
abundant in the vicinity of houses, breeding in roadside puddles 
or any pool of water in the open. 


432 Records of the Indian Museum. { VoL. iV, 


C. pulcherrima, Theob. 


App. RreF.—Blanch., Moust., 215, 2. 

App. Loc.—Kokand (Turkestan) [¢. Univ. Coll. Helsingfors] : 
Lahore. 

Type 2 in British Museum, @ apparently still unknown. 


C. punctulata, Don. 


Anopheles id.; Myzomyia id. ; auct. 

Nyssorhynchus td, Blanch., Moust., 208. 

Anoph. tessellatum, 'Theob., Monog. Culic., 1, 175 (wom. nud). 

App. REF.—Leices., Culic. Malaya, 27,7 92. 

App. Locs.—Kuala Lumpur, Klang (at both places in 
bungalow) [¢. Leicester]; Sumatra, Borneo, Taiping, Papua. 


N.B.—‘‘ It proves to be a Cellia and not a Myzomyra, as it ap- 
peared from Donitz’s description. Blanchard erroneously placed 
it in Nyssorhynchus”’ (Theob., Monog., iv, 109). 

James and Liston (Monog. Anoph. Mosq. India, 2nd Ed.) des- 
cribe and figure parts of a ‘‘ punctulata James and Liston,’ under 
their proposed new genus Nyssomyzomyia without any reference 
to Donitz’s species of this name. ‘The species cannot be taken 
from Donitz’s credit if the two are identical, and, if not, two spe- 
cies with the same name cannot co-exist in the same genus. 


NEOCELLIA, Theob. 


Monog. Culic,, iv, III, 7 2. 


N. dudgeoni, Theob., 1907. 
Monog. Culic., iv, 112, 2. 
Loc.—Kangra Valley (4,500 ft.), June, July [Dudgeon}. 
Described from several ? ¢. 
Type in British Museum. 
N. indica, Theob., 1907. 
Monog. Culic., iv, I11,¢7 9. 


Loc.—Dehra Dun (foot of Mussoorie Hills), February, March. 
Described from I~” and 32 9. 
Type in British Museum. 


N. intermedia, Rothwell, 1907. 
Entomologist, Feb. 1907, 2 . 


Theob., Monog. Culic., iv, 115, 2; v, 73, 7 2, fig. 20, wing’. 
Loc.—Deesa (W. Centr. India), January, August. 
Type in British Museum. 


1912.] E. BRUNETTI: Catalogue of Ortental Culicidae. 433 


The @ described by Theobald (v, 73) from Ferozepore, India 
(18 # @ and 109 ? ¢ taken by Adie). 


ALDRICHINELLA, Theob., rgro. 


Monog. Culic., v, 77, nom. nov. for 
Aldrichia, Theob., 1903. 

Aldrichia has been preoccupied by Coquillett in Bombylidae 
since 1894 (Trans. Amer. Ent. Soc., xxi, 93), so Theobald has just 
forestalled me in renaming it by proposing Aldrichinella. 

In James and Liston’s new edition the correction is not 
made. 

The type (A. error, a unique specimen) was in the British 
Museum, but has subsequently been broken. Col. Alcock, how- 
ever, says it is not a good species at all, being a Myzomyia rossit, 
Giles, with the abdomen of some different species attached. 


BIRONELLA, Theob. 


App. REF.—Theob., Monog. Culic., iv, 120, 7; fig. # wing, 
pe 122) 


B. gracilis, Theob. 


N.B.—The ¢& being unknown, Mr. Theobald judges this genus 
to belong to the Anophelina, but some details of its characteristics 
make him uncertain as to which subfamily should really in- 
clude it. 

Type in Hungarian Museum. 


CALVERTINA, Ludlow, 1909. 


Can, Ent., xli, 234, emen. from Calvertia, Ludl., loc. cit., 22. 


C. lineata, Ludlow, 1908. 
Can. Ent., xl, 50 (Chagasia id.). 


Calvertia lineata, Ludl. 

Calvertina id., Theob., Monog. Culic., v, 77. 

N.B.—Described first under Chagasia, Calvertia was erected 
for it, but this is preoccupied by Warren in Lepidoptera and by 
Calvertius, Sharp, in Coleoptera ; there is also a Calveria, Carp., in 
Echinoderma., 

The species described from a single perfect specimen of which 
the author does not state the sex. From Pangasinan (Phil. Is.) 
in August. Its position here is uncertain, and it may be better 
placed near Pyretophorus. 


434 Records of the Indian Museum. [Vor. IV, 


‘* ANOPHELES,”’ sensu lato. 
‘* Anopheles’”’ arabiensis, Patton. 


App. REF.—Theob., Monog. Culic., v, 81, @ 2 , fig. 34, o @ 
palpi. 

N.B.—Theobald seems vague in his opinion on this. From 
damaged 2 @ sent him by the author of the species, he considered 
them identical with A. wellcomei, Theob., but Patton had already 
sent the species to Dr. Stephens, who compared it with the type 
of wellcomei and pronounced them distinct. In vol. v, 82, Theo- 
bald says: ‘‘ it is certainly not an Anopheles,’ and immediately 
after he says a damaged specimen sent him by Patton could not 
be separated from wellcomei, Theob. In his last volume he puts 
the species under ‘‘ Anopheles.” 


‘* Anopheles ”’ culiciformis, Cogill. 


Theobald confesses inability to trace this species. 


‘* Anopheles ’’ deceptor, Don. 
Removed to Myzomyva. 


‘* Anopheles ”’ pictus, Lw., 1845. 
Dipt. Beit.;“Poseny py. 


Theobald still (Monog. Culic., iv) thinks this the same as 
Grassi’s pseudopictus, but defers a definite opinion until he can 
compare specimens from Rhodes (Asia Minor). Dr. Thin records 
it from Haut Tonkin and Harioi. ‘‘In both cases M. sinensis is 
evidently referred to as an allied species’’ (Theob., Monog. 
Culic., iv, 124). Theobald also says here that Giles is wrong in 
considering Myzomyia leptomeres, Theob., as a synonym of ictus. 


** Anopheles”’ subpictus, Grassi. 
This species cannot be traced. 


t 


‘* Anopheles ’’ vincenti Laveran. 


bf 


Evratum.—My ‘‘ correction’’ of Theobald’s quotation of date 
and volume (1gor and lili) is an error, as both his references are 
correct; yet in vol. v, 84, he perpetuates my previous error as 
regards the volume by quoting xxiil. 


‘* Sub-Family MEGARHININAE.” 


App. REF.—Blanch., Moust., 218, figs. 184-185. 
Dr. Leicester (Culic. Malaya, 48) is averse to the subdivi- 
sion of this ‘‘ sub-family ’? of Theobald, and observes that though 


1912.) E. Brunetti: Catalogue of Onental Culicidae. 435 


the Culicidae may be divided into four sub-families, principally 
on the comparative length of the palpi in the sexes, the division 
is an arbitrary and not a natural one, and from his observations 
I entirely agree with him. Blanchard (Moust., 218-219, figs. 184- 
185) gives an extensive account of the characters of this group. 


MEGARHINUS, Rob. Desv. 


App. Rer.—Blanch., Moust., 220, 221 (tab. of all spp.). 
Theob., Monog. Culic., iv, 128 (tab. of spp.):; 
v, 96 (tab. of spp.). 


N.B.—Theobald says (loc. cit., v, 95) the genus is not Orien- 
tal, being only found in North and South America and the West 
Indies. 


M. amboinensis, Dol. 
M. lewaldii, Ludlow. 
M. minimus, Theob. 
M. splendens, W. 


The first two are definitely referred to Toxorhynchites, the 
last two probably belong there also. 


TOXORHYNCHITES, Theob. 


Type of genus. T. brevipalpis, Theob., from Natal. 
App. REF.—Leices., Culic. Malaya, 59. 


T. amboinensis, Dol. (Megarhinus). 


App. REF.—Blanch., Moust., 266,07 @. 


T. argenteotarsis, Ludlow, 1906. 
Can Ent, sxseviitsi307 79. 


Theob., Monog. Culic., v, 100, 2. 

Type in Army Medical Museum, Washington. 

Described from 5? ¢ from Margosatubig, Mindanao, Phil. 
Is., June and July. Is near spectosus, Skuse, and marshalli, Theob. 


T. gilesii, Theob. 


Monog. Culic., 1, 227, % 2 ; note op. cit., V, 99. 

Described originally as distinct, it was relegated to a syno- 
nym of zmmisericors, but is reinstated. 

Iocs.—Sikhim, Sylhet, 7-vi-05; 13-vii-o5 [Hall]; Calcutta, 
Ceylon, Upper Burma, Singapore [/inlayson'. 


436 Records of the Indian Museum. [ VOLE. LV, 


T. immisericors, Wlk. 


App. ReF.—Theob., Monog. Culic., iv, 4, 9; v, 97. 
Blanch., Moust., 230,07. 


App. Locs.—Calcutta (Museum gardens and zoological gar- 
dens), i, vi, vii, viii; Bhim Tal(W. Himal.), 19g—22-x-06 [all 
Annandale] ; Aijal (3,600 ft.), Lushai Hills, 24-1v-04 ; Sylhet, 5-v-05 ; 
Chittagong, 3-ix-08 ; Peradeniya, Ceylon (1,600 ft.), 24-xii-07, bota- 
nical gardens, not uncommon, and Pundaluoya, Ceylon (4,000 ft.) 
[Green and Bainbrigge Fletcher]; Andaman Is., 19-vi-08, 8-vii-08 
[Ray White]. 

Also from Sikhim, Burma, Malacca, Trincomalee Hot Wells, 
Macassar, Mysore, North Ceram, Waigiou. 

N.B.—Theobald says that his figure of the pupa (Monog. 
Culic., iii, 123, fig. 67) is not quite correct. He also describes 
the 2 (Rec. Ind. Mus., iv, 4), and says that the type (o) in the 
British Mus. is in bad condition. 

Mr. E. E. Green gives the life-history (with a plate) of this 
species (Spol. Zeyl., ii, pt. viii, 159 to 164) (1905). He says the 
larvae prey first on those of their own race before proceeding to 
devour those of other species, and that he does not know of the 
adult biting. Females were kept alive for eleven days on sliced 
bananas. 

Mr. C. A. Paiva gives a very interesting account of the habits 
of the larva (Rec. Ind. Mus., v, 187) from personal observations 
and experiments. It iscommon in the outskirts of Calcutta during 
June and July in earthen pots. He finds that the larva will 
devour that of any other species if present before attacking those 
of its own kind, whereas Mr. E. E. Green thought it ate its own 
species first. They are sluggish and remain at the surface of the 
water, seizing other larvae as they come within reach. A curious 
thing is that the larvae of other species actually attack the 
immisericors larvae, seizing their abdominal bristles from behind. 
Stegomyia fasctata the carrier of yellow fever is greedily devoured 
by I. immisericors larva, which renders it a valuable ally in 
destroying the former in the event of that disease being introduced 
into India, the more so as the adult is not known to bite man. 

N.B.—Megarhinus gilesit, Theob., and subultfer, Dol., to be 
eliminated from synonymy, as both are now regarded as good 
species belonging to Toxorhynchites. 


T. inornatus, WIk. 
App. REF.—Blanch., Moust., 223,07 ?. 
Type o 2 in British Museum. 
Loc.—Papua. 
T. javaensis, Theob., I9II. 
Tijd: v. Enee lives233 90. 


Java, aunique 2. Type in Amsterdam Museum. 


1g12.] E. BRUNETTI: Catalogue of Orvental Culicidae. 437 


T. leicesteri, Theob. 
Evratum.—1804 is given for 1904 in my catalogue, p. 325. 
App. REF.—Theéob., Monog. Culic., iv, 142,72. 
Leices., Culic. Malaya, 59,7 2. 

App. Locs.—Papua. 

N.B.—Leicester says that the colours fade so much after 
death as to make identification very difficult, and says the larvae 
are found in bamboos. 


T. lewaldii, Ludlow (Megarhinus). 


App. REF.—Theob., Monog. Culic., iv, 139, (Miss Ludlow’s 
description copied), adding ‘‘ may be a Toxorhynchites.”’ 
Type in Army Medical Museum, Washington. 


N.B.—lWarvae taken on April Ist hatched on the roth. 


T. metallicus, Leices. 


Erratum.—In my catalogue correct metallica to metallicus. 

App. REF.—Leices., Culic. Malaya, 61,7 2. 

N.B.—Leicester says it is bred from bamboo and also that in 
the adult the colours fade quickly. ‘‘I have not taken the adult 
of this mosquito; it is entirely sylvan and fairly widely distri- 
buted in the Peninsula ”’ (Leicester). 

Theobald also notes the rapid fading of colours after death, 
especially in the abdominal bands of the? . 


T. minimus, Theob. (Megarhinus). 


App. REeF.—Theob., Monog. Culic., iv, 138, # (full descrip- 
tion). 

The author originally suggested it might be a Toxorhynchites, 
and noted its small size. 

Type in British Museum. 


T.splendens, W. (Megarhinus). 


App. Rrer.—Blanch., Moust., 225 (Megarhinus). 


N.B.—I expect one of the more recently described species 
will eventually prove to be synonymous with this. 


T. subulifer, Dol., 1857. 
Nat. Tijd. Ned. Ind., xiv, 382 (Megarhinus). 


Theob., Monog. Culic., 1, 242. 

Amboina. For a time considered synonymous with ?7se- 
vicors, in which case Doleschall’s name has the priority over 
Walker’s. 


438 Records of the Indian Museum. [VoL. IV, 


TEROMYIA, Leices., 1908. 
Culic. Malaya, 49. 


Leicester says in the above reference ‘‘ nov. gen.,’’ yet adds 
‘“TIn Theobald’s Monograph it is stated that in members of this 
genus there are no upright scales on the head.’ 

He adds, ‘“‘ Five species are here described, and they are all 
apparently new species, though quasiferox may prove to be merely 
a variety of M. ferox, or immisericors.’’ 


T. acaudata, Leices., 1908. 
LOCsICi.5 49,012. 


The examples from which this species was described were 
obtained by Dr. Finlayson of Singapore, all of them being bred 
from larvae found in pitcher plants in the neighbourhood of 
Singapore. The author notes that the colours fade very con- 
siderably after death. 


T. ater, Daniels, 1908. 


Studies from Instit for Medic. Research (Fed. Malay States), 
lii, 205,07 2. 


““ Notes on the Mosquitoes on the river and coast district 
of the Eastern side of the Peninsula.’’ 


N.B.—This is issued as a succeeding paper to Dr. Leicester’s 
elaborate Monograph on the ‘“‘ Culicidae of Malaya,’’ with con- 
tinuous pagination. 

Bred from larvae found in the pitcher plant Nepfenthes raffle- 
stana, on the east coast of Pahang. 


T. funestus, Leices., 1908. 
Culic. Malaya, 58, o. 


‘* Described from one @ bred from a pupa taken in a bamboo 
jungle 6 miles from Kuala Lumpur. A very funereal looking mos- 
quito ; some of the more beautiful colours are only seen with a 
lens ’’ (Leicester). 

T. magnificus, Leices., 1908. 


Culic. Malaya, 54,7 2. 


‘Bred from larvae obtained from the water collected in 
living bamboos, which had been pierced by an insect borer, the 
only entrance to the water being through the small hole thus 
made.’’ Leicester compares his new species with splendens and 
notes minor differences, but it must be remembered that at 
the time Wiedemann wrote, the limited number of known species 
made unnecessary the lengthy and detailed descriptions required 
in the present state of our knowledge. 


1912.] EE. Brunetti: Catalogue of Oriental Culicrdae. 439 


T. quasiferox, Leices., 1908. 
Culic. Malaya, 51,7 @. 


It seems advisable here to quote the author. 

‘“Mr. Theobald, to whom a specimen of this mosquito was 
sent, suggests that it is probably Megarhinus imm¢sericors, first 
described by Walker, and in his recently published vol. iti, he has 
placed M. tmmisericors in the genus Toxorhynchites. 

‘“* He does not seem to have described the female. 

‘* Tf this is zmmisericors, I am at a loss to understand how it 
can be placed in the genus Toxorhynchites, as the palpi are 
distinctly five-jointed. As mentioned in the remarks on the 
generic characters, I do not think the separation of Toxorhyn- 
chites from Megarhinus is based merely on the palpi, though this 
is the only character given by Theobald, but even on that ground 
alone, this mosquito would be excluded. It now comes in 
LTeromyia.’”’ 

Leicester adds that it probably breeds in the pitcher plant. 


T. raris, Leices., 1908. 
Culic. Malaya, 56,¢. 


‘* Described from a @ bred from a white larva taken in. 
water collected in a bamboo in the jungle at Ula Klang. 
It is a very distinct species ’’ (I,eicester). 

Apparently rare and very near magntficus. 


WORCESTERIA, Banks. 


Theobald (Monog. Culic., v, 110) says this genus, erected on 
minute difference in the palpi, is not valid and comes within reach 


of Toxorhynchites. 
Sub-Family CULICINAE. 


App. Rrers.—Mr. Theobald (Monog. Culic., iv, 147) tabulates 

3 genera, mentioning that others have been 
subsequently described. 

Blanchard (Moust., 231) gives the sub-family 
characteristics. , 

Leicester (Culic. Malaya, 64) includes, after his 
notes on the sub-family, a table of all the 
known genera up to 1905, though many of 
them had not then been found in Malaya. 

Col. Alcock divides this sub-family, which he 
designates Culicales, into groups of genera 
(sub-genera, presumably) as follows: the 
Culex, Stegomyia, Aedes, Uranotaenia, Psoro- 
phora and Mucidus groups. Under these 
respective headings are given in this cata- 
logue the so-called ‘‘ genera’? embodied in 
each. 


440 Records of the Indian Museum. [Vo.L. IV, 


MUCIDUS, Theob. 


App. REF.—Leices., Culic. Malaya, 69. 

Col. Alcock includes under his ‘‘ genera of the Mucidus type ”’ 
Mansonia, Blanch., Mansonioides, Theob., Etorilepidomyia, Theob. 
(is this the same as Eftorleptiomyta ?), Orthopodomyta, Theob., 
Aedimyia, Theob., Finlayia, Theob. He considers this group of 
sub-genera or species links the Culicinae with the Anophelinac. 


M. laniger, Wied. (Culex 7d.). 


App. REF.—Blanch., Moust., 244, 2. 
Type in Wiedemann’s collection. 


M. mucidus, Karsch. 


App. REF.—Leices., Culic. Malaya, 69,7 2. 

Redescribed by Leicester from a @ and 2? bred from larva 
taken in marshy ground near a patch of jungle near Kuala 
Lumpur. 

Banks has recorded it from the Philippines. 


M. scatophagoides, Theob. 

App. REFr.—Blanch., Moust., 245, 2. 

App. Loc.—Bauria, Bengal, 17-vili-o7 [Tyrie]; Damukdia 
Ghat, E. Bengal, 22-viii-o7; Purnea, 5-viii-o7 [Paiva]; all in 
Indian Museum. 

Type in British Museum. 


EKRINOMYIA, Leices., 1908. 
Culic. Malaya, 71. 


The author says, “‘ between Culex and Mucidus, the larva 
very near Megarhinus.”’ 


E. aureostriata, Leices., 1908. 
Culic. Malaya, 71,0 @. 


Described from 4 ~~ and 3 2 2 taken as pupae ina small 
hole in marshy ground at Klang. 


BLANCHARDIOMYIA, mihi, nom. nov. 


Syn.—Desvoidya, Blanch., preoccupied. 
App. Rrer.—Theob., Monog. Culic., iv, 163 (table of the 4 
known species). 
Blanch., Moust., 265. 
Leices., Culic. Malaya, 74. 


1912.]| KE. Brunetti: Catalogue of Oriental Cultcidae. 441 


N.B.—‘* Desvoidya”’ was a nom. nov. for Armigeres, Theob., 
practically preoccupied by Hartmann in 1840—1842 (Armiger) in 
Mollusca. Ss 

Moreover Desvoidia, Meade, Ent. Month. Mag., xxvill, 179 
(1892), in Tachinidae antedates Blanchard’s genus, for which 
1 propose the title Blanchardiomyia. I should consider it, with 
other ‘‘ genera ” in this family, at most a sub-genus. 


B. apicalis, Theob., Igto. 
Rec. Ind. Mus., iv, 5,2. 


Theob., Monog. Culic., v, 143,2. 

Described from a single perfect 2? from Balighai near Puri, 
Orissa, taken by Dr. Annandale, 24-x-08. In the Indian Mu- 
seum. 


B. aureolineata, Leices., 1908. 
Culic. Malaya, 79, °. 


‘“ A very distinct Desvoidya; described from a series bred 
from larvae found in water collected in the shells of a fruit in 
jungle at Ampang.” Leicester in a footnote says ‘‘ The descrip- 
tion of the ~ will be found on the slip at the end under Addenda.”’ 
However, in my copy of his work there is no such slip. 


B. fusca, Theob. 


App. REF.—Leices., Culic. Malaya, 78, 2 , near ‘‘ obturbans 
and panalectros.” 

App. Locs —Sylhet, March to June [Lt.-Col. Hall]; Wushai 
Hills, Assam, August [Macleod]; Calcutta, May, and August to 
December [all Indian Museum f¢. Theobald |. 

Type in British Museum. 

N.B.—In vol. v Theobald says that he previously erro- 
neously placed this species as a variety of obturbans : in that 
volume he considers it a good species. 

He previously stated, ‘‘ all variations in colour between the 
true obturbans of Walker, and the fusca of Theobald, seen in these 
specimens, and hence the latter species is sunk as a variety.”’ 


B. joloensis, Ludlow. 
Mr. Theobald (Monog. Culic., iv, 165) admits this as a good 
species. 
Type in Army Medical Museum, Washington. 
B. jugraensis, Leices., 1908. 
Culic. Malaya, 77, 7 ¢. 


‘‘ Larvae in bamboo in Ampang jungle, and water collected 
in a fallen leaf in jungle at Jugra; also on the East Coast and 


442 Records of the Indian Museum. [VoEatV, 


elsewhere. I have received specimens from Borneo ’’ (Leicester). 
Near obturbans, Wik. 
B. obturbans, WIk. 


Evvratum.—The reference to Walker’s Culex ventralis (synony- 
mous with obturbans) should be Jour. Linn. Soc., iv, 91 (1860). 

App. REF.—Blanch., Moust., 266,7 ?. 

Leices., Culic. Malaya, 75,07 2 (with notes on 
varietal forms). 

Locs.—Naini Tal [Gzles] ; Sylhet | Hal/] ; Lushai Hills, Assam ; 
Madras [Cornwall|; Travancore [James]; Behar, Bengal [Green| ; 
Mozufferpur, India; Selangor, 28-x-99 | Butler]; Singapore, 25- 
viii-99 ; Perak [Wray]; Madulsima, Ceylon, 26-ix-07 [Green] ; Sema- 
rang, Java, 1, il,-iii, 1904 and viii-o5 [Jacobson]; Waria Riv., 
Brit., Papua [Dr. Fleming Jones]; Amboina; Celebes; Mysol ; 
Waigiou ; North Ceram ; Tinghai, Formosa; West lake, Hankow, 
China, 28-viii-o7 [Cornford| ; Foochow [ Rennie]. 

Theobald (Rec. Ind. Mus., iv, 4) gives the following data for 
this species and “‘ varieties ” from specimens in the Indian Museum. 

Sukna, I-viii-o8, in deep jungle, Kurseong, 5-vii-o8 [both 
Annandale]; Calcutta, i, iii, vil, x, xii, Rajmahal, Bengal, 31-vii-07 
| Hodgart] ; Trivandrum, 14-xi-08 [Annandale]. 

Taken by me in Calcutta, I11—25-iii-o8 ; 12-iv-08 ; 8-viii-07 ; 
all in bedrooms ; Meerut, 25-iv-05 ; Batavia, 27-vi-06 to 9-vii-06. 

Type in the British Museum. 

N.B.—Mr. Theobald mentions this species being bred in a 
tumbler of water in the Indian Museum by Mr. Tipper of the 
Geological Survey of India. Miss Ludlow has recorded it from 
the Philippines. It breeds freely in the flowers of Heliconia brasi- 
hiensis. ‘‘ Bred from large larvae from under overhanging rock, 
in a deep pool of a clear running stream.’’ 


B. panalectros, Giles. 


(Armigeres panalectoros, Giles, in Theob., Monog., ii, 317.) 

App. REF.—Blanch., Moust., 266, 7 (pamnalectros). 

App. Loc.—Semarang, Java, vill-05. 

N.B.—Theobald (Rec. Ind. Mus., iv, 5) notes that the co- 
type ‘‘ is nothing but an immature, large Culex fatigans, Wied., 
with distinct abdominal banding.’’ 


BREVIRHYNCHUS. Theob., 1908. 
Rec. Ind. Mus., ii, 293. Monog. Culic., v, 144. 


B. annulipalpis, Theob., rgio. 
Rec. Ind. Mus., iv, 6,2. Monog. Culic., v, 148. 


App. Loc.—Maddathorai, 16-xi-o8 [Annandale]. 
Described from a single perfect @ in the Indian Museum. 
Theobald erroneously quotes 1903 as date of capture. 


19i2.] EE. BRUNETTI: Catalogue of Oriental Culicidae. 443 


B. apicalis, Theob., rgro. 
Rec. Ind. Mus., iv, 7, 2. Monog. Culic., v, 149. 


Described from a single @ in the Indian Museum collection 
from Sylhet, 26-vii-05 [ Lt.-Col. Hall). 


B. magnus, Theob., 1908. 


Rec.-Ind.. Mus., 1i, 293, 7 9: iv, pl.i, wing, pl. iii, wing 
scales. | 
Theob., Monog. Culic., v, 145, 7 @, fig. 51 wing, 52 hea , 
side view of abdominal segments. 
Types (one ~ and one @ only) in Indian Museum collection. 
‘* A most marked and beautiful species ; easily told by the quaint 
proboscis and abdominal markings ’’ (Theobald). 
Locs.—Sylhet, May [Z¢.-Col. Hall]; Sukna, 1-vi1-08, in thick 
jungle [Annandale]; Maddathorai, S. India, 17-xi-08 [ Annandale. | 


QUASISTEGOMYIA, Theob., 1906. 
2nd Rep. Gordon Coll. Well. Labs., p. 69. 
Theob., Monog. Culic., iv, 165. 


Q. gardneri, Ludlow. 


Removed here from Stegomyta by Theobald (Monog. Culic., 
iv, 170). 


App. Locs.—Pampanga (Phil Is.) [Whitmore]: Mindoro, 
Bulacao, Phil. Is. 


N.B.—Miss Ludlow (Mosq. Phil. Is., 10) put this species in 
Pseudostegomyia, admitting subsequently to Theobald that it was 
a purely clerical error, intending it for Quasistegomyia. 

Type in Army Medical Museum, Washington. 


KINGIA, Theob., roro. 
Monog. Culic., v, 135. 
K. annandalei, Theob., rgro. 
Rec. Ind. Mus., iv, 10 (Stegomyia id.). 


One 2 from Sukna (500 {t.), vii-o8 [Annandale]. 
Type in Indian Museum. 


STEGOMYIA, Theob. 


Theob., Monog. Culic., iv, 170 (list known spp. I9),—I71 
(table of spp.). : 


444 Records of the Indian Museum. [VOL TVs 


App. Rrer.—Blanch., Moust., 247 (generic characters), fig. 194 
(larva), p. 248 (table of species). 
Leices., Culic. Malaya, 81, with table of Malayan 
species. 
N.B.—Col. Alcock includes Brevirhynchus, Theob., and Har- 
pagomyia, Theob., in his ‘‘ genera of the Stegomyia type.” 


S. albipes, Theob., Igro. 


RecsindyMuseAve Eh. oe 
Monog. Culic., v, 169, @. 
Described from a perfect @ in the Indian Museum taken by 
Dr. Annandale at Maddathorai, 17-xi-08. 


S. albolateralis, Theob. 


Rec. Ind. Mus., ii, 289, 2; iv, pl. i, wing, pl. iii, wing scales. 

Theob., Monog. Culic., v, 179, fig. 67, wing. 

Loc.—Sylhet, September [Hal/|; Lushai Hills, Assam, in 
July. 

Both in the Indian Museum, from five 2 ? in which collec- 
tion the description was drawn up. 

Type in Indian Museum. 


S. amesii, Ludlow. 


App. ReFr.—Theob., Monog. Culic., iv, 191 (Miss Ludlow’s 
description copied). 

App. Locs.—Oras, Samar, Tacloban, Leyte, Twin Peaks, 
Banquet Luzon [all t. Theobald]. 

N.B.—Type in Army Medical Museum, Washington. 


S. annandalei, Theob., 1gto. 
Rec. inds Mus: av. 105-2: 


Described froma single perfect @ taken by Dr. Annandale 
at Sukna, vii-o8. In Indian Museum. Near minutissvma. 


S. annulirostris, Theob. 


App. Re¥F.—Theob., Monog. Culic., iv, 173. 
Type in British Museum. From Peradeniya, Ceylon. 


S. argenteomaculata, Theob., 1907. 
Monog, Culic., iv, 184, 2 


Described from two 2? @ (not in good condition). An easily 
recognized species. 


1912.) EE. BRUNETTI: Catalogue of Oriental Culicidae. 445 


Loc.—Narcondam Isles (Bay of Bengal), 80 miles from Anda- 
mans and Nicobars ; taken by G. Rogers. 
Type in the British Museum. 


S. assamensis, Theob., 1908. 


Rees Inds Mus: 11,290, 9 . 


Monog. Culic., v, 174, 2. 

Theobald says ‘‘ described from a single @,’’ in the Indian 
Museum collection, but gives two localities, Sylhet [Hal/] and 
Pallode, South India. The former is dated 13-iv-05, the other 
one 15-x1-08 [Annandale]. 

Type in Indian Museum. 


S. aurostriata, Banks, 1900. 
Phil. Jour. Sci., i, 995. 


No sex is given, either by Banks or Theobald, who (Monog., 
v, 181) recopies the description with the note that it is clearly a 
distinct species. 

Loc.—Negros Occidental, Phil. Is., Mt. Siya Siya, Canlaon 
Volcano (760 metres), 24-vi-06. 

Type in the entomological collection, Bureau of Science 
Manila. 


S. brevipalpis, Giles. 


App. REF.—Blanch., Moust., 264, 7 9. 


N.B.—This author says the species is ‘“‘ Like a Simulium ;”’ 
which does not agree with Theobald’s expression ‘‘a_ typical 
Culex.’’ 


S. crassipes, V. Wulp. 
App. ReFr.—Blanch., Moust., 250, ?. 


S. desmotes. Giles, 1904. 
Journ: Trop; Med: vii, 367,49; 


Received by Giles from ‘the Philippines: Theobald places it 
doubtfully here. 
Type in British Museum. 


S. dissimilis, Leices, 1908. 
Culic. Malaya, 91, 7 2. 


Described from a series bred from water in the hollow of a 
tree in Ampang jungle. Distinct from all other Stegomyias by the 
gold-scaled mesonotum in the 7. 


446 Records of the Indian Museum. [ VOE. ahve 


S. fasciata, F. 


App. REF.—Leices., Culic. Malaya, 85,0 9. 

App. Locs.—Ceylon [Green]; Garvet, Java [Prof. Marlett], 
Soekaboemi, Java; Celebes, Siam, Perak, Malay Ports, Papua, 
Batavia. 

It occurs in Calcutta in January, and from May to October (not 
so common as scutellaris) ; Lucknow, November ; Purnea, August 
[Paiva]; Lushai Hills [Macleod]; Puri, 18—19-i-09; Mandalay, 
I2-iii-08 ; Rangoon, 25-ii-08, in house, biting by day ; common on 
board ship, Bay of Bengal, between mouth of Hooghly River and 
Rangoon, 22—23-ii-08. I have taken it in Calcutta, i, vi, vii, 
viii, ix, in bedrooms and other places and in the hotel at Lucknow, 
7-vili-05 ; Madras Town, 31-x-08 [Hodgart]; on board ship off 
Coconada, 15-iv-08 [Paiva], and Bhim Tal (4,500 ft.) in September. 

Miss Ludlow recordsit under the name calopus, M. (Mosq. Phil. 
Is., 33), from a very long series of localities in the Philippines 
where it occurs all the year round. 

N.6.—Although Blanchard and Coquillett assume the syno- 
nymy of calopus, Mg., Mr. Theobald doubts its identity with 
fasciata, F. (Monog., iv, 177). 

Owing to Villiers in 1789 adopting the specific name fascrata 
for a Culex, Mr. Theobald fears a change in the name of this well- 
known species may be necessary. Meigen described a fasctata in 
1805, which Theobald adds as a synonym 

Meigen’s calopus (1818) is next on the list, but its identity 
with fasciata, F., appears uncertain. The next name identified 
with the species is frater, Rob. Desv., and Theobald thinks that 
this is the name that may have to be adopted. 

However, as Villiers’s description is unintelligible, and the 
type has long ago ceased to exist, he proposes to abolish Villier’s 
species and retain the name fasciata, F., for this species. This,-as 
he says, will save endless confusion. 

Anyway if Villiers’s description is useless and his type des- 
troyed, there is no reason to assume the species was not a true 
Culex, in which case the question of synonymy drops. Moreover, 
the Kertesz catalogue does not mention this species of Villiers 
at all. 

This species is the sole carrier of yellow fever. 

Mr. Howard says ‘‘ we may expect to find this species every- 
where in the moist tropical zone, or at all events, when intro- 
duced at any point within the low moist tropics it may be 
expected to establish itself.”’ 

In Malaysia the species seems to be confined to the ports. 
Leicester notes that the larvae are found in bathroom tubs in 
houses at Klang, Singapore, Penang, Pangkor-Haut and other 
places. He notes the dense scaling on the clypeus, which, he says, 
no previous writer has noted, and also mentions the variety 
luctensis as occurring in Malaysia. 

Should the identity of fasctata, F., with ‘calopus, Mg., be 


1912. | E. BRUNETTI: Catalogue of Oriental Culictdae. 447 


proved, the species also occurs in South Europe, North, West 
and East Africa, Madagascar, Palestine, Tahiti and New Caledonia. 
Theobald records the typical form from Khartoum, the Nile, 
Greece and Cyprus. 


S. fusca, Leices., 1908. 
Culic. Malaya, 92,7 2. 


‘¢ A small species bred from larvae from water in leaves of 
an atap palm in mangrove swamp at Port Swettenham. Adult a 
blood sucker, and common in jungle where atap palms occur ’ 
(Leicester). 


S. gardneri, Ludlow. 


Removed to Quasistegomyia by Theobald (Monog., iv, 170). 


S. gracilis, Leices., 1908. 
Culic. Malaya, 81, 7. 


Larva found in water in bamboos. Adults numerous in 
bamboo jungle. 


S. imitator, Leices., 1908. 
Culic. Malaya, 89, 2. 
“Described from 2 2 @ from jungle 5 miles from Kuala 
Lumpur,’ which is apparently its only locality. 
S. leucomeres, Giles, 1904. 
Journ. Trop. Med., vii, 367, @. 


Loc.—Phil. Is. Type in British Museum. 

Banks records it from Pampanga, but Theobald says the 
species is an uncertain one, the type being in bad condition, but 
probably a Stegomyia. 


S. mediopunctata, Theob. 


App. REF.—Theob., Monog. Culic., iv, 187, @. 
Type in British Museum. 


S. microptera, Giles. 


App. RreF.—Blanch., Moust., 263. 

This author adds ‘‘ N. W. Prov. India’’; in houses during 
the rains ; at one time referred by Giles to Wyeomyia. 

Theobald (Monog., v, 607, Appendix) says, ‘‘ Type appears 
to be lost.’’ 


448 Records of the Indian Museum. hVOL AY. 


S. minutissima, Theob., 1910. 
Rec. Ind. Mus., iv, 9, 2; v, 168, 2, fig. 61, wing. 


N.B.—Though the o sign prefaces the description of the 
species, apparently only the @ is known. 
Type in Indian Museum. 


S. nivea, Ludlow. 


Referred to Scutomyia. 


S. periskelata, Giles. 


App. RreF.—Blanch., Moust., 264. 


N.B.—Theobald says (Monog., v, 155) that he does not under- 
stand this species, which is placed here provisionally. The type 
is notin the British Museum. Further on (loc. cit., 607, App.) he 
suggests that the name of the species should be dropped. 


S. perplexa, Leices., 1908. 
Culic. Malaya, 83, 7 @. 


Described from one ~ and several @ @ from jungle near 
Kuala Lumpur in May, October and November. The autho: 
seems uncertain of its true position in this genus as it has affinities 
with Scutomyia, and he suggests it may be a hybrid. 


S. pipersalata, Giles. 


App. Rer.—Blanch., Moust., 264,07 2. 
Theobald (Monog., v, 607, App.) is doubtful if a 
Stegomyia. 


Type in British Museum. 


S. pseudonivea, Theob. 


App. ReF.—Theob., Monog. Culic., iv, 188, 2; v, 176, 0, fig. 


64, head. 
Type o in British Museum, type @ in Hungarian Museum 


Four of each sex were taken by Lowis in the Andamans. 
S. punctolateralis, Theob. 
Type in British Museum. 


S. scutellaris, W1k. 


App. RreFr.—Blanch., Moust., 257,c. 
Leices., Culic. Malaya, 86,7 2. 


I9I2.] E. BRUNETTI: Catalogue of Oriental Cultcidae. 449 


App. Loc.—Sylhet, April [Had/]; Lushai Hills, May to July 
[Macleod]; Manipur, July [Gourlay|; Calcutta, March, July to 
October, common during the hot weather in the rains, disappear 
ing in winter, active by day; Katihar, N. Bengal, October 
[Paiva]; Purnea, viii, ix [Paiva]; near Puri, x-0o8; Lucknow, 
4-iv-05 [Brunetti] ; Sukna, I—2-vii-08 ; Mandalay, ii, iii-o8 ; Bhim 
Tal, breeds in hollow trees in jungle, ix-o6; Trivandrum, 14-xi-08 ; 
Maddathorai, 18-x-08 [all Annandale]; Madras Town, 30-x-08 
[Hodgart]; Shahjahanpur [Giles]; Victoria gardens, Colombo, 
26-iv-08 [Paiva]; Singapore, 21-vi-06 [Brunettt]. 

Sarawak, Papua generally, Upper Burma, Foochow, Hankow, 
21—28-vi, Seychelles, Mauritius, Pitcairn Is., Honolulu. 

All the above specimens identified by Theobald are in the 
Indian Museum. 

I have taken it in Calcutta as late in the year as I0-xi-04. 

Type in the British Museum. 


N.B.—The species breeds freely in the flowers of Heliconia 
brasiliensis. Dr. Barker says that at Sarawak it is abundant in 
the neighbouring thick undergrowth, but that it seldom enters 
houses in the daytime, and not at all at night. Common in Cal- 
cutta in hot weather and rains, disappears in winter. Is the most 
abundant species in Mauritius. 

Dr. Leicester notes in his description of the species, one or 
two points not mentioned by Theobald, adding that the insect 
breeds as freely in bath tubs as in the jungle. 


Sub-species samarensis, Ludlow. 


Theobold retains this (Monog., iv and v) as a variety of 
scutellaris, Wlk., and disputes Banks’s suggestion that possibly 
intergradations may occur between scutellaris and fasciata. 

Type in Army Medical Museum, Washington. 


““S. albopictus,’’ Skuse. 


Definitely accepted as synonymous with scutellarts. 


S. sexlineata, Theob. 


Further corroboration of the identity of the Philippine Island 
specimens received from Banks, with this species, described from 
Trinidad. 


Type in British Museum. 


S. striocrura, Giles, 1904. 


No sex is mentioned, the fype is not in the British Museum, 
and Theobald is doubtful of its specific validity. 


450 Records of the Indian Museum. [Voev ing 


S. thomsoni, Theob. 


App. Rer.—Monog. Culic., iv, 174. 
From N. W. Prov. India. Tye in British Museum. 


S. tripunctata, Theob., 1908. 


Rec. Ind. Mus., ii, 288, 2; iv, pl. i, wing, pl. iii, 
wing scales; v, 182, fig. 68, wing. 


Loc.—Lushai Hills, Assam, 6-vi-04 [| Macleod}. 


N.B.—Described from two ? 92 , very near S. amesiz, Ludlow. 
Type in Indian Museum. 


S. w-alba, Theob. 


App. REF.—Theob., Monog. Culic., iv, 180, 2 , fig. 151 (head, 
thorax, femur). 


PSEUDOSKUSEA, Theob., 1907. 
Monog. Culie "iv. 192,07 o- 
P. multiplex, Theob. 


App. RreF.—Theob., Monog. Culic., iv, 192,0. 
Removed here by Theobald from Skusea. 
Type in Hungarian Museum. 


P. nigrotarsis, Ludlow, 1908. 
Cans Ets ext 52 aor 


Loc.—Infanta, Tayabas (Phil. Is.), October, a unique speci- 
men. 


SKUSEA, Theob. 


App. REF.—Blanch., Moust., 416. ° 

In Monog., iv, 542, Theobald definitely places this genus in 
Aedinae and retains it there in vol. v, but in view of possible 
further alterations in the sequence of genera in this family, I 
prefer to retain the order adopted in my previous catalogue, as 
near as possible, merely for the sake of convenience. 

Leicester (Culic. Malaya, 117) says, ‘‘ This genus was origin- 
ally placed im the sub-family Aedeomyinae by Theobald in vol. iii 
of his Monograph, and my genus Amauromyia!' exactly corres- 
ponds to it, but in the Genera Insectorum, Fasc. 26 (1905), Theo- 
bald has transferred Skusea to the Culicinae, as the 7 @ have 
long palpi. ‘The genus is unrepresented in Malaya.’’ 


1 IT can find no reference to this genus. 


IQI2. | E. BRuNneEtTt1: Catalogue of Oriental Culicidae. 451 


S. culiciformis, Theob. 


App. REF.-—Theob., Monog. Culic., iv, 546, 2 , fig. 251 wing @. 


S. diurna, Theob. 


App. REF.—Theob., Monog. Culic., iv,547, 2. The type was 
bred by Dr. Durham in September. 
Type in British Museum. 


S. funerea, Theob., var. ornata, Theob. 


App. Rer.—Theob., Monog. Culic., iv, 542, fig. 248 wing 2. 
Type in Hungarian Museum. 


S. mediofasciata, Theob., 1907. 


Monog. Culic., iv, 544, 0% 9 , fig. 249, wing 2 , 250, 
@ genitalia, pl. vii, wing scales 2. 
pl. ix, ¢ genitalia. 
Syn. Pseudoskusea mediolineata, Ludlow (¢. Ludl., Can. Ent., 


xl, 332). 
Loc.—-India [Christophers]. Described from I7 3 @ 2. Very 


neat Skusea funerea, Theob. Miss Ludlow says it has been received 
from the Philippines. Theobald does not give mediolineata as 
synonymous with his medzofasciata in vol. v. 

Type in British Museum. 


S. pseudodiurna, Theob., 1g10. 
Rec. Ind. Mus., iv, 32, o. 


Theob., Monog. Culic., v, 491, @. 
A unique specimen ; in the Indian Museum, from Sukna, I-vii- 


08 [Annandale]. Very near S. diurna. 


S. pseudomediofasciata, Theob., 1gIo. 
Monog. Culic., v, 489, 0. 


From Peradeniya and Hakgala, Ceylon, iti and iv, 1907 


[Green]. 
Type in British Museum. 


S. uniformis, Theob., Igfo. 
Rec. Ind. Mus., iv, 33, 2. 


Theob., Monog, Culic., v, 491, 2. 
Type in the Indian Museum ; a unique specimen trom Pallode, 


S. India, 15-xi-08 [Annandale }. 


452 Records of the Indian Muscum. [Vox. IV, 


SCUTOMYIA, Theob. 


App. ReF.—Theobald, Monog. Culic., iv, 196, 197 (short 
description and tabulation of the only known 
five species). 

Leices., Culic. Malaya, 105. 


S. albolineata, Theob. 


App. REF.—Theob., Monog. Culic., iv, 197, 2. 
Leices., Culic. Malaya, 105,07 ?. 
N.B.—Nothing is said by Theobald as to Giles’s species of 
this name (vide my Catalogue, p. 336). 
Type in British Museum. 


S. nivea, Ludlow. 


App. RErF.—Leices., Culic. Malaya, 87,7 2°. 

Leicester described from a series of adults and some bred 
from bamboo water in jungle. ‘‘ Entirely sylvan, although ap- 
pearing in houses close to jungle, fairly common, and a_ vicious 
biter.” 

Type said to be in the Army Medical Museum, Washington, 
but Theobald says (Monog., v, 203) it has been given by Miss 
Ludlow to the British Museum. 


S. notoscripta, Skuse. 
App. REF.—-Theob., Monog. Culic., i, 286, @ , fig. 84 (wing 
2), fig. 85 (wing scales); ili, 145. 
Blanch, Moust., 257,0 ° . 
App. Locs.—Muiria, Seleo, Berlinhafen, Friedrich-Wilhelms- 
hafen (all Papua) and Ins. Graget [all locs. ¢. Biro]. 
“India” is given by Theobald as doubtful, although Giles 
reports it from that country. 
S. notoscripta, sub-species samarensis, Ludlow. 


Also occurs at Kuranda, Queensland, taken by Dr. Bancroft. 


S. sugens, W. 
App. REF.—Blanch., Moust., 256, 2. 


LEICESTERIOMYIA, mihi, nom. noy.! 


Syn. Chaetomyia, VWeices., Culic. Malaya, 100, 1908 (preoccu- 
pied). 


} The present name is suggested merely as a substitute for Chaetomyta, but 
it must be understood that I do not consider it as of generic rank, any more than 
the great majority of the so-called ‘‘ genera ’’ in this family. 


Igi2. | E. BRuNE?Tt1: Catalogue of Oriental Culicidae. 453 


’ 


Near ‘‘ Desvotdya’ 
in bamboo water. 


and Leicesterta ; purely sylvan, and breeds 


N.B.—The generic name Chaetomyia being preoccupied in 
Tachinidae by Brauer and Berganstamm,! I propose Leicesterio- 
myia as a nomen noyum. 

Gi flavas Leices. Fas: 
Culiec. Malaya, 101,07 9. 


Described from a series bred from larvae from bamboo water 
and in cocoanut shells in jungle in Malaysia. Sylvan, a vicious 
day biter. 


DASYMYIA, Leices., 1908. 
Culic. Malaya, 102. 


‘There is no mosquito this species could possibly be mis- 
taken for,’ the author adding that it might be found to belong 
to the Aedeomyina, and notes its affinity to Mimomyia. 

D. fusca, Leices., 1908. 
Culic. Malaya, 102, 7 9. 


Aco in jungle five miles from Kuala Lumpur, and a 9? in 
bungalow at Bukit Kutu. Appears to have affinities with Stego- 
mytia, Scutomyia, Uranotaenia, Etorleptiomyia and Mimomyra. 


CONOPOMYIA, Leices., 1908. 
Culic. Malaya, 113. 

Copious notes on this genus are given by this author, who is 
uncertain where to place it, and as to whether it belongs to the 
Culicinae or Aedeomyinae. I therefore leave it here where he 
temporarily places it. 

C. aurea, Leices., 1908. 
LOG Cis EIG.. oF °9.. 

Described from one @ (taken in a bungalow in Kuala Lum- 
pur) and one @ , in jungle at Raub. 

C. hybrida, Leices., 1908. 
Boer cu., 115, 7 2. 


Described from a series. A @ from a bungalow in Kuala 
Lumpur and other 7 #7 anda @ sent to Leicester by Dr. Finlay- 
son of Singapore. 


1 Denk. Ak. Wien., lviii, 311 (1892). 


ad 


454 Records of the Indian Museum. [Vor. IV, 


C. metallica, Leices., 1908. 
LOC CHAAEET R= opOhs 


Described from a series from larvae from marshy ground near 
Kuala Lumpur and elsewhere. 


PSEUDOCARROLLIA, ‘Theob , 1910. 


Rec. Ind. Mus., iv, 12, near Carrollia, Lutz. 


d 


Theob., Monog. Culic., v, 186. 


P. lophoventralis, Theob., 1gro. 


Rec: Ind. Mus:5 ive-t3,.09 ¢. > Monog, Culie- v7 186..0"- 


Described from a single perfect 2 taken by Mr. Paiva at 
Purnea, Bengal, 6-viii-o7, resting on the under side of a leaf of a 
lichi tree during the day. 

Type in Indian Museum. 


) 


LEICESTERIA, Theob. 


App. REF.—Theob.. Monog. Culic., iv, 201. 
Leices., Culic. Malaya, 94. 


L. annulitarsis, Leices., 1908. 
Culic. Malaya,99, 7 @. 


Apparently sylvan and local in jungle 5 miles from Kuala 
Lumpur ; quite a distinct species, a bamboo breeder ; only one ¢. 


L. apicalis, Theob., 1908. 


Rec. Ind. Mus., i, ZOL GOs ive epliel, wins, 
pl. 111, wing scales. 
Theob., Monog. Culic., v, 213, 7 2, fig. 86, wing. 


Described from one ~ and two 2 2: ‘‘ two hatched from 


larvae and one caught.” Lushai Hills, Assam (1,500 ft.), May 
| Macleod}. 


Types in Indian Museum. 


L. cingulata, Leices., 1908. 
Culic. Malaya, 97, @. 


Described from three @ 2 ‘(April and June), of which one 
was bred from a larva from bamboo in jungle five miles from 
Kuala Lumpur. 

A vicious biter, but apparently very local. 


Un 


1g12. | E. BRUNETTI: Catalogue of Oriental Culictdae. 45 


L. dolichocephala, Leices., 1908, em. mihi. 
Culic. Malaya, 95, 7 ¢ (dolicocephala). 


In jungle round Kuala Lumpur and at Bukit Kutu, probably 
a bamboo breeder, a vicious biter in daytime and at sunset. 


L. longipalpis, Leices., in Theob. 


App. REF.—Theob., Monog. Culic., iv, 201, 7 9°. 
Leices., Culic. Malaya, 94, 7 2. 
Types in British Museum. 


HULECOETOMYIA, heob., 1907. 


App. RreF.—Monog. Culic., iv, 220 (Hulecoeteomyia). 
Leices., Culic. Malaya, 107. 


H. fluviatilis, Leices., 1908. 
Culic. Malaya, ili, a 9. 


Very near jugraensis and frilineata, the larva, however, is 
quite distinct. Probably rare, as it has only been found in one 
locality, Ulu Gombak jungle, 13 miles from Kuala Lumpur. 


H. jugraensis, Leices., 1908. 
Culic. Malaya, 109, 7 2 [Helecoeicomyia td., lapsus). 


Described from a series bred from water in fallen leaves in 
jungle at Jugra. Very near H. trilineata, Leices., but the larvae are 
quite different. J,eicester adds: *‘ the name ¢rilineata is an unfor- 
tunate one, as there are really seven distinct lines on H. trilineata 
whereas in H. jugraensis there are only three.’’ 


H. pseudotaeniata, Giles. 


App. Rre¥F.—Blanch., Moust., 261, °@. 
Theob., Monog. Culic., v, 222, describes o@ for 
first time, fig. 89, wing, 90, head o@, gr, 

genitalia o. 

Ieicester says it will breed in small collections of rain water 
near houses. Theobald (Rec. Ind. Mus., ii, 291) mentions 3 7 @ 
and2 2? 2 ‘‘hatched 29-vili-o4’’ from Tungleh (? ) in the Lushai 
Hills. 

Also recorded from Dehra Dun by Thomson. 

Banks records it from the Philippines. 


H. trilineata, Leices., in Theob. 


App. REF.—Theob., Monog. Culic., iv, 220, 7 °@. 
Leices., Culic. Malaya, 107, 7 @. 


456 Records of the Indian Museum. [Vo.. IV, 


App. Loc.—Kurseong, 3-vii-o8 [Annandale]; Bukit Kutu 
(May), Bamboo jungle 5 miles from Kuala Lumpur, April, May, 
July, October, December; Ulu Gombak, 13 miles from Kuala 
Lumpur (both ¢. Lezcester). 

The slightly damaged types came from the former jungle. 

Leicester says it is a very distinct species, a bamboo breeder, 
not common, quite sylvan and a blood sucker. 


PHAGOMYIA, Theob. 
P. gubernatoris, Giles. 


App. RreF.—Blanch., Moust., 261, 2 (Stegomyta). 


HOWARDINA. Theob. 
App. ReF.—Blanch., Moust., 415. 


H. chrysolineata, Theob., 1907. 
Monog. Culic., iv, 218, 2, pl. i, wing scales, °. 


A unique ? from Peradeniya, Ceylon [Green]. 
Type in British Museum. 


H. greeni, Theob. 


App. Rre¥F.—Blanch., Moust., 416. 
Type in British Museum. 


H. himalayana, Giles, 1904. 
Journ. Trop. Med. (1904), 384, ¢@. 


Loc.—Naini Tal {in August}, bred. Giles puts it doubtfully 
here and Theobald states that Mr. Carter suggests, after examining 
the type, that it would be better placed in Pseudohowardina. 


PSEUDOHOWARDINA, Theob., 1907. 


Monog. Culic., iv, 223, 2. 


P. chrysoscuta, Theob., 1907. 
Op. ctt., v, 228, 2 , fig. 94, wing. 


A unique specimen from Peradeniya, iv-07 [Green]. 
Type in British Museum. 


CULICIOMYIA, Theob., 1907. 


Monog. Culic., iv, 227, 7 92, pl. ili, wing scales °. 


LOuz: | KE. BRUNETTI: Catalogue of Oriental Culicidae. 457 


C. annulata, Theob., 1907. 


Op. cit., 230,07 &, fig.64, wing @ , 65, genitalia; pl. i, wing 
scales ©. 


Loc.—Kuching, Sarawak [Dr. barker|; taken in company 
with C. inosnata, Theob. ‘‘ Looks like a small Culex piptens.’’ 
Type in British Museum. 


C. annuloabdominalis, Theob., rgro. 


Monog. Culic., v, 236, 7 @, fig. 102, head occiput, 103, head. 
Loc. —Peradeniya and Hakgala, Ceylon, i and v-07 [Green]. 
Tvpe in British Museum. 


C. ceylonica, Theob., 1907. 
Op. ctt., 236, 7 @, fig. 70, wing, ¢. 


Described from a perfect 7 and 2 from Peradeniya and 
Maskeliya, Ceylon (February and April) [Greev]. 
Type in British Museum. 


C. inornata, Theob., 1907. 
% 


Loc. cit., 227, 7 2, fig. 61, head scales, 62, wing ? , 63, 7 


genitals. 


1oc.—Kuching, Sarawak | Dy. Barker|, in a house, November. 
Type in British Museum. 


N.B.—Miss Ludlow records it from the Philippines (Can. 
Bit. -xli, Oi): 


C. minutissima, Theob., 1907. 
EGCACi.; 235.02" 


Lovc.—Peradeniya, Ceylon, February [Green]. 
Tvpe in British Museum. 


. . s - 
C. nigerrima, Theob., 1gto. 
Monog. Culic., v, 233, @ , fig. 100, wing. 


A pertect unique specimen from Trincomalee, Ceylon, taken 
October 1907 by Green. 
Type in British Museum. 


C. ‘pulla, Theob. 


App. REr.—Theob., Monog. Culic., iv, 232, fig. 66, head 
scales. 

Removed here from Culex. 

Type in British Museum. 


458 Records of the Indian Museum. [VoL aie 


NEOMACLEAYA, Theob., 1907. 
Monog. Culic., iv, 238, °. 


N. indica, Theob., 1907. 
Loe "cits, 238) 42% 


Loc.—India [Christophers]. ‘* At first sight resembles Skusea 
funerea Theob.’’ Philippines [¢. Ludlow]. Woodlands, Cevlon, 
Q-x-07. 

Type in British Museum. 


Var. simplex, Theob. 


Reéc: Ind’ Mus. 2017 "9": 
Loc.—Sylhet, June [Hall]. A single @. Type in Indian 
Museum. 


DANIELSIA, Theob. 


App. REF.—-Theob., Monog. Culic., iv, 240. 
Leices., Culic. Malaya, 117. 
N.b.—It may be noted that there is a Daniela, Koch, 1891, 
in Coelenterata. 


D. albotaeniata, Leices., in Theob. 


App. REF.—Theob.,: Monog. Culic:, iv, 241, + 9. fg. 72 
(cephalic and scutellar adornment). 
Weices, (Culics Malaya 117,507 12 


Leicester describes it in the above work from a series bred 
from larvae from bamboo water. Sylvan, a vicious day biter. 


LEPIDOTOMYIA, Theob., 1905 


REFs.—Gen. Insect., Fasc., 26, p. 22. 

Theob., Monog. Culic., iv, 249. 

Leices., Culic. Malaya, 132. 

Ervratum.—Delete line 3, p. 339 of my Catalogue. 


N.B.—‘* An error has occurred here. The Lefidotomyza ot 
Theobald, in his paper on the Hungarian Museum Culicidae, re- 
ferred to the genus Reedomyia, Ludlow, to which the species 
Lepidotomyia alboscutellata Theob., belongs. The true Lepidotomyza 
contains only one species, magna Theob., and comes very near 
Danielsta’’ (Theobald). 

Leicester, however, in his Culicidae of Malaya (p. 132) retains 
alboscutellata in Lepidotomyia without comment. 

Type in British Museum. 


1912.] E. BRuNET?II: Catalogue of Ortental Culicidae. 459 


L. magna, Theob. 


App. REF.—Theob., Monog. Culic., iv, 250, 7 @?. 

Described from two perfect specimens ; a true Lepidotomyza. 
Recorded from Bombay, I9-viii-o2, by James. Tye in British 
Museum. 


L. taeniata, Leices., 1908. 
Culic: Malaya, 133, 07:2. 


Described from a large series bred from larvae taken in water 
collected in ruts in a jungle waggon track. ‘The only time this 
author has seen the larvae. 


THEOBALDIA, Nev. Lemaire. 


Theobaldinella, Blanch., 1905, Moust , 390, nom. nov. (gen. 
chars. given). 

N.B.—Blanchard changed the name from Theobaldia to Theo- 
baldinella on account of Theobaldius of Neville in Mollusca ; but 
Mr. Theobald retains the spelling as originally written. which in 
accordance with zoological ruJes is permissible. 


T. annulata, Schrk. 
Beitr. zur. Naturgesch.. 97 (1776). 


App. REF.—Culex annulatus, Blanch., Moust.. 280. 7 @, 
fig. 206, ungues and genitalia, fig. 207, adult larva. 

Apparently a hill species. lanchard records it from 4,000 ft. 

(Brianon, France), also from 8,000 feet in Mexico (Durango 
State). 


T. spathipalpis, Rond. 


App. Rrer.—Theob., Monog. Culic , iv, 276 (larva described). 
Blanch., Moust., 283, 7 @ , fig. 209. ungues and 
genitals. 


N.6.—-Blanchard says that Ficalbi thinks the adult does not 
bite, but lives on plant juices. The larva has been found during 
winter { ? in Sardinia). Blanchard thinks it may carry ‘‘ undulat- 
ing’’ fever in Malta, adding that, at least in Gibraltar, where it 
abounds, it is infested by a microbe closely resembling Micrococcus 
melitensis. 


PECOMYIA, Theob. 


Geitonomyia, Leices., 1908, Culic. Malaya, 134. 


P. caeca, Theob. (Culex td.). 


App. Rer.—Blanch., Moust., 305, 2 (Culex id.), 
Leices., Culic. Malaya, 135, 7 9 (Geitonomyia id.) 
App. Locs.—-Fed. Malay States (Ipoh-Parak}, Philippines 


460 Records of the Indian Museum. [ VorE.-Dvs 


N.B.—Re-described by Leicester from a series bred from 
larvae in water in pools and waggon tracks at Ulu Klang near 
Kuala Lumpur. 

‘** As the scutellar scaling and the palpi are quite distinct from 
those of a Culex, I have no hesitation in placing this mosquito in 
a separate genus ” (Leicester) However, Leicester had probably 
overlooked Theobald’s genus Pecomyza, to which the latter author 
referred this species (Monog. Culic., iv, 268, fig. 86, wing 2. pl. i, 
wing scales @ }). 

Tvpe in British Museum. 


P. maculata, Theob. 


App. RrEF.—Theob., Monog. Culic., iv, 266, 7 2; figs. 82, 
83, 84, 2 wings. 
Type in British Museum. 


PARDOMYIA, Theob., 1907. 


Monog. Culic., iv, 280. 


P. aurantia, Theob., 1907. 
DL OCNCUiaS 2005 Or 


Loc. —Kuching (Sarawak), November [Dr. Barker]. Received 
also from Kuala Lumpur. TJ ve in British Museum. 


P. quadripunctis, Ludlow. 


No reference to the description of this species is given, of 
which Theobald (Monog., v, 608) copies the original description 
It is from Parang, Mindanao (Phil. Is.}, Oct. 26. 


PSEUDOGRAHAMIA, Theob. 
P. aureoventer, Theob., 1910. 
Rec dndyMus iv 270 oS 
Monog. Culic., v, 551, 2. 


A unique ¢ , at present in the British Museum. 
Pallode (Travancore. S. India), 16-xi-08 [Annandale |. 


PSEUDOGRABHAMIA, Theob. 
P. maculata, Theob. 


App. REF.—Theob:, Monog. Culic:, iv, 314, @ 2%, fig. 10g 
wing @. 
Id., Journ..<Bom. Nat: Bist..90¢:, xvi, 2447075 2 


Igi2.] E. BRUNETTI: Catalogue of Oriental Culicidae. 461 


App. Loc.—Madras Town, 30-x-o08 [Hodgart, Ind. Mus.]. 
Type in British Museum. 


GRABHAMIA, Theob. 


App. RE¥F.—Theob., Monog. Culic., iv, 284 (list of known 
sp.), 285 (table of leg bands). 
Blanen..)Moust.,. 396, oc 9. 


N.B.—Feltidia, a genus by Dr. Dyar, for some American 
species, is an absolute synonym of Grabhamia, being founded on 
jamaciensis, the very species which is the type of Grabhamia. 


G. ambigua, Theob. 


Type in British Museum. 


G. ochracea, Theob. 


App. Rt¥r.—Theob., Monog. Culic., iv, 300. 
Described from two perfect 2? @. 
Type in British Museum. 


G. spenceri, Theob. 


7 ype in British Museum. 


G. sollicitans, WIk. 


App. RreF.—Theob., Monog. Culic., iv, 291 (life-history notes), 
figs. 97-98 (larval characters}. 


N.B.—As in that work Mr. Theobald makes no further men- 
tion of the Formosan example, on which the species was intro- 
duced into my Catalogue, it should be, pro fem., eliminated from 
the list of Oriental species. 

G. taeniarostris, Theob., 1907. 
Monog. Culic., iv, 299, 2. 

Loc.—Peradeniya, January [Green |. 

Type in British Museum. 

CULICADA, Felt., 1904. 
Mosq. of New York State, App. p. 39Ib. 


Theob., Journ. Econ, Biol. (1905), i, 26 
Id., Monog. Culic., iv, 318. 


462 Records of the Indian Museum. [ VOEAEV, 


C. eruthrosops, Theob., 1910. 
Monog. Culic., v, 299, 2 , fig. 140, wing. 


Trincomalee, Ceylon, xi, 1900. 
7 vpe in British Museum. 


C. minuta, Theob., 1907. 
Monog. Culic., iv, 338, 2. 


Loc.—India [Christophers|. Described from a perfect ?. 
Type in British Museum. 


C. suknaensis, Theob., 1910. 
RecInd. Mus., ‘vi;21, 12). 


Monog. Culic., v, 297, @ , fig. 139, wing. 

Described from four perfect 2 @ from Sukna, I—2-vii-o08 
[Annandale|. Near C. ntpponit, Theob. The specimens were 
taken in dense jungle. and bit during the day. 

Type in Indian Museum. 


THEOBALDIOMYIA, mihi, nom. nov. 


Syn. Leucomyta, Theob., 1907, Monog. Culic., iv, 372, pl. ix, 
y genitalia. 
Type of genus (=sub-genus ¢t. miht), Culex gelidus, Theob. 
N.B.—Leucomyia is preoccupied in 1892 by Brauer and 
Bergenstamm in Sarcophaginae (Denk. Ak. Wien., lviii, 368). 
I therefore propose the name Theobaldiomyia, with the view that 
the group represents, at most, a sub-genus 


T. argentea, Ludlow. 


Taentorhynchus argenteus, Ludlow. 
Td. rd., Theob., Monog. Culic., iv, 487, 9 
‘copies Miss Ludlow’s descr.}; v, 426, fig. Ig1, wing. 
Described from several taken by Dr. Whitmore in September 
at Angeles (Pampanga, Phil. Is.). 
Type in Army Medical Museum, Washington. 


N.B.—In vol. v ‘Theobald says that Miss Ludlow informs 
him that it is a Leucomyra. 


T. gelidus, Theob. (Leucomyia). 


App. Rrer.—Leices., Culic. Malaya, 147, 7 @ |Leucomyia]. 

App. Locs.—Calcutta, August; Calcutta, 6-x-o04 [ Brunett7] ; 
Rajshahi, E. Bengal, 1—6-ii-o7; Rangoon, 25-ii-08 ; Travancore 
and Cochin States, xi-o8 [Annandale]; Purnea, ix-x [Paiva]; 


1912. | E. Brunetti: Catalogue of Oriental Culicidae. 463 


Madras Town, 30—31-x-08 [Hodgart|; Maskeliya, Ceylon (April) 
[Green]. 

N.B.—Leicester re-describes the @ from a long series bred 
from pools and taken in bungalows at Kuala Lumpur. He notes 
that Theobald classes this species with those with an unbanded 
proboscis instead of with the banded ones. He says the o has 
not before been described, but Theobald did so at the time of 
establishing Leucomyia. 

Type in British Museum. 


T. gelidus, var. bipunctata, Theob., 1907. 
Monog. Culic., iv, 374, 7 (Leucomyta id. 1d.}. 
Loc.—India [Mai. Aldrich]; Sarawak [Dr. Barker}. 


T. gelidus, var. cuneata, Theob. 


Evratum.-—My reference to the original description of this form 
is incorrect, as it should be, Theob., 1901, Monog. Culic., ii, 22. 

App. Rer.—Blanch., Moust., 316, @. 

App. Locs.—Balighai, near Puri, 23-x-08, at light; Calcutta, 
Aug. to Dec., ‘‘ not uncommon in houses and at light, and in the 
open, on shrubs and in railway carriages; Travancore State, 5 
and 19-xi-08 [all A nnandale| ; Calcutta, 6-x-04 [ Brunetti|; Katihar, 
Bhogaon, ix, x; Purnea, viii, x [Paiva]; Sylhet (May) [Hal/]; and 
between Bolpore and Rampore Haut, Bengal, in August in railway 
carriage [Paiva]; western base of W. Ghats, Travancore, 19-xi-08 
[Annandale|; Malabar, 4-xi-o8 [Annandale]; Madras, 30-x-08 ; 
Pangasinan (Ph. Is.). 


T. sinensis, Theob. 


(L gelhida, var. sinensis, Theob.). 

Theobald (Rec Ind. Mus., iv, 20) raises this variety to 
specific rank, and adds the locality Balighai, near Puri, 23-x-08, 
at light [ Annandale}. 


T. (2?) whitmorei, Giles. 


See Taeniorhynchus, td. 


LOPHOCERATOMYIA. Theob. 
App. Rer.—Theob., Monog. Culic., iv, 471 (generic notes). 
Leices., Culic. Malaya, 119. 
L. bicornuta, Theob., gto. 
Rec. Ind. Mus... iv, 25. 


Theob., Monog. Culic., v, 412, o, fig. 178 antennal organs, 
179 wing, 180 genitalia, 181 head, 182 antennal organs in profile. 


464 Records of the Indian Museum. [ VOLSGEV, 
Type in Indian Museum. A single (dissected) male from 
Kawkareik, base of Dawna Hills, 4-i1i-08 [Annandale]. 


Closely related to fraudatrix, Theob. 


L. brevipalpis, Theob. 


Monog. Culic., iv, 477, @, fig. 12 @ proboscis, palpus, basal 


antennal segment, antennal organs, ungues. 
App. REF.—Leices. Culic. Malaya, 129 (copies Theobald’s des- 


cription, as h> has not seen the species in Malaysia). 


L. eminentia, Leices., 1908. 
Culic. Malaya, 131, o. 
Described from a single ~ from jungle near Kuala Lumpur. 
‘Very distinct and easily recognised.’ 


L. fraudatrix, Theob. 


or fig. 208 


App. REF.—Theob., Monog. Culic., iv, 474, oN 
palpi, 209 wing @ , 210 antennal organs o~, 211 wings o, 


Type in Hungarian Museum. 


L. mammilifer, Leices., 1908. 
Culic. Malaya, 128, o& @. 


Described from a series of o” o™ and one ? bred from larvae 


from pools in jungle near Kuala Lumpur and at Raub. 
A somewhat distinct species, according to the author. 


L. minor, Leices., 1908. 
OG Cia T2ZONI HN? 
Described froma series bred from bamboo water. Sylvan, 


and the smallest of the Ma'ayan species, and dull coloured. 


I. niger, eices.,. 1906, 
LOC ICU AZ 2 as 
Described from a series from larvae from ponds at Kuala 
Lumpur. 
L. rubithoracis, Leices., 1908. 
LOcecti 5 STEOR ae: 12%, 


Described from a series from ponds in Kuala Iumpur. 
Easily known by its brilliant red thorax, 


1912. | E. Brunetti: Catalogue of Oriental Cultcidae. 465 


L. sylvestris, Leices., 1908. 
POCNClE 25. (Qe. 


Bred from larvae from still ponds in Malayan jungle. 


L. taeniata, Leices., 1908. 
TOG: CU X27 Agen: 


._ Bred from larvae from ponds in the open near Kuala Lumpur 
and Klang. A very distinct species. 


L. uniformis, Theob. 


App. REeF.—Theob., Monog. Culic., iv, 473, @ 9. 
Id., Journ. Bomb. Nat. Hist. Soc., xvi, 245. 
Type in British Museum. 


L. variata, Leices., 1908. 
Culic)Malaya.a2r. 0 9. 


Described from a large series bred from larvae mostly found 
in small ponds and puddles in swamps and ponds. 


RADIOCULEX, Theob., 1908. 


Rec. Ind. Mus., ii, 295. 
Theob., Monog. Culic., v, 192. 


R. clavipalpus, Theob., 1908. 


Rec. Ind. Mus., ii, 295, 7 2, iv, pl. i, wing, pl. 11, head. 

iheob:, -Monoge. Culic:, v, 193, 'o 2. 

Loc.—Calcutta, Nov. and Dec. ; and onein July ; Berhampur 
(Murshidabad Distr.}, Bengal, 1-i-o8 jl/oyd|; Katihar, 4—5-x-08 
[Paiva]; Rangoon, 25-ii-o8 [A nnandale|; Vaikam Coast of Travan- 
core State, 5-xi-08 [ Annandale}. Common inbrushwood in the cold 
weather in Calcutta. 


N.B.—‘‘ Described from a long series, some taken at light, 

. the marked black shiny thorax with the clear-cut yellow 

area on it, and the quaint marginal cell will at once identify it” 
(Theob.). 


Tvpe in Indian Museum, co-types in British Museum, 


CULEX, L. 


For sub-division of even the restricted genus Culex, wide 
Theobald (Monog. Culic., iv, 387). Blanchard divides the genus 
into nine sections (Moust., 269) and arranges the groups of species 
in further analytical tables, one for each section, incorporating 


466 Records of the Indian Museum. [Vor. IV, 


the specific descriptions in the tables. - Theobald (vol. v) admits 
nearly 200. 

Leicester also (Culic. Malaya, 138) divides the Malayan species 
into groups, in an analytical table. 


C. albolineatus, Giles. 


7 vpe in British Museum. 


C. albopleura, Theob., 1907. 
Monog. Culic., iv, 456, 2. 


Loc.—India [Christophers}. Described from a perfect unique 
specimen. 
Tvpe in British Museum. 


C. albus, Leices., 1908. 
Culic. Malaya, 148, @. 


One 2 from a bungalow at Kuala Lumpur. 


Calis; Theobs, 1903. 
Monog. Culic., iit 
Loc.—Christmas Island. 


N.B.—Described from a series bred by Dr. Durham from 
larvae from salt pools. Miss Ludlow says it occurs in the Philip- 
pines. 

Type in British Museum. 


1 Oba iieetregh Neigh 


y 


C. angulatus, Theob. (angulata emend.). 


App. REF.—Blanch., Moust., 362, @. 


C. annuliferus, Ludlow. 


See /udlowi, Blanch., nom. nov., annuliferus being pre- 
occupied. 


C. annulioris, Theob., ror. 
Monog: Culiesi1; 371,.-2%. 
N.B.—Described froma single @ from Mashonaland, but it 
has since been found to occur in the Philippines. 
C. annulus, Theob. 


App. RreF.—Blanch., Moust., 293, 2 . 


Leices., Culic. Malaya (reproduces Theobald’s description of 
thei"), 


IgI2. | E. Brunetti: Catalogue of Oriental Culicidae. 467 


C. arabiensis, Patton, 1905. 
Journ. Bomb. Nat. Hist. Soc., xvi, 633, 7 2. 


In the crater, Aden. 


N.B.—Not, correctly speaking, within the Oriental Region, 
but included because I included Patton’s species from Arabia in 
my Catalogue. 


C. argentinotus, Banks, IgIo. 
Phil. }outneocl,, 1V,.547, 2.2. 


Loc.—Rizal (Phil. Is.). Types 7 2 (No. 11,460) in the ento- 
mological collection, Bureau of Science, Manila. 


L. auratus, Leices., 1908. 
Culic. Malaya, 153, 2. 


One of the largest species of Culex; a vicious biter ; some 
affinity with C. occidentalis and C. flavifrons. 


C. aureostriatus, Dol. 


App. REF.—Blanch., Moust., 310 @. 
ce 


N.B.—Blanchard gives Tokio as well as Amboina, in 
houses ”’ as localities, but the species is not included as Japanese 
in the recent Palaearctic catalogue. Theobald in his last volume 
stili retains it doubtfully in Culex. 


Cy biro:, Theob. 


App. REF.—Theob., Monog. Culic., iv, 390, 7 9%, fig. 165, 
wing o, 164, wing 92. 

Described from 3 7 @ and3 2 @. Tyfein the Hungarian 
Museum. 


Gcaecus, Iheopb: 


Type in British Museum. 


C. cantans, Meig. 
Erratum.—Delete my note about C. maculatus, Mg., in my 
Catalogue. 
C. christophersi, Theob., 1907. 
Monog. Culic., iv, 453, 7 @. 


Loc.—India [Christophers]. Described from rt @ and several 
aeor 
Type in British Museum. 


468 Kecoras of the Indian Museum. [Vou. IV, 


C. concolor, R. Desv. 


App. REF.—Blanch., Moust., 365, 7 9. 
Leices., Culic. Malaya, 154, 7 9. 

App. Locs.—Sylhet, ii, iv, v, xi [Hall]; Manipur, viii 
[Gourlay|; Calcutta, I-viii-o7; 6-x-04; 4-xi-o6 [all Brunetti]; 
ili, vi, vil, vili, ix, x, xi; Port Canning; Purnea, 4—6-viii-07 
[Pawa]; Damukdia Ghat (N. Bengal), vii; Lucknow, 4-ix-05 
Brunetti]; Kulattupuzha (Travancore), Ig-xi-o8, in bungalows 
[Annandale]; Samarang [Jacobson]; Phrapatoon, Siam, viii-o6 
{Woolley}; Andaman Is. [Ray White]; also in the Straits, Burma, 
and China. 

Leicester says, ‘‘ Fairly abundant in Malay Peninsula, very 
variable, larvae in stagnant pools, stable tanks or drainage.” 

Theobald observes that Giles erroneously refers it (‘‘ Journ. 
Trop. Med.,’’ 1904, p. 383) to Taeniorhynchus. 


C, fatigans, W. 


App. Syn.-—Desvotdea panalectros, Giles; ¢.. Theob., Rec. 
Ind. Mus., iv, 5. 

ADD. REF.—Theob., Monog. Culic., iv, pl. xvi (larval chars.). 

Blanch., Moust., 353, 7 2, fig. 230 @ 92 (as pungens), 
231 2,232 head o» @ (anxifer), 233 adult larva and 
nymph (pungens), 234 head of larva above and below 
(pungens). 

Leices.,, Culic: Malaya, 157,10 (2). 

App. Locs.—In the Rec. Ind. Mus. (ii, 298) Theobald gives a 
long list of dates and localities from specimens in the Indian 
Museum collection, and adds a further list in Rec. Ind. Mus., 
IV LZ: 

Between the two reports nearly all parts of India are repre- 
sented by this common species, with specimens from Nepal, Man- 
dalay, Rangoon, Moulmein, Manipur; Soerabaya, Java, 16—25- 
vii-06 [Brunetti|, Manila, 10—16-iii-06 [Brunetti], and at sea be- 
tween the mouth of the Hooghly River and Rangoon, 22—~23-11-08 
[Annandale]. 

Personally, I have taken it in Calcutta in February, March, 
April, July, October, November and December ; Meerut 25-iv-05 
(common in bedroom), Lucknow 7-vill-05, 4-ix 05, 2-xil-04, in bed- 
rooms and at the old Residency, Agra 28-iii-o5, Rangoon 24-x1i-04 
to 3-105 ‘common in bedroom), 9-ii-06, Penang 3—S-viti-06, 
Singapore 2I-vi-06, Batavia 27-vi and 9-vii-06, Soerabaya 16—25- 
vii-o6 common, Shanghai 16—25-vii-o6, where it was literally 
swarming in a ditch in front of a hedge facing the west ; Hankow 
22~-206-iv-06, Manila 10—1r16-iii-o6. Nearly all the specimens are 
in inferior condition, and were named by Mr. Theobald. 

Theobald in his 5th volume adds the following localities :— 

Phrapatocn, Siam, 1018-1; 19-iii; 30-xil-07; viii and xi-06 
[Dr. Woolley]; Sarawak. Outside the East it has a very wide 
range, Japan, Natal, Mombasa, Pemba Is. (E. Africa) ; Zanzibar, 


IgI2.| E. BRUNETTI: Catalogue of Oriental Culicidae. 469 


Soudan, Egypt, British Central Africa, Senegambia, Mauritius, 
Seychelles, Australia, Fiji, and the southern part of North America. 

N.B.—-Both Theobald and Blanchard consider the species iden- 
tical with pungens, W., and this latter name being given first by 
Wiedemann, although on the same page, should perhaps be 
adopted. Inthe absence of absolute certainty of identification and 
on account of the confusion that would be caused by the change, 
neither author adopts pungens. It is doubtful if the type of 
pungens still exists in any state sufficiently good to decide so close 
a point of identity. 

Moreover, my own personal opinion, not as a culicidologist 
but as being fairly intimate with the magnificent work done by 
Wiedemann, is that that author must have had two distinct 
species before him, for he certainly would not have regarded such 
minute differences as specific, as are to-day delighted in by the 
workers in Culicidae. 

In a paper just published by Mr. F. W. Edwards! fatigans, 
W., is sunk (with some species of Theobald’s) as a synonym of 
pipiens, Ll. If the two are identical it is very strange no previous 
author has ascertained it. 

The species carries Filariasis. 

Miss Ludlow’s tables show that it is common in all parts of 
the Philippines all the year round. 

Leicester says it occurs in houses all over the Malay Peninsula, 
the larva breeding in any small collection of water near houses. 


C, fatigans, var. trilineatus, Theob. 


Admitted as a good species. 


C, foochowensis, Theob. 
App. Rrer.—Blanch., Moust., 344, # 2 (fouchowensts). 
Type in British Museum. 
C. fragilis, Ludlow. 


App. REF.—Theob., Monog. Culic., iv, 424, 7 2. 
App. Locs.—Oras (Samar, Phil. Is.}, Aug. 6th [¢. Ludlow]. 


N.B.—Type in Army Medical Museum, Washington. 


C. fuscanus, Wied. 


App. Rer.—Blanch., Moust., 275. 


C. fuscocephalus, Theob., 1907. 
Monog. Culic., iv, 420, 9. 


Loc.—Peradeniya, March and September [Green]; Hakgala, 
Ceylon, 5-xi-07 (Green) ; Pallode, 15-xi-o8 [Annandale |. 


! Bulletin of Entomological Research, ii, 262 (Oct. 1911). 


470 Records of the Indian Museum. [Voz. IV, 


Described from 2 @ @. 
Type in British Museum. 


C. graminis, Leices., 1908. 
Culic. Malaya, 158, @ 9. 


Described from a large series from larvae from open bamboo 
in jungle ; sylvan species. 


C. gnophodes, Theob. 


App. REF.—Leicester copies Theobald’s description, ? . 
Type in British Museum. 


C. halifaxii, Theob., 1903. 


Placed with uncertainty here, only one specimen known 
(Theob., Monog. Culic., v, 350). Tvpe in British Museum. 


C. hirsuteron, Theob. 


Type in British Museum. 


C. impellens, Wk. 


ADD. REF.—Blanch., Moust., 294, @. 
Leices., Culic. Malaya, 142 (copies Theobald’s 
description). 


App. Locs.—-A long list from specimens in the Indian Museum 
(Theob., Monog. Culic.,iv, 15). Thaumaspur, Nepal, 1o—14-ii-08, 
Sukwani, 15—16-1i-08 ; Moulmein, Peb.; Mandalay, 11—12-iii-08 ; 
Rangoon, 24—25-1i-08 ; Lucknow, 21-iv-07 [all three Annandale] ; 
Chittagong, 26-vii-08 [Hall]. Sukna, in deep jungle; Puri, x-o8, 
common; Port Canning, 9-x-08, also several localities in Travan- 
core and Cochin States collected 4—25-xi-08, all taken by Dr. 
Annandale. 

Calcutta, I-ili-vi to ix; on board ship 5 miles off Alleppey, 
Malabar Coast, 4-v-08 [Paiva]. Also taken by me in Calcutta 
(bedroom), I-iv-08; 10-11-07, June, July: Lucknow, 7-viii-05 ; 
Rangoon, 24-xii-94 to 3-1-05, and Batavia, 27-vi-06 to 9-vii-06. 


C. imprimens, WIk. 
App. ReF.—Blanch., Moust., 306. 
Type (remnant) in British Museum. 
C. infula, Theob. 


ADD. ReF.—Blanch., Moust., 298, ?. 
Leices., Culic. Malaya, 146 (copies Theobald’s 
description). 
Type in British Museum. 


1912. | E. BRUNETTI: Catalogue of Oriental Culicidae. 471 


C. japonicus, Theob. 


Theobald (in vol. v, 391) admits this species from Tokio and 
Ceylon, making the latter a distinct variety which he terms ceylon- 
ica. The types of both forms are in the British Museum. He 
places ceylonica doubtfully in Culex, eliminates aureostriatus. Dol., 
from the synonymy, placing the latter separately and doubtfully 


in Culex. 
C. longifurcatus, Theob. 


See pseudolongifurcatus, Theob., nom. nov. 


C. longipalpis, V. Wulp. 


App. REF.—Blanch., Moust., 318, @. 
N.B.—This author correctly notes that although Van der 
Wulp says the palpi have only two joints, that author shows four 


in his figure. 
C. longipes, Theob. 


See macropus, Blanch., nom. nov. 


C, loricatus, Leices., 1908. 
Culic. Malaya, I51, @. 


One ¢ taken in bungalow at Kuala Lumpur. Very distinct. 


C. ludlowi, Blanch., nom. nov. 


Syn. C. annuliferus, Ludlow, 1903 (Journ. N. Y. Ent. Soc., 


it, FAT), 
App. REF.—Theob., Monog. Culic., iv, 388, 7 @. 
App. Loc.—Mangarin and Dagupan (Phil. Is.), Feb. to April. 


N.B.—The species was described from 234 7 o@ and 28 @ 9. 
but many were in bad condition. Blanchard renames the species, 
annuliferus being preoccupied by Em. Blanchard in 1852 for a 
species from Chili. 


C. luteola, Theob., rgro. 
Monog. Culic., v, 378, ¢. 
Peradeniya, x-1900. A single perfect 2, in the British 
Museum. A very distinct species. 
C. luteolateralis, Theob. 


The genus Bankstella, Theob., is established for this species, 


472 Records of the Indian Museum. [Vou IV, 


C. macropus, Blanch., nom. nov. (1905). 
Comp. Rend., lili, 1045. 


Blanch., Moust., 327. 
For longipes, Theob., preoccupied. 
Type in British Museum, a unique specimen from Singapore. 


C. mediolineatus, Theob. 


App. Rer.—Blanch., Moust., 369, 2. 
Type in British Museum. 


C. microannulatus, Theob. 


App. Syn.—C. volandi, D’Emm. de Ch., Ann. Trop. Med. et 
Par., ii, No. 3, 259 [1908]. 

App. REF.—Blanch., Moust., 292, 7 @. 

Leices., Culic. Malaya, 140, ~7 9. 

App. Locs. —Calcutta, July to November, common; Purnea, 
August ; Gopkuda Is., Lake Chilka, August ; Sylhet, i, ii, iv, v, 
xi, xii; between Bolpore and Rampore Haut, Bengal, August, in 
railway carriage [all ¢. Theob. in Ind. Mus.}. 


N.B.—Weicester describes it from a series sent him by Dr. 
Finlayson from Singapore, mostly taken in houses. I have only 
taken it once, in Calcutta, I—10-viii-o8. Green has taken it at 
Trincomalee and Hakgala, Ceylon, 24-viii to 9-ix-o7. It occurs 
in Mauritius. 

Type in British Museum. 


C. mimeticus, Noé. 


App. RreF.—Blanch., Moust., 271, 7 @. 
Leices., Culic. Malaya, 139, 7 2. 

App. Loc.—Lushai Hills, r-vi-o4, one 2 [Macleod]; Theog, 
Simla district, 2-v-o7, one @ [Annandale]; Thaumaspur, Nepal, 
18—20-ii-08, one @ ; Peradeniya, 17-ix-07 [Green]. 

N.B.—Weicester says he describes the o for the first time, 
but this is not so. He says it occurs in any roadside pool or 
marshy ground. Blanchard notes its occurrence in Italy and 
Palestine, also Malacca. 


C. minimus, Leices., 1908. 


Culic. Malaya, 160, 7 @. 


Described from a series bred trom larvae from mud holes full 
of water, in swampy ground in Kuala Lumpur. 


16124 E. BRUNETTI: Catalogue of Oriental Culicidae. 473. 


C. minor, Theob., 1908. . 
Rec. Ind. Mus., ii, 298, 7 9? ; iv, pl. 1, wing. 


Monog. Culic., v, 363, ¢ 2, fig. 150, wing. { 
App. Locs.—Sylhet [Hall]: Lushai Hills. June, July [Mac- 
leod|; Calcutta, December [ Annandale]. 


N.B.—Described from 3 ¢ @ and 2 9 @ in the Ind. Mus. 
coll. ‘A very small obscure species, easily told by its unbanded 
abdomen.’’ 

Type in Indian Museum. 

C. multimaculosus, Leices., 1908. 
Culic. Malaya, 155, 7 2 

Described partly from jungle examples and partly from larvae 
from roadside ditches. 

C. nigricephalus, Leices., 1908 (emend. mihi). 
Culic. Malaya, 149, 7 @ (nigricephala). 


Bred from paddy swamps near Batu Gajah by Dr. Daniels. 


C. pallidostriatus, Theob., 1907. 
Monog. Culic., iv, 410, o 9, fig. 175, wing o. 


Loc.—Peradeniya, December [Green]; India [Christophers. 
Described from 2 7 @ anda @°. 
Type in British Museum. 


C. pallidothorax, Theob. 


EKmend. by Theob. in Monog., iv, 446, from pallidithorax. 
ADD. REF.—Theob., Monog. Culic., iv, 446, 7 @ 
‘* Something like C. fatigans, Wied.’’ 
Type in British Museum. 


C. parascelos, Theob., tgro, 


Rec. Ind. Mus., iv, 18, @. 


Theob., Monog. Culic., v, 379. 

Described from 2 ¢ 2? from Madras Town, 30-x-08 [Hodgart]. 
A very marked species. 

Type in Indian Museum. 


C. perplexus, Leices., 1905. 


Culic. Malaya, 150, 7 9. 


Bred from larvae from marshy edges of lake near Kuala 
Lumput. 


474 Records of the Indian Museum. [VOE.sEV; 


C. pettigrewii, Theob., rgro. 
Recwitid? Mus. av, 5s - 


Theob., Monog. Culic., v, 351, 2. 

A perfect ¢@- from Ukhrul, Manipur. viii-o8, taken by the 
Rev. W. P. Pettigrew. 

In the Indian Museum. 


C. pseudolongifurcatus, Theob., nom. nov., Igro. 
Monog. Culic., v, 366. 


A new name for his /ongifurcaius (Rec. Ind. Mus., iv, 19, ~ 2), 
that name being preoccupied by Becker in 1903. Described from 


I 7 and 2 ¢ 2 from Dahawangahary Hill, Bengal—Nepalese 
Frontier, 16-ii-08. 
Type in the Indian Museum. 


C. pseudostenoetrus, Theob., 1g1o. 
Monog. Culic., v, 343, 2, fig. 154 wing. 


Two @ @ from Hakgala, Ceylon, v and viii, 1907 [Green]. 
Whereabouts of type not mentioned. 


C. pulchriventer, Giles. 


ADD. REF.— Blanch., Moust., 338, 7 @. 
Type in British Museum. 


C. pullus, Theob. 


Removed to Culiciomyta. 


C. quasipipiens, Theo! . 


ADD. REF.—Blanch., Moust., 344, @ . 
Type in British Museum. 


C. quasiunivittatus, Theob 


Type in British Museum. 


C. reesii, Theob. 


App. REF.—Blanch., Moust., 361, 7 ¢. 


C. rizali, Banks. 


Theobald (Monog., v, 391) notes on its differences from japo- 
nicus, to which it is closely allied. He puts it doubtfully here. 


1912.] EE. Brunerri: Catalogue of Oriental Culicidae. 47 


Un 


C. sepositus, Leices., 1908. 
Culic. Malaya, 152, 2. 


A single @ from jungle near Kuala Lumpur. Very distinct. 


C. sericeus, Theob. 


ApD. REF.—Blanch., Moust.. 362, ° 

App. Locs.—Taken by me at Meerut, India, 25-iv-05, Luck- 
now (the Residency), 2 xii-04; Rangoon, 24-xii-04 to 3-1-05, in 
bedroom ; Shanghai, 8—r10-v-06, inditch. The specimens identified 
by Theobald with some doubt owing to their rubbed condition. 


C. sitiens, Wied. 


App. REF,—Blanch., Moust., 293, °. 
Leices., Culic. Malaya, 143, 7 @. 
App. Locs.—Blanchard adds Sumatra, Celebes, Malacca ; 
Theobald adds Calcutta—Aug. and Sept. Philippines (¢. Ludlow). 
Leicester describes it from a series bred from marshy ground 
and from adults taken in the bungalows in the Malay Peninsula. 


C. stenoetrus, Theob , 1907. 
Monog. Culic., iv, 395, @. 


1oc.—Maskeliya, Ceylon, April [Green]. 
Type in British Museum. 


C. taytayensis, Banks, Ig1o. 
Phil. Journ. Sci., iv, 545, @ 2. 


Loc.—Rizal (Ph. Is.). Type 7 2 in entomological collec- 
tion, Bureau of Science, Manila. Bred from larvae from the water 
ot the esteros. 


C. tigripes, Grand. et Char. 


App. Locs.—Manipur (bungalow), August [Gourlay|; Sylhet, 
ii, iv, v-xii-04 [Hall]; Mandalay, 11-11-08 [ Annandale]; Calcutta, 
July to November ; Damukdia Ghat [Riv. Ganges, E. Bengal), 
July; Port Canning, Dec.; Kurseong, 4-viii-o8. I took it in 
Calcutta, Aug. 1908 and Dec 1905; Kirindi, 20-xio8; Weligama, 
'3-i-08 ; Dondra, 28-iv-o8. and Mandulsima, 14-xii-o8 (all four in 
Ceylon). Also occurs in South, Central and West Africa. 

The larvae are carnivorous and cannibalistic. Patton found 
it at Aden in a tank of rain water feeding on C. fatigans larvae. 

Theobald describes three varieties, none being Oriental. 


C. tipuliformis, Theob. 


App. Locs.—Blanch., Moust., 363, 2. 
Type in British Museum. 


476 Records of the Indian Museum. [Vor. IV, 


C. trilineatus, Theob., 1gor. 
Monog; Culic., ii, 159, 2. 


Blanch., Moust., 330, 2. 

Loc.—Thayetmyo, Upper Burma. First regarded as a variety 
of fatigans. Theobald admits it (Monog., v, 359) as distinct. 

Type, a unique specimen, in the British Museum. 


C. trimaculatus, Theob 


Type in Hungarian Museum. 


C. tritaeniorhynchus, Giles. 


App. REF.—Blanch., Moust., 294, 7 @?. 
Type in British Museum, from Madras. 


C. uncus, Theob. 


App. REF.—Blanch., Moust., 350, 2. 
Type in British Museum. 


C. uniformis, Leices., 190s. 
Culic. Malaya, 159, ¢. 


Two @ ¢ from marshy ground near Batu Gajah. Very near 
C. virtdis. : 
C. univittatus, Theob. 


App. RE¥F.—Blanch., Moust., 321, 2. 
Type in British Museum. 


C. vagans, Wied. 


App. Rrer.—Blanch., Moust., 304, 9 (Foochow). 
Theob., Ree: Ind, Mus:; iv 14-0 += Monog. Culic. v1 347.0% 
Theobald describes the ~ of this species, as the present 


example referred to (Madras Town, 31-x-08, Hodgart) is the only 
one of this species he has seen. It is in the Indian Museum. 


C. viridiventer, Giles. 


ADD. REF.—Blanch., Moust., 346, 7 @. ‘The 2 is said not 
to bite. 
Type in British Museum. 


C. vishnui, Theob. 
App. REF.—Blanch., Moust., 292, 7 2. 
Theob., Monog. Culic., iv, 387, 7, fig. 162, wing 
?; 163, o genitalia; pl. v, wing scales, ¢. 


1g12.] E. BRuNET?II: Catalogue of Oriental Cultcidae. 477 


Leices., Culic. Malaya, 141, ~ @ (description of 
@ copied from Theobald). 

App. Locs.—Sylhet, Jan. and Feb. [Hall]; Port Canning, 
July ; Gopkuda Is., August; Lake Chilka, August; Ferozepore, 
Punjab (Adie). 

Leicester has taken one 2 from jungle near Kuala Lumpur 

Type in British Museum. 


BANKSIELLA, Theob., 1907. 
Monog. Culic., iv, 468. 


B. luteolateralis, Theob., 1gor. 
Monog. Culic., 11, 71 (Culex 2d.). 


App. REeF.—Blanch., Moust., 278, 7 2. 
eices |; Culic. Malaya, 160% ‘or. 
Theob., Monog. Culic., iv, 469, ° 


N.B.—Banksiella Giemuced for this species, which, origi- 
nally described from the Soudan, has been recorded by Theobald 
from Sylhet, 23-xi-04 and 5-x-04, taken by Lieut.-Col. Hall, and now 
in the Indian Museum. Leicester describes it from larvae from the 
margins of small pools in grounds of the Institute of Medical Re- 
search at Kuala Lumpur. Taken there also in the adult stage. 
Occurs in several parts of Africa, where it varies considerably. 

Type in British Museum. 


TRICHOPRONOMYIA, Theob. 
App. REF.—Theob., Monog. Culic., iv, 479. 


Ds panel Theob. 


App. REF.—Theob., Monog. Culic., iv, 479 o,. fig. 213a, 
apex of proboscis; b, scales; fig. 214, wing o. 


TRICHORHYNCHOMYIA, mihi., nom. nov. 


Syn. Trvichorhynchus, Theob. 

App. Rer.—Theob., Monog. Culic., iv, 270. 

Comes between the Stegomyian group and true Culex (Theo- 
bald). 


N.B.—Trichorhynchus was preoccupied as far back as 1887 
by Balbiani in Protozoa. The above name is therefore suggested 
in its place, with the presumption that it only represents a sub- 
genus, at most. 


T. fuscus, Theob. 
ADD. Rer.—Theob., Monog. Culic., iv; 270, @. 


b 


Evratum.—In my catalogue ‘‘a single perfect @’’ is an error 
for 9. 


478 Records of the Indian Museum. [VoL. IV, 


Type in British Museum, according to Theobald (loc. cit., vol. 
v), but in a previous letter to me he says in the ‘‘ Hungarian 
Museum.” 


PSEUDOTHEOBALDIA, Theob., 1907. 


Monog. Culic:, iv, 271. 


bf 


P. niveitaea, taai Theob., 1907. 
Loc. ctt., 272, fig. 87, » genitalia; 88, wing o. 


Loc.—Dehra Dun, February, March [Thomson]. 
Described from two perfect 7 o. 

Theog, Simla district, 2-v-07 [Annandale]. 

Type in British Museum. 


TAENIORHYNCHUS, Arrib. 


App. REeF.—Theob., Monog. Culic., iv, 483 generic and egg 
notes. 
Blanch. Moust., 381, fig. 244 generic chars. 7 9; 
table of spp. p. 383. 
Leices., Culic. Malaya, 163 table of six Malay 
spp. 
T. ager, Giles. 

App. REF.—Blanch., Moust., 385, 7 @. 

Leices., Culic. Malaya, 168, 7 2. Recorded from 
Kuala Lumpur, and Westlake, Hankow, 27-v1-07 
| Cornford}. 

App. Locs.—Sylhet, Jan. to May [Hall]; Miss Ludlow says 
it has been recorded from the Philippines. Calcutta, 2-11-08 ; 
Balighai, near Puri, 26-x-08 ; Mandalay, 12-11-08 ; Pallode, S. India, 
14-xi-08 ; Kerumadi (S. end of Vembanaad Lake, Travancore 
State), 6-xi-08 [all Annandale}. 


N.B.—Weicester describes it from a large series bred from 
larvae from water at marshy edges of lakes, and entangled in 
floating green weed at the sides of a running stream. 


T. argenteus, Ludlow. 


Removed to Theobaldiomyia, nom. nov. for Leucomyia. 


T. (2) aureosquammatus, Ludlow, 1909. 
Can Ent., xli, 234, @ |! 


Theob., T (?) aureosquamata (lapsus ?) Monog., v, 425. 
Loc.—Parang, Mindanao (Phil. Is.), December. 
Tvpe in Army Medical Museum, Washington 


|! Theobald (Monog., v) erroneously quotes the page as Iol. 


1912.| E. BRUNETTI: Catalogue of Oriental Culicidae. 479 


T. aurites, Theob. 
T. brevicellulus, Theob. 


Both referred to Chrysoconops. 
Type of latter in British Museum. 


T. confinnis, Arrib., 1891. 
Dipt. Argent. La Plata, 49. 


Theob., Monog. Culic., ili, 289. 
Recorded from Chaca, Formosa, possibly incorrectiy, as it is 
a South American species. 


T. conopas, Frnfid. 


Referred to Chrysoconops. 


T. domesticus, Leices., 1908. 
Culic. Malaya, 169, 7 ¢. 


Described from a series bred from larvae, and from adults 
from bungalow at Kuala Lumpur and elsewhere. 


T. epidesmus, Theob., 1910. 
Rec. Ind. Mus , iv, 22-9. 


Theob., Monog, Culic., v, 429, 2. 
Described from a single perfect 2 in the Indian Museum 
taken by Mr. Paiva at Bhogaon, Bengal, 2-x-o8. 


T. lineatopennis, Ludlow. 
App. RrrF.—-Theob., Monog. Culic., iv, 489, ?. 


N.B.—The types (in the Army Medical Museum, Washington) 
were labelled ‘‘ inside screens of screened houses,’’ Sept. 13, I4. 


T. luteoabdominalis, Theob., gro. 
Rec. Ind. Mus., iv, 23 @. 
Theob., Monog. Culic., v, 423. 


A single perfect @ from Katihar, Purnea (Bengal), 5-x-08 
[Paiva]. Very near T. epidesmus. 
Type in Indian Museum. 


T. ochraceus, Theob. 


Referred to Chrysoconops. 


480 Records of the Indian Museum. [ VOL: iV, 


T. pagei, Ludlow, 7 @. 


Given in Theobald’s vol. v, 618, without reference to author’s 
description, which is copied verbatim. 
From Parang, Mindanao, Phil. Is., Oct. 27. 


T. tenax, Theob. 


App. REeF.—Blanch., Moust., 386, @. 
Leices., Culic. Malaya, 167, @. 

App. Locs.—Sylhet, March to June [Hal/]; Manipur, August 
[Gourlay|; Balighai, near Puri, 23-x-08, at light, one @ [Anunan- 
dale}. 

N.B.—Leicester describes from one 9 taken ina bungalow at 
Kuala Lumpur. 

Type in British Museum. 


T. tenax, var. ocellata, Theob., 1907. 
Monog. Culic., iv, 488, @. 


Loc.—Kuching (Sarawak, Borneo), November [Dr. Barker}. 
This form apparently also occurs in China (v. Theob., Monog., iii, 
259). 

N.B.—There isa var. maculipes, Theob. (Monog., iv, 488), from 
African localities (White Nile, etc., v, pl. iv, wing scales), and one 
termed maculipes arabiensis by Patton found in the Aden Hinter- 
land. 

Tvpe in British Museum. 


T. whitmorei, Giles. 


Theobald says (Monog., v, 431) that Mr. Carter has examined 
the type and reports it a distinct ‘‘ Leucomyia.’? See Thec- 
baldiomyta. 

Type in British Museum. 


CHRYSOCONOPS, Goeldi. 


Os. Mosq. no Para, I14. 
Theob., Monog. Culic., iv, 491. 


C. aurites, Theob. 


Removed here from Taentorhynchus. 

App. Loc.—Kuala Lumpur, 10-xi-o2 and 25-v-02 [Durham]. 
Miss Ludlow says it has occurred in the Philippines. 

7 ype in British Museum. 


1912.] E. BRUNETTI: Catalogue of Oriental Culicidae. 481 


C. brevicellulus, Theob. 
Removed here from Taeniorhynchus. 


ADD. REF.—Blanch., Moust., 389, 7 @. 
Leices., Culic. Malaya, 163, 7 92. 

App. Loc.—Ceylon [Green]; Sylhet, Saugar, Manipur Haut 
(Feb., May, June), Manipur, Aug. [Gourlay]; Calcutta, August; 
Philippines (¢. Lud/., Can. Ent., xli, 234). 

Leicester describes from a large series taken in bungalows at 
Kuala Lumpur. He says the & is described for the first time, 
but Blanchard antedates him. 

Type in the British Museum. 


C. conopas, Frnfld. 


App. REF.—Blanch., Moust., 387, 2 (conopus). 
Leices., Culic. Malaya, 166, 9. 
N.6.—Blanchard spells it conopus, but Theobald reverts to 
the original spelling. Miss Iudlow says it has been recorded from 
the Philippines. Leicester describes from a single @ from Klang 
jungle, saying it is a very distinct and easily recognized species. 


C. ochraceus, Theob. 


App. REF.—Leices., Culic. Malaya, 164. 

Described from specimens taken in bungalows at Kuala Lum- 
pur. 

Referred here from Taeniorhynchus by Theobald. 

Type in British Museum. 


C. pygmaeus, Theob., 1908. 
Rec. Ind: Mus. i. 300, 2. 


Loc. ctt., iv, 25, @ pl. i, wing, pl. ili wing scales: Monog. 
Culic., v, 435, 7 @, fig. 192 wing, 193 head @, 194 genitalia o, 
195 wing @. 

Described from a single perfect 2 from Sylhet [Hal/]. 

The @ described later, from examples from Purnea, Raj- 
mahal, and Calcutta in July and August. 

Type » 2 in Indian Museum, co-fype @ in British Museum. 

Dr. Annandale says the eyes of the @ are iridescent in life. 


d 


MANSONIA, Blancl 


App. REF.—Blanch., Moust., 375, generic chars. ~ 2. 
Leices., Culic. Malaya, I71. 


482 Records of the Indian Museum. [VoL. IV, 


M. annulata, Leices., 1908. 
Culic. Malaya, 174, 9. 


Described from examples from bungalows at Kuala Lumpur 
and elsewhere near rivers. Near uniformis. 


M. annulifera, Theob. 


ApD. REF.—Blanch., Moust., 380, ?. 

Leices., Culic. Malaya, 174, @ (copies Theobald’s 
description). 

App. Locs.—Sylhet, May, June [Hall]; Manipur [Gourlay] ; 
Calcutta, Aug. and Dec.; Port Canning, Dec. [Annandale]; Pur- 
neah, Oct. ; Bhogaon, 7-vili-og [both Bengal, and Paiva]. 

N.B.—At one time placed by Theobald in Mansoniotdes:! 

Type in British Museum. 


M. annulipes, Wlk. 


App. Syn.—Mansonia nero, Dol., t. Blanch., Moust., 380, ¢@. 

ADD. REF.—Leices., Culic. Malaya, 172, 7 @. 

This latter author describes both sexes from a series taken in 
jungle near Klang, Kuala Iumpur, Port Swettenham and Jugra. 

Very local, troublesome in jungle; probably a river breeder, 
according to Leicester. 

I took one in Calcutta, 6-x-04. 

Type in British Museum. 


M. arabica, Giles, 1905. 
Journ. Trop. Med., May 1906, 130. 


Theob., Monog. Culic., v, 451. 
Loc.—Isle of Barham, North Arabia. 
N.B.—Properly speaking, should not be included in Oriental 
lists. 
M. chrysogona, Knab, 1909 (November). 
Entom. News Philad., xx, 386, 7 @. 


‘“Chrysoconopas aurites,”’ Ludlow. 

Described from one ~ and two ? @? from Parang, Mindanao 
(Phil. Is.), 31-v-06. 

Type (cat. No. 12,626) in United States National Museum. 

In describing this species Miss Ludlow was under the impres- 
sion she had Taeniorhynchus aurites, Theob., before her (¢. Kab). 


M. seguini, Laveran (Panopliies 1d.). 


Blanchard admits this species asa good one (Moust., 380, @ ), 
from Hanoi, Tonkin, taken in the military hospital during the 


1 ** First Rep. on Culic. in Ind. Mus. ColJ.,’> Rec. Ind. Mus., ii, 287—302 
(1908). 


Ig12.] E. Brunetti: Catalogue of Oriental Culictdae, 483 


fever season (July to September), where it was found to suck 
blood. ‘Theobald (Monog. Culic., iv) notes that he has seen no ex- 
ample of the species, and that from the description he considered 
it identical with wntformis. 

In the ‘‘ Genera Insectorum” the name seguwini is not men- 
tioned. Theobald’s latest suggestion is ‘‘ close to, if not, wniformis, 
Theob.” 


” 


M. septempunctata, Theob. 


App. Rer.—Theob., Monog. Culic., iv, 494, ?. 
Closely related to WM. annulipes, W1k. 


M. uniformis, Theob. 


App. REF,—Blanch, Moust., 379, 2. 

Leices., Culic. Malaya, 171, @ 9. 

App. Locs.—Sylhet 1, ii, ili, v, vi, vii, xii [Hall]; Mani- 
pur, June and Sept.,in stable and bungalow; Gopkuda Is., 
Lake Chilka, August; Bhogaon, 30-ix-o8 and Oct.; Katihar, Oct. 
[both Paiva]: Balighai, near Puri, 23-x-08; Travancore and Cochin 
States, 19-xi-o8 and 4-xi o8, ‘‘ very common in bungalows;’’ Cal- 
cutta, 2-vii-07; Rangoon, 25-11-08 [all Annandale]; Kuala Lumpur. 


N.B.— Leicester says it isin some parts of Kuala Lumpur the 
most troublesome mosquito after C. fatigans, W. He describes the 
@ for the first time. He has only bred one example of the species ; 
from a larva taken in a swamp. 

Very common throughout the Philippines all the year round, 
according to Miss Ludlow’s extensive tables, though she notes it is 
much more abundant during the winter. 

It is recorded from many parts of Africa, also Madagascar 
and Australia, and it is a very variable species. 

Tvpe in British Museum. 


MANSONIOIDES, Theob., 1907. 


Monog. Culic., iv, 498. ‘‘ closely related to Mansonia.’’ 


M. annulifera, Theob. 


Replaced in Mansonia. 


M. septemguttata, Theob., 1907. 
Monog. Culic., iv, 499, 2 , figs. 226, 227 wings @. 


Loc.—Sarawak [ Dr. Barker]. 
Type in British Museum. 


484 Records of the Indian Museum. [VoL. IV, 


ETIORLEPTIOMYIA, Theob. 


Syn. O’Reillia, Ludl., 1905. Can. Ent., xxxvii, Ior. 

App. REF.—Theob., Monog. Culic., iv. 505. 

Leices., Culic. Malaya, 178 (‘‘ position undetermined”’). 

Spelt Etorleptiomyia originally and placed in Corethrinae, but 
Theobald now considers it ‘‘undoubtedly culicid,’’ though the 
proboscis is characterised as very weak. 


E. completiva, Leices., 1908. 
Culic. Malaya, 178, o. 


Leicester thinks that the unique specimen sent him by Dr. 
Finlayson of Singapore, and from which this description was 
drawn up, represents the ~ of some species of this genus. 
Theobald only describes the ? of his genus. 


E. [uzonensis, Ludl. 


App. REF —Theob., Monog. Culic., iv, 506, @. 
Type in Army Medical Museum, Washington. 


MELANOCONION, Theob. 


App. RreF.—Theob., Monog. Culic., iv, 507, further generic 
notes. 
Glen Herrick, Ent. News Philad. (1905), p. 282. 
Blanch., Moust., 395, 7 2. 
Leices., Culic. Malaya, 136. 


M. juxtapallidiceps, Theob., rgto. 
Monog. Culic., v, 456, @. 


A single @ inthe British Museum, from Trincomalee, Ceylon, 
taken by Green, Oct. 1907. 


M. ornatus, Theob. 


App. REF.—Theob., Monog. Culic., iv, 508 @ , fig. 231 wing °. 


M. pallidiceps, Theob. 


App. REF.—Theob., Monog. Culic., iv, 509 @. 


M. uniformis, Theob. 


App. ReF.—Leices., Culic. Malaya, 136, 7 @. 

Redescribed by Leicester from bred examples from hollow 
bamboo and other jungle trees. Probably entirely sylvan. Leices- 
ter notes that his present description corrects some errors in his 


IgI2. | E. BRUNETTI: Catalogue of Oriental Culicidae. 485 
previous one sent to Mr. Theobald for publication in ‘‘ The Entomo- 
logist.’’ 

OCULEOMYIA, Theob., 1907. 


Monog. Culic., iv, 515. 


O. fulleri, Ludl., 1909 
Cane Bate, xii, 975. 9. 


Theob., Monog. Culic., v, 478 @ (Ludlow’s descr. copied). 
Loc.—Parang, Mindanao, Phil. Is., October. 


N.B.—Miss TL,udlow spells the genus Oculiomyza. 


O. sarawakii, Theob., 1907. 


Monog. Culic.,iv, 515 @ , fig. 236, head and @ haltere; 237, 
wing 2; pl. vi, wing scales 9°. 
Loc.—Sarawak [Dr. Barker}. Described from a unique speci- 
men. 
Type in British Museum. 


POPEA, Ludlow. 


This genus is now placed by Theobald (Monog., iv) between 
Leicesteria and Howardina. It is, by the way, almost preoccupied 
by Poppea, Stal., 1867, in Hemiptera. 

P. lutea, Ludlow. 


Type in Army Medical Museum, Washington. 


RACHIONOTOMYIA, ‘Theob. 


App. REF.—-Theob., Monog. Culic., iv, 518. 
In this volume Theobald places this genus next to Oculeomyia 


R. ceylonensis, Theob 


ADD. REF.—-Theob., Monog. Culic., iv, 518, @. 
Type in British Museum. 


FINLAYA, Theob. 


App. RerF.—Blanch., Moust., 415. 


N.B.—Mr. Theobald (Monog., iv) considers this genus and 
Orthohbodomyia as intermediate between Culicinae and Aedinae. 


F. aranetana, Banks. 


Loc.—Negros Occidental, Phil. Is., 17—24-vi-1g06 


486 Records of the Indian Museum. (VoL. IV, 


F. poicilia, Theob. 


App. Rrer.—Blanch., Moust., 415. 


N.B.—Theobald notes that the figure of wing scales on 
page 283 (Monog., ili) is slightly incorrect, and shows a corrected 
wing in vol. iv, 520, fig. 238, with other notes. Papuan specimens 
differ a little from Malayan ones. The species is recorded from 
N. Queensland by Dr. Bancroft. 

Type in British Museum. 


ORTHOPODOMYIA, Theob. 


App. Rrer.—Theob., Monog. Culic., iv, 527. 
Leices., Culic. Malaya, 175. 
Theobald admits this as intermediate between Culicinae and 
‘ Aedinae.”’ Leicester (Joc. cit.) notes its affinities with Aedeo- 


myinae. 
O. albipes, Leices. in Theob. 


App. REF.—Theob., Monog. Culic., iv, 527, 7 @. 
Leices.,, Culick Malaya, 1760517024: 
Leicester describes in above work from a long series bred from 
larvae from bamboos. Sylvan, probably not a blood-sucker. 
Type in British Museum. 


Of maculata, Mheobss 19x0: 
Rec. dards. 1visn2G., e orn 


Monog. Culic., v, 473, &. 
A single »: Indian Museum, from Maddathorai, 17-xi-08 
[| Annandale). 
O. maculipes, Theob., 1gro. 


Monog. Culic., v, 470, fig. 208 head @ , 209 wing, 210 
wing @ (2 error for o~). 

Loc.—Andaman Is., 22-vii-o8 [Lowis and Ray White]; Pera- 
deniya, Ceylon, 5-vii-og [Green]; Maddathorai, S. India, 17-x1-08 
[Annandale]. 

Tvpe @ Indian Museum, ? British Museum. 


O. nigritarsis, var. 


Leicester notes (Culic. Malaya, 177) a new variety of this 
species, stating its affinities with O. albifes, but I have found no 
mention of any species of the nate of nzgritarsis. 

The specimen was taken in the hollow of a tree on a small 
island, Pangkor-Haut, by Dr. Daniels. 


REEDOMYIA, Ludlow. 


Syn. Lepidotomyia, Theob., Ann. Mus. Hung, ili, 80. 
App. REF.—Reedomyta, id., Monog. Culic., iv, 257, 7 2. 


1g912.]} E. Bruner: Catalogue of Oriental Culicidae. 487 


N.B.—Theobald (l.c., iv, 249) notes that his Lepzdotomyza in 
Ann. Mus. Hung., iii, referred to, and is a synonym of Reedomyta. 

His second genus under this name stands good, with at pres- 
ent four Oriental species. 


R. alboscutellata, Theob. 


Removed here from ‘‘ Leptdotomyra.”’ 
App. REF.—Theob., Monog. Culic., iv, 261 ? , fig. 80 wing @. 
Leices , Culic. Malaya, 132, 7 2 (Lepidotomyza id.). 

The latter author redescribes it from a series of adults from 
jungle near Kuala Lumpur, and from bred specimens from larvae 
from a jungle pool. Theobald spells this species alboscutella in 
vol. v, 257; presumably in error. 

Type in Hungarian Museum. 


R. lowisii, Theob., 1910. 


Monog. Culic., v, 257, 7 @; fig. 121 ¢ head, 122 wing, 
123 head o , 124 wing o. 
Loc.—Andaman Isles [Lows and Ray White]; Galle, Ceylon, 
6-iv-07 [Bainbrigge Fletcher]. 
Type in British Museum. 


R. niveoscutellata, Theob. 


ADD. REF.—Theob. 
fore ungues. 

Resembles Culex pipiens. Miss Iudlow says it has been 
recorded from the Philippines. 

Tvpe in British Museum. 


Monog. Culic., iv, 259, 7 @ ; fig. 79 @ 


’ 


R. pampangensis, Ludlow. 


App. ReF.—Theob., Monog. Culic., iv, 258, 2. 
Type in Army Medical Museum, Washington. 


Sub.-Family AEDEOM YINAE. 


App. Rers.—Aedeinae, Blanch., Moust., 398 (table of genera) 

Aedinae, Theob., Monog. Culic., iv, 537. 

Aedeomyinae, Leices., Culic. Malaya, I8o. 

Leicester gives notes on sub-family, also table of genera, in 
which he includes Deinocerites, Theob., Aedeomyia, Theob., Aedes, 
Mg., Atoretomyia, Leices., gen. nov., Acalleomyia, Leices., gen. 
nov., Verrallina, Theob., Mimomyia, Theob., Uranotaenia, Theob..,! 
Ficallia, Theob., Hodgesia, Theob., Zeugnomyta, Leices., gen. nov., 


' Incorrectly attributed to Theobald instead of Arribalzaga, 


488 Records of the Indian Museum. [Vor. IV, 


Colonemyia, Leices., gen. nov., Topomyza, Leices., gen. nov., Haema- 
gogus, Theob., Skeivomyza, Leices., gen. nov. Several of these genera 
are not represented in the East. 

Inhis ‘‘ genera of the Aedes type’ Col. Alcock sinks Mimomyta, 
Theob., and Pseudoskusea, Theob. 


SKUSEA, Theob. 


By some authors referred to this sub-family, but herein it is 
retained in its position in my Catalogue. 


LEPTOSOMATOMYIA, Theob. 


Evratum.—In my reference to this genus (Cat., p. 362) change 
Dp: S0-to pe mo: 


L. lateralis, . Theob. 


App. REF.—Theob., Monog. Culic., iv, 548 7 ; fig. 252, head, 
scutelium ungues & 

The type is partly dissected. I am uncertain whether it 
reposes in the British Museum or the Hungarian Museum. 


AEDEOMYIA, Theob. 


App. RrF.—Blanch., Moust., 403, ~ @ , fig. 255, var. generic 
chars. 
eices, Culic: Malaya. rom 
This author says he cannot agree with Theobald in the rela- 
tionship of this genus to Aedes. ‘‘Even the = palpi, the only 
point of agreement, are very different from those in Aedes.’’ 


A. catasticta, Knab, 1909. 


Ent. News Philad., xx (November), 387, ?. 


He only describes the 2 , apparently, but he must have had 
both sexes present as, in comparing it with A. sguamitpenna, Arrib.., 
he says ‘‘the @ genitalia of the two species show specific differ- 
ences: « 

A. squamipenna, Arrib. 


(Aedeomyia squammipenna, Theob., Monog., vol. ii, ¢. Leices.) 


App. REF. 


Blauch.. Moust., 404. 9 (squamtpennts). 
Leices., Culic. Malaya, 182, @ 2 (squammtpenna). 

App. Locs.—Example; taken occasionally during the year in 
bungalows at Kuala Lumpur; Calcutta, at ight, November; base 
of Dawna Hills, 2-iti-08 [Annandale]; at light at sea, 4 miles off 
Tuticorin, 25-v-08 [ Parva]. 

N .B.—Weicester reverses the correct quotation of this species, 
making it appear asif Arribalzaga’s name was a synonym, whereas 
the species was first described by him. 


1912. | E. Brunerrt: Catalogue of Oriental Culicidae. 489 


PSEUDOGRAHAMIA, Theob., rgto. 


Rec. Ind. Mus., iv, 26, near Grahamia, Theob. 


P. aureoventer, Theob., 1gto.' 
boexcil, 275. 2 . 


Loc.—Pallode, Travancore State, S. India, 16-xi-08, a single ? 
[Annandale]. 

‘“A very marked and beautiful species which cannot be con- 
fused with any other culicid.’’ 


N.B.—Care must be taken to avoid confusion through the 
similarity of these generic names, Grahamia, Pseudograhamia, Grab- 
hamia and Pseudograbhamia. 


SQUAMOMYIA, ‘'Theob., IQI0. 
Ree. Inds. Mus; 1v,-28. 


Theob., Monog. Culic., v, 529. 


S. inornata, Theob., 1910. 
Rec. Ind. Mus., iv, 28, 7. 


Theob., Monog. Culic., v. 529, &. 
Described from a single ~ in the Indian Museum from the 
Dawna Hills (2—3 000 ft.), 2 or 3-iii-08 [Annandale ,. 


AEDES, Mg. 


App. REeF.—Blanch., Moust., 399,77 @. 
Leices. , Culic. Malaya, 183. 

‘This genus is closely related to Culex and Melanoconion on 
the one hand, and less so to Atovetomyia amongst the Aedeomyinae. 
In fact, nothing could better show how unscientific is a classifica- 
tion based on palpi, than that genera so closely related as Culex, 
Melanoconion and Aedes should be placed in different families ”’ 
(Leicester). 


ae 


N.B.—-Leicester, I hope, means ‘‘ sub-families. ”’ 


A. butleri, Theob. 


N.B.—In his Monog., iii, 295, Theobald created the genus 
Verralina for Aedes butleri and two other species; and Blanchard 
(Moust., 417) retains the genus, but in the ‘‘ Genera Insectorum ” 
Theobald does not mention it, referring butlert to the *‘ uncertain 
‘position ”’ section of the species of Aedes. 

Type in British Museum. 

| This species has inadvertently been dealt with before, see p. 460. The 
mistake was discovered too late for correction in the text.—Ed. 


490 Records of the Indian Museum. [VOL. IV 


A. malaya, Leices., 1908. 
Culic. Malaya, 184,°¢?. 


Bred from larvae from pond in Kuala Lumpur. 


A. nigrescens, Theob., 1907. 


Monog. Culic., iv, 540, fig. 246 wing 2, 247 o genitals, 
pl. vi, wing scales @. 


Loc —Castle Rock (Canara Dist., S.-West India), Jan. to 
March. 


‘A very small species, looking like a Melanoconion’’ (Theob.). 
Type in British Museum. 


AIORETOMYIA, Leices., 1908. 


Culic. Malaya, 185. ‘‘ Near Verrallina and Howardina.” 


A. aedes, Leices., 1908. 
(LOGIC. COR ore 


One o& bred from a larva from a pond in Kuala Lumpur, the 
species doubtfully referred by the author to this genus. 


A. ostentatio, Leices., 1008. 
OG RCIL A elOGk ee 


Described from a series of 2 92, some from larvae from a 
jungle pool, and some taken as adults. Sylvan species, a vicious 
day biter. 


A. perdita, Leices., 1908. 
EOC MEU OZ oe 
Description drawn up from recollection from a single ~ taken 
at midday in jungle five miles from Kuala Lumpur. 
The tv£e is lost. 
A. singularis, Leices., 190%. 
LOCC SO. ors 


A single # from a bungalow in Kuala Lumpur. 


A. taeniata, Leices., 1908. 
EOC RCIEG OOD 


Two @ @ only taken at midday in jungle near Kuala Lumpur. 


1912. ] E. BRUNETTI: Catalogue of Oriental Culicidae. 491 


A. varietas, Leices., 1908. 
LOCK Ctt., T8554 2%. 


Fairly common in Malayan jungle, where the @ bites viciously. 
Occurs at Kuala Lumpur, Ulu Gombak, Pangkor Haut and Klang. 
Leicester only bred one example, a ~. 


AGAEEEBOMYIA, Weices., 1908. 
EDC. 6tl_. TOA: 


A. obscura, Leices., 1908, emend. mihi. 
Loc. ctt., 194, & @ (obscurus). 


No notes or dates are given. 


VERRALLINA, Theob.; 1903. 
Monog. Culic., iii, 295. 
ADD. REF.—Leices., Culic. Malaya, 196. 


V. butleri, Theob. 
(Aedes. id.) Theob., Monog. Culic., ii, 230 


App. Re¥F.—Leices., Culic. Malaya, 196, 7 2. 

The o@ described for the first time. The commonest mosquito 
in the mangrove swamps; not unlike Stegomyia fusca, Weices. : 
larva living in brackish pools in mangrove swamps. 

N.B.—Al\though Theobald created Verrallina for this species, 
he ignores the genus in the ‘‘ Genera Insectorum,”’ placing butleri 
in the ‘‘uncertain position ’’ section of Aedes. But in vol. v he 
replaces it in the present genus 

Type in British Museum. 


V. fragilis, Leices., 1908. 
Culic. Malaya, 199, o@. 


A sylvan species in jungle near Kuala Lumpur. 
N.B.—Possibly the same species as malayi; if so, the name 


fragilis must stand. 
V. imitator, Leices., 1908. 
EGeaciis 201, 9 . 


Two 2? ¢ from jungle five miles from Kuala Lumpur. 
Types rather damaged, but distinct. 


492 Records of the Indian Museum [Vor. IV, 


V. indecorabilis, Leices. 


, 1908. 
Loc.-ctt., 


ZOO Or Sie 


Bred from larvae from small jungle pool near Kuala Lumpur. 
Very near zmitator. 


Wi malayi, Leices., 1908. 
Loe. cit. Oo ree 
One ¢ in jungle ten miles from Kuala Lumpu 
? 2 of fragilis (Leicester). 


V. virilis, Leices., 1908. 
OG. Cite, MOF 


One o& from jungle a few miles from Kuala Lumpur 
Near V. malayi 


BOLBODEOMYIA, Theob., 1910 


Ree: Inds Mus. av oi. 


B. complex, Theob 
I FASC: 


5, IQLO. 


SS Slite ieaacee 

Monog. Culic., 

o, 255 wing @. 
Loc.—Dawna Hills 


V, 581, 7 @, fig. 253 @& genitalia, 254 wing 
, 2—3,000 ft., I— 3-11-08 [ Annandale]. 
Types (a unique pair) in Indian Wereere 


N.B.—There is a Bolbodimyia by Bigot, 1892, in Tabanidae 
(Wien. Ent. Zeit., xi, 162). 


MIMOMYIA, Theob. 


REF.—Blanch., Moust. 
IGeIces:: 


ADD. Moust., 419. 
Culic. Malaya 


a, 202; notes, apparently 
criticising its place in this sub-family 


M. chamberlaini, Ludlow 
Vide Ludlowia. 


M. minuta, Theob., 1908. 


Rec. Ind. Mus. 
Theob., Loc. cit., iv, 30, 2 ; pl. i wing, pl. 111 wing scales 
Id., Monog. Culic., v, 531, o 9, fig. 226 wing. 
Locs.—Sylhet, 27-xi-o4 [Hall]; Calcutta, 
4-vili-07 [| Annandale]. 


Types in British Museum. 


Sites Oiectore 


30-vii and 3 or 


1Q12. | FE. BRuNnET?I1: Catalogue of Oriental Culicidae. 493 


RUNCHOMYIA, Theob. 


Syn. Binotta, Blanch., 1904 (Archiv. de Parasit , viii, 478). 
Blanchard’s name, suggested, on account of alleged preoccu- 
pation (Rhynchomyia, R. Des., in Muscinae), cannot stand, the names 
not being identical. 


R. philippinensis, Giles. 


App. Rer.—Theob. , Monog. Culic., v, 555 (copies Giles’s descr.). 

Type in British Museum. Theobald has not seen it, but says 
that Banks says ‘‘ this species is identical with Uvanotaenia 
nitidoventer Giles, but both are incorrectly placed as to genera.”’ 
Mr. Carter also thinks the present species neither a Runchomvyia 
nor a Uvanotaeni«. 


LU DLOWIA, Theob:, 2907. 
Monog. Culic., iv, 193, o. 


Established for Mimomvyia chamberlaint, Iadlow, also to 
comprise a Soudanese species. 


L. chamberlaini, Ludlow. 
(Mimomyra td., Ludlow.) 
App. REF.—Theob., Moiiog. Culic., iv, 194, 7 (Miss Ludlow’s 
description copied). 


N.B.—This author adds that since vol. iv of his Monograph 
went to press Miss Ludlow has sent him a description of the @. 
Type in British Museum. 


L. minima, Ludlow, 1907. 
Can Hit, .cx xis gare an 12. 


Theob. 
description). 
Type in Army Medical Museum, Washington. 


, Monog. Culic., v, Ig1, 7 2 (copies Miss Ludlow’s 


ANISOCHELEOMYIA, Theob. 


App. REeF.—Theob., Monog. Culic., iv, 570. 


A. alboannulata, Theob. 
App, RreF.—Theob., Monog. Culic., iv, 573, o, fig. 267 wing 
@: pl. vii wing scales 9 
N.B.—Type (unique) in British Museum. ‘* The species is a 
very beautiful and marked one, and cannot be confused with any 
other ” (Theob.). 


494 Records of the Indian Museum. [Vor. IV, 


A. albitarsis, Ludlow. 


App. REF.—Theob., Monog. Culic., iv, 576, 9 (Miss Ludlow’s 
description copied). 

N.B.—Type in Army Medical Museum, Washington. Theo- 
bald thinks its place in this genus doubtful. 


FICALBIA, Theob: 


App. REF.—Blanch., Moust., 418. 
Leices., Culic. Malaya, 228. 


F. longirostris, Leices., 1908. 
Culic. Malava, 228, 7 ¢. 


Bred from larvae from stagnant water at Kuala Klang in 
January ; a very distinct species. Apparently slightly aberrant. 


F. minima, Theob. 


App. REF.—Blanch., Moust., 418 


N.B.—First placed in Uvanotaenia, then referred here. 

As Theobald (Monog., v) retains a species of this name under 
both genera, it is left here for the present. 

Type in British Museum. 


F. simplex, Theob. 
ADD. REF.—Blanch., Moust., 418. 
Theob., Monog. Culic., v, 541, 2 , fig. 235 wing. 


B] 


App. Loc.—Two ¢@ @ from Trincomalee, xi-1906 {Green|}. 
Type in British Museum. 


PSEUDOURANOTAENIA, Theob., 1905. 
Journ. Econ. Biol.,; i, 33. 


Theob., Monog. Culic., iv, 566, fig. 262 (p. 567) wing o. 


P. parangensis, Ludl., 1909. 
Can Ents xii 249. 


Theob., Monog. Culic., v, 524 (copies Miss Ludlow’s descrip- 
tion). 
Loc.—Parang, Mindanao (Phil. Is.). 


P. triangulata, Lud!., 1908. 


Can-wlnt. oxi 331 oe 


Theob., Monog. Culic., v 
tion). 
I1oc.—Reine Regente, Mindanao (Phil. Is.) ; February. 


525 (copies Miss Ludlow’s descrip- 


b 


1g12.]| E. Brunetti: Catalogue of Oriental Culicidae. 495 


URANOTAENIA, Arrib. 


App. Rer.—Blanch., Moust., 406, @ @ generic chars. 
Leices.. Culic. Malaya, 203 (genus incorrectly 
attributed to Theobald). 
Table of 18 new Malayan species. 


U. argyrotarsis, Leices., 1908. 
Culic. Malaya, 214, 7 @. 


Described from series bred from larvae from a pool in a patch 
of jungle five miles from Kuala Lumpur, and one adult on surface 
of same pool. Very distinct. 


UW? atra, Cheob- 
App. REF.—Theob., Monog. Culic., iv, 563, 2 . 


U. bicolor, Leices., 1908 
Culic. Malaya, 225, 7” @. 


Bred from larvae from the marshy edges of a jungle stream 
in Kuala Lumpur. Leicester has not captured the adult, which 
is the largest Malay species after maxima. 


U. bimaculata, Leices., 1908. 
L0G. c1t., 226, 9x 


Fairly common in damp places of jungle at ‘‘ The Gap,”’ 
Kuala Lumpur, the only place it seems to occur at. 


U. bimaculiala, Leices., 1908. 
Lotscit., 208. a 3 . 


The o from Raub jungle; the @ described from a series 
caught over jungle pools (presumably at Kuala Lumpur). Near 
untmaculiala. 


U. caeruleocephala, ‘heob., var. lateralis, Ludlow. 


In vol. v Theobald makes it a species. 


U. campestris, Leices., 1908. 
Culic. Malaya, 212, 7 2°. 


Very near Verrallina pygmaea, Theob. Captured on swampy 
ground. 


406 Records of the Indian Museum. [Vor aayE 


U. cancer, Leices., 1908. 
LOC. Ct, 2n5,on Oe 


Abundant in crab holes at Port Swettenham, and among 
Nepah paims at that place, and Klang. larvae in crab holes or 
stagnant pools. Male very distinct. 


U. ceylonica, Theob., 1910. 
Monog. Culic., v, 503, 2 , fig. 213 wing. 


Loc.—Galle, Ceylon, 10-iv-07 | Bainbrigge Fletcher]. 
Type (a unique specimen) in British Museum. 


UW: fusca, Veices., 1608: 
Culic. Malaya, 227, 7 @. 


Described from a series bred from larvae from a pool at 
Sungei Limbing, Malay Peninsula, taken by Dr. Daniels. 


U. lateralis, Ludl. 


Phil. Journ: Scix 4. .coo: 
Syn. U. caeruleocephala, Th., var. lateralis, Ludl. Mosq. Phil. 
Ses palo: 


U. longirostris, Leices., 1908. 
IBOGS Che ODE 7e oe 


Only one specimen knowr, bred from a pupa taken in a pond 
(presumably at Kuala Lumpur). 


U. lutescens, Leices., 1908. 
Loc. Cit..222 


A bamboo sylvan breeder, Leicester has not taken the adult 
form. Presumably from Kuala Lumpur. 


U. maculipleura, Leices., 1908. 
Culic. Malaya, 223, °. 


Only one specimen known, taken by a jungle stream six 
miles from Kuala Lumpur. 


U. malayi, Theob. 


ADD. REF.—Blanch., Moust., 410. 
Type in British Museum. 


1gI2.| E. BRuNE?TII: Catalogue of Oriental Culicidae. 497 


U. maxima, Leices., 1908. 
Culic. Malaya;.221, 2% 


Described from specimens taken at ‘‘ The Gap.’’ Selangor. 
The largest Malay species. Dr. Leicester possesses a ” which may 
be that of this species. 


U. micans, Leices., 1908. 
oc. Cit. 200. ar 9: 
Described from a series taken on marsh land, Malaysia. 
U. minima, Uheob: 
See Ficalbia 1d. 
U. modesta, Leices., 1908. 
Culic: Malayay 2718, vo" 2. 
Described from a series bred from larvae from water in a 
hollow tree in Ampang jungle. 
U. nitidoventer, Giles. 


See Runchomyia philippinensts. 


Wiaenivea,eices. 1908: 
LOG Ct... 2EL, 0". 
Only one specimen known, taken by Dr. Leicester in his 
bungalow at Kuala Lumpur. 
U. nivipleura, Leices., 1908. 
FOC: (Cita. 2G. os. 


Described from a single pair: o@ taken by Leicester at ‘* The 

> . . 
Gap,” Selangor, @ sent by Dr. Finlayson, bred from a larva in 
a pitcher plant at Singapore. 


U. powelli, Iudl., 1909 
Cana) nites aiinee ss, 42. 


Theob., Monog. Culic., v, 519, ?. 
The locality is given as Yayabas (Phil. Is.), which is probably 
a misprint for Tayabas. January. 


U. testacea, Theob. 


App. REF.—Theob., Monog. Culic., iv, 560, 2 , fig 258 wing @. 
App. Loc.—Base of Dawna Hills, 4-iii-o8 [Annandale]; Phil. 
Is. [¢, Ludlow]. 


498 Records of the Indian Museum. [VoLz TVs, 


U. trilineata, Leices., 1908. 
Culic. Malaya, 204, 2 


I,arvae found in quiet pools, adults in neighbourhood of jungle 
pools. The only species in Malaysia with a banded proboscis. 


U. unilineata, Leices., 1908. 
Culic. Malaya, 220; 7 9. 


In jungle close to water, Leicester has found females full of 
blood. Malaysia. 


U. unimaculata, Leices., 1908. 
Culic. Malaya, 210, 2. 


Near water in jungle, Malaysia. oo unknown. 


ZEUGNOMYIA, Leices., 1908. 
Culic. Malaya, 231. 


Links Uranotaenta to Colonemyta and Skeiromyia, and thence 
to Wvyeomyia. 


Z. gracilis, Leices., 1908. 
Culic. Malaya, 232, 7 @. 


Faitly common in certain localities, especially October to 
December ; a vicious biter. Occurs all the year sparsely at Jugra 
and Kuala Lumpur. Larvae in water in fallen leaves, feeding on 
other larvae. (Lezcester). 


COLONEMYIA, Leices., 1908. 


Culic. Malaya, 233. 


C. caeruleocephala, Leices., 1908. 
LOC NCE eS ae OF, 


Described from a series of both sexes bred from larvae from 
bamboo water at Bukit Kutu and Ulu Klang (Malaysia). 


C. hybrida, Leices., 1908. 
Woenctt @e2a7 
One 2 at Bukit Kutu; quite distinct. 


1912.] E. Brunerri: Catalogue of Oriental Culicidae. 499 


C. mendacis, Daniels, 1908. 


Studies from Instit. for Med. Research (Fed. Malay States), 
iii, 266, 7-2. 


‘* Notes on the Mosquitoes on the river and coast district of 
the eastern side of the Peninsula,’? C. W. Daniels. 


N.B.—Sequential to Leicester’s paper. 


Bred from larvae from pitcher plants on East Coast, Malay 
Peninsula. 


TOPOMYIA, Leices:, 1908. 
Culic. Malaya, 238. 


Very near Zeugnomyta and Colonemyta. 


T. argyropalpis, Leices., 1908. 
0c. tis, 242) oF 2% 


A unique o from a stream at “‘ The Gap,”’ Selangor, in April. 
The unique 2 by a jungle stream five miles from Kuala Lumpur 
in March. 


T. argyroventralis, Leices., 1908. 
Loc. ctt., 240, 7 Q. 


Described from 2 7 @ and a series of @ @ from ‘‘ The Gap,”’ 
Selangor, near jungle streams. Possibly the two sexes described 
under this name by Leicester are not of the same species, but 
T. tipuliformis may be the real ~. (Leicester.) 


N.B.—If this should prove the case, the name must stand 
for the o@ according to zoological precedent, and ‘“‘ argyroven- 
tralis@”’? be sunk as the 2 of ‘‘ tepuliformis;’’ the present sug- 
gested o& of argyroventralis being either renamed or allowed to 
retain that specific name. 

T. decorabilis, Leices., 1908. 
Loc. ctt., 239, 2 


One o@ and @ at ‘‘ The Gap,’’ Selangor, in jungle. 


T. durbitans, Leices., 1908. 
Bore 240, 2., 


A unique 2 from ‘‘ The Gap ”’ in May. 


T. gracilis, Leices., 1908. 
Loc. ctt., 244, oo. 


‘*The Gap, ’’ Selangor, May. 


500 Records of the Indian Museum. [VOrL.sIvsS 


T. minor, Leices., 1908. 
Lockctie 236,019". 


In jungle near Kuala Lumpur near streams. Both sexes, 
when settled, have a habit of dancing up and down like certain 
Tipulidae. 


T. nigra, Leices., 1908. 
OCCt 2A 5 oe 


Near jungle streams at Ampang, six miles from Kuala Lumpur, 
May. 
T. rubithoracis, Leices., 1908. 


Loc. ciut., 243, 7 &. 


Described from two @ o@ and a series of 2? 2. 


T. tipuliformis, Leices., 1908. 
LOC. Cts, 2AF, Oe 


Several ~ @ near a stream in ‘“‘ The Gap’”’ jungle, and in the 
dried bed of a stream at Raub. Leicester suggests that this may 
be the o of ‘‘ vubithorax’’ (; lapsus). He also previously suggests 
it is the same as argvroventralis (vide note under argyroventralts). 

Abdomen very thin, longer proportionately than in any other 
species of the family, and in general appearance closely resembling 
several species of Tipulidae. 

SKEIROMYIA, Leices., 1908. 


Culic. Malaya, 248. 


S. fusca, Weices:, 1906; 
LOE. Ctt.5 ZAGA NC). @. 


Leicester has not seen the adult, except when bred, from 
larvae, which are common in bamboo. 


HODGESIA, Theob. 


App. REF.—Leices., Culic. Malaya, 229. 


N.B.—To the generic definition, Leicester proposes to add 
‘antennae pilose in @ and @?.’’ 


H. malayi, Leices., 1908. 
Culie; Malaya, 23n8 oo 


Described from a series bred from jungle pools near Kuala 
Lumpur. 


1912.] E. Brunetti: Catalogue of Oriental Culicidae. 501 


H. quasisanguinae, Leices., 1908, 
oe. cit. 230, 2% . 


In jungle near Kuala Lumpur; a vicious biter, 


H. sanguinea, Theob. 


Type in British Museum. 


WYEOMYIA, Theob. 


App. RrEF.—Leices., Culic. Malaya, 250. 


N.B.—Leicester admits a sub-family Wyeomyinae, with the 
following genera: Wyeomyia Theob., Phoniomyta Theob., Dendro- 
myia Theob., Runchomyia Theob., Sabethes, Sabethoides, Goeldia, 
Limatus, Malaya, Leices., all gen. nov. ; giving a table of them, 
several however not being Malayan. 


W. aranoides, Theob. 
App. REeF.—Blanch., Moust., 425. 
Type in British Museum. 
W. funerea, Leices., 1908. 
Culic. Malaya, 252, 2. 


A unique @ from jungle six miles from Kuala Lumpur. 


W. greenii, Theob. 


App. REF.—Theob., Monog. Culic., iv, 596. 
Blanch., Moust., 423. 
App. Loc.—Peradeniya [Green]. 


N.B.—Mr. Theobald omitted this spectes from the ‘* Genera 
Insectorum’’ but mentions it, as above, for a good species. 
Type in British Museum. 


W. metallica, Leices., 1908. 
Culic. Malaya, 251, ?. 


A unique @ in bungalow at Bukit Kutu, Malaysia. 


W. nepenthicola, Banks, IgIo. 
Phil Journ. sci. iv, 550, 7 ¢. 


Loc.— Benguet, Trinidad (Phil. Is.}. Bred from larvae in 
pitchers of Nepenthes alata, Bl. 

Type # @ (No. 8159) in entomological coll., Bureau of 
Science, Manila. 


502 Records of the Indian Museum. [Vor IV, 


PHONIOMYIA, Theob. 


bf 


Evratum.—“ Vol. ii’’ is a misprint for vol. ili in my Catalogue , 
p. 365. 
App. REeF.—Blanch., Moust., 425. 
Leices., Culic. Malaya, 253. 


N.B.—The name of this genus is likely to be confounded with 
Phonomyia established in Tachinidae by Brauer and Bergenstamm 
in 1894. 


P. bimaculipes, Theob. 
App. Rer.—Theob., Monog. Culic., iv, 600, @. 
‘A very distinct and beautiful species ” (Theob.). 
P. caeruleocephala, Theob., rgto. 

Monog. Culic., v, 577, 7 @ , fig. 252 wing. 
Loc.—Hakgala, Ceylon, iii-o7 [Green]. 
Types, a unique pair, in British Museum. 

-P. indica, Theob. 


App. REF.—Theob., Monog. Culic.,iv,601, 7, fig. 275 wing @. 

Redescribing the species in this volume, the author repeats, 
“‘Described from a perfect o,’ and again makes no direct 
reference to the @, yet he figures a wing marked @. Is this in 
error ? 

In vol. v he says simply that the ¢yfe is from Singapore. 

Leicester records a ~ and two 2 2 from jungle, six miles 
from Kuala Lumpur. 


POLYLEPIDOMYIA, Theob. 
App. REeF.—Theob., Monog. Culic., iv, 625. 


P. argenteiventris, Theob. 
App. REF.—Theob., Monog. Culic., iv, 625, fig. 292 wing @. 
N.B.—The tyfes were taken November and December 1892. 


DENDROMYIA, Theob., 1903. 
Monog. Culic., ili, 313. 
Blanch., Moust., 426. 
Leices., Culic. Malaya, 254. 
Syn. Heinzmannia, Ludlow. 


N.b.—Heinzmannia is sunk for Dendromyia. Mr. Theobald 
says (Monog., iv, 603) that, due to some error of his in writing to 


1912.]| E. Brunert1: Catalogue of Oriental Culicidae. 503 


Miss Ludlow, she founded her genus on a species which is an un- 
doubted Dendromyia. 
D. achaetae, Leices., 1908. 
Culic. Malaya, 257, 2: 


Described from a single example; genitalia very concealed, 
and sexes very alike in this genus. 


D. aureochaeta Leices., 1908. 
EE OGwOth. 255; 9 
A series of @ 2 taken by day in jungle near Kuala Lumpur 
in April and May. 
D. communis, Leices., 1908. 
Loc. ctt., 254, 2. 


Described from one ~ and a series of 9 2 from ‘‘ The Gap,” 
Selangor. 


D. scintillans, Ludlow. 
(Heinzmannia id., id.) 


This species was described from a perfect 2 , except that the 
antennae are missing. 
Type in Army Medical Museum, Washington. 


PHILODENDROMYIA, Theob., 1907. 


Monog. Culic., iv, 623. 


P. barkeri, Theob., 1907. 


Monog. Culic., iv, 623, @ , fig. 289 head, scutellum, palpus o , 
fig. 290 wing o , pl. vii, wing scales @. 


Loc.—Sarawak, in July [Dr. Barker]. Described from two 
¢ & taken in a house. 
Type in British Museum. 


MALAYA, Leices., 1908. 
Culic. Malaya, 258. 


M. genurostris, Leices., 1908. 
Loc. cit., 258, o. 


Taken by Dr. Daniels in bungalow in Malay Peninsula. 


504 Records of the Indian Museum. [Vo.. IV, 


HARPAGOMYIA, Melj., 1900. 
Tijd. voor. Ent., lii, 165. 


N.B.—The author places this simply in ‘‘ Aedeinae,’’ without 
any more definite note as to its exact position, so I place it here at 
the end of this sub-family. 


H. splendens, Meij., 1909. 
Tijdav. Ent? lit, 167, °0 9 “pli x, 1 —on(vare figs): 
Loc.—Java. 


N.B.—The author notes that this mosquito is eaten by the 
widely distributed ant Crematogaster difformis, Smith. 


Sub-Family CORETHRINAE. 


Corethyva and its allies form a sub-family of Culicidae, and 
cannot morphologically be separated from this family. The 
absence of a biting mouth, and the absence of scales on the body 
are quite secondary characters. 

Moreover, the discovery in Ceylon quite recently by Major 
MacDougall, R.A.M.C., of a new genus which Dr. Annandale has 
described as Ramcta! irrevocably links together, on account of its 
undeniably intermediate nature, the two groups Culicinae and 
Corethrinae. The short, feeble proboscis and absence of scales on 
the head, body and legs approximate it to the Corethrinae, the 
presence of scales on the distal half of all the longitudinal veins 
connects it with the Culicinae. T'wo peculiarities of venation 
distinguish it: the Ist longitudinal vein ends soon after the 
middle of the wing, running parallel to the auxiliary vein, instead 
of reaching the distal margin as in both Culicinae and Corethrinae, 
in this peculiarity resembling Phlebotomus ; and, secondly, the 2nd 
longitudinal vein begins almost in a line with the origin of the 3rd 
and comparatively close to it instead of some distance before it, as in 
both Culicinae and Corethrinae. Only the male is known, but the 
larva and pupa were also obtained and from the characters of 
these Dr. Annandale considers its total affinities lie mainly with 
the Corethrinae. 

Two short papers of mine® thoroughly: investigate the syno- 
nymy of Corethra, Mg., Sayomyia, Coq., and Chaoborus, Lichten- 
stein, with the result that it is found that no species of Corethra, 
Mg., occurs in the East, the genus being apparently restricted to 
two or three European species. The genus Sayomyza is antedated 
by Lichtenstein’s Chaoborus, as admitted by Coquillett himself, 
and both aszatica, Giles, and cornfordi, Theob., certainly belong here, 


1 Thus named, by special request of Major MacDougall, after the Royal 
Army Medical Corps. 


2 ** Synonymy in Corethrinae,’’ Rec. Ind. Mus., iv, 317, and vi, 227. 


1912. | E. BRUNETTI: Catalogue of Oriental Culicidae. 505 
whilst there can be no reasonable doubt that mantlensis, Sch., is 
also congeneric. 
RAMCIA, Annandale, fort. 
Spolia Zeyl., vii, pt. xxviii, p. 187 (Aug., Ig1T), 


R. inepta, Annandale, IgIr. 


Spolia Zeyl., vii, pt. xxviii, p. 189, fig. (p. 188) and plate giving, 
larva, pupa, wing, and other characters. 
Type in the possession of Major A. J. MacDougall, R.A.M.C. 


CHAOBORUS, Lichtenstein, 1800. 
Syns. Sayomyta, Coq. ‘‘ Corethra’’ Auct. 


C. asiatica, Giles. 


App. Locs.—Calcutta Zoological Gardens, May, July, August, 
‘*common, resting on damp walls during daytime and flying to 
light at night’? [Annandale]; also occurs in Calcutta, Septem- 
ber, November and December; Sibpur (near Calcutta), August ; 
Katihar, Purnea District, at light. 


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Acalleomyia Leices. 56 
acaudata Leices. 
achaetae Leices. 

aconita Don. (aconitus) 
Aedeomyia Theob. 
AEDEOMYIINAE Leices. 
aedes Leices. 

Aedes Mg. 

AEDINAE Theob. 

ager Giles 

Aioretomyia Leices. 
aitkenii James in Theob. 
albipes Leices. in Theob. 
albipes Theob. 
albirostris Theob. 
albitarsis Ludl. 
alboannulata Theob. 


alboannulatus James and List. 


albolateralis Theob. 
albolineata Theob. 
albolineatus Giles 
albopictus Skuse 
albopleura Theob. 
alboscutellata Theob. 


albotaeniata Leices. in Theob. 


albotaeniatus Theob. 
albus Leices. 
Aldrichia Theob. 
Aldrichinella Theob. 
alis Theob. 
Amauromyia Leices, 
ambigua Theob. 
amboinensis Doles. 
amesii Ludl. 
angulata Theob. 
Anisochelecmyia Theob. 
annandalei Theob. 
annandaiei Theob. 
annularis Wulp 
annulata Leices 
annulata Theob. 
annulata Theob. 
annulatus Blanch. (Culex) 
annulifera Theob. 
annuliferus Ludl. 
anuulioris Theob. 
annulipalpis Theob. 
annulipes W1k. 
annulirostris Theob. 
annulitarsis Leices. 
annuloabdominalis Theob. 
annulus Theob. 
Anopheles Mg. s. lato. 
; ids idoes. Str. 
ANOPHELINAE 
apicalis Theob. 


Teromyia 
Dendromyia 
Anopheles 


= AEDINAE 
Aioretomyia 


Taeniorhynchus 


Anopheles 
Orthopodomyia 
Stegomyia ais 
Myzomyia nit 
Anisocheleomyia 
id. ots os 
=Myzorhynchus  albotaeniatus 
Theob. 
Stegomyia 
Scutomyia 
Culex .. 
=Stegomyia scutellaris Wik. 
Culex 56 
Reedomyia 
Danielsia 
Myzorhynchus 
Culex .. 
=Aldrichirella © 


Culex ; ‘ ; 


Grabhamia cit 
Toxorhynchites .. 
Stegomyia 3¢ 
Culex sa 


Kingia ee 
Stegomyia 
Myzomyia ; 
Mansonia =f aie 
Culiciomyia % <U 
Trichopronomyia 

Theobaldia 

Mansonia 

Culex 

Culex 

Brevirhynchus 

Mansonia 

Stegomyia 

Leicesteria 

Culiciomyia 

Culex 


Blanchardiomyia (Desvoidya) .. 


508 Index. 


apicalis Theob. ee 36 
apicalis Theob. as a 
arabica Giles a 55 
arabiensis Patton 

arabiensis Patton F 

aranetana Banks ails 

aranoides Theob. 

argentea Ludl. (Taenio. argenteus) 
argenteiventris Theob. Se 
argenteomaculata Theob. 

argenteotarsis Ludl. 

argenteus Ludl. 


argentinotus Banks oe 
argyropalpis Leices. 

argyrotarsis Leices. 5c ore 
argyroventralis Leices. 3 oe 
Armigey Hartmann 

Armigeves Theob. 

asiatica Giles 

asiatica Leices. 

assamensis Theob. 


ater Daniels are a 
atra Theob. 

aurantia Theob. bs ate 
auratus Leices. a ays 


aurea Leices. 

aureochaeta Leices. 

aureolineata Theob. 

aureosquammatus Ludl. ae 
aureostriata Leices. Sg ate 
aureostriatus Doles. a6 we 
aureoventer Theob. ne 

aurites Ludl. (Chrysoconops) 

aurites Theob. oe 
aurostriata Banks 

auxifer 

azriki Patton 


Banksiella Theob. oe oO 
barbirostris v. Wulp Bie 
barianensis James Ac 

barkeri Theob. aS 

bicolor Leices. i 

bicornuta Theob. 

bimaculata Leices. & < 
bimaculiala Leices. oe oe 
bimaculipes Theob. 

Binotia Blanch. ae 
bipunctata Theob. ae 


biroi Theob. 

Bironella Theob. Be aC 

Blanchavdiomyia Brun. nom. nov. for 
Desvoidya. ae oie 

Bolbodeomyia Theob. Xe 50 

brevicellulus Theob. ; 

brevipalpis Giles 

brevipalpis Theob. 

brevipalpis Theob. 

Brevirhynchus Theob. 

butleri Theob. 20 ote 

butleri Theob. ate 56 


Page 
Brevirhynchus ae we 443 
Leicesteria or be élgul 
Mansonia oe ve AOL 
** Anopheles ”’ ss oa 434) 
Culex oe 467 
Finlaya ai 485 
Wyeomyia 56 501 
Theobaldiomyia .. e402 
Polylepidomyia .. sie 502 
Stegomyia ine -- 444 
Toxorhynchites ye 435 
Theobaldiomyia (Leucomyia), 
argentea Sc eeOz 
Culex 467 
Topomyia 499 
Uranotaenia 495 
Topomyia 499 
v. Blanchardiomyia _ 441 
id. : 441 
Chaoborus 505 
Lophoscelomyia 427 
Stegomyia 445 
Teromyia 438 
Uranotaenia 495 
Pardomyia 460 
Culex. 407 
Conopomyia 453 
Dendromyia set 1503 
Blanchardiomyia (Desvoidya). AAT 
Taeniorhynchus 478 
Ekrinomyia 440 
Culex 467 
Pseudograhainia see 400 
=Mansonia chrysogona Knab.. 482 
Chrysoconops 479 
Stegomyia 445 
=Culex fatigans We 468 
Myzomyia : 418 
* 
ae 477 
Myzorhynchus 424 
Anopheles be 415 
Philodendromyia 503 
Uranotaenia : 495 
Lophoceratomyia .. 463 
Uranotaenia 495 
id. 5c 495 
Phoniomyia S502 
=Runchomyia Theob. =. 4 498 
var, aa pane gem eee 
gelidus : 7403 
Culex .. 467 
SC 433 
Sc 440 
5 5 492 
Chrysoconops : 481 
Stegomyia. te 445 
Lophoceratomyia 464 
Toxorhynchites 435 
4c 442 
Aedes 489 
Verrallina 491 


+ 


caeca Theob. 

caecus Theob. 
caeruleocephala Leices. 
caeruleocephala Theob. 
caeruleocephala Theob. 
calopus Mg. 

Calvertia Ludl. 
Calvertina Ludl. 
campestris Leices. 
cancer Leices. 

cantans Mg. 

Carrollia Lutz 
catasticta Knab. 

Cellia Theob, 
ceylonensis Theob. 
ceylonica Theob. 
ceylonica Theob, 
Chactomyia Leices. 
Chagasia Cruz 
Chaoborus Lichtenstein. 
chamberlaini Ludl. 
christophersi Theob. 
christophersi Theob. 
Christophersia James. 
Chrysoconops Goeldi. 


Chrysoconopas aurites Ludl. | 


chrysogona Knab. 
chrysolineata Theob. 
chrysoscuta Theob. 
cingulata [eices. 
clavipalpis Theob. 
cohaesa 
Colonemyia Leices. 
communis Leices. 
completiva Leices. 
complex Theob. 
concolor Rob. Desy. 
confinis Arrib. 
conopas Frnfld. 
Conopomyia Leices. 
Corethva Mg. 
CORETHRINAE. 
crassipes v. Wulp. 
Culex L. 

Culicada Felt. 
culicifacies Giles 
culiciformis Cogill 


culiciformis James and List. 


culiciformis Theob. 
CULICINAE. 
Culictomyia Theob. 
cuneata Theob. 


Danielsia Theob. 
Dasymyia Leices. 
deceptor Don. 
decorabilis Leices. 
Dendromyta Theob. 
desmotes Giles 
Desvoidya Blanch. 
dissimilis Leices. 
diurna Theob. 
dolichocephala Leices. 
domesticus Leices. 
dthali Patton . 


Index. 


-“- 


Peconiyia ar 
Culex aC 
Colonemyia 
Uranotaenia Ac 
Phoniomyia 


=Stegomyia fasciata 
v. Calvertina 


Uranotaenia 
id. ae 
Culex” 7 FP, 


Aedeomyia 


Rachionotomyia 
Culiciomyia : 
Uranotaenia a 
v. Leicesteriomyia .. 
v. Calvertina 


Ludlowia 
Culex 
Myzomyia 


=Mansonia chrysogona Knab .. 


Mansonia. ei 
Howardina sts 
Pseudohowardina .. 
Leicesteria 
Radioculex 


var. Anoph. formosaensis 


Dendromyia ats 
Etorleptiomyia sje 
Bolbodeomyia 

Culex 
Taeniorhynchus 
Chrysoconops 


Stegomyia 


Myzomyia 

«* Anopheles ”’ 
Stethomyia 

Skusea oe 


Var. of ‘Theobaldiomyia (Leuco- 


myia) gelida 


* 


Myzomyia 
Topomyia 


Stegomyia 


—Blanchardiomyia nom. nov. Ae 


Stegomyia 

Skusea 

Leicesteria 
Taeniorhynchus 
Myzomyia és 


F. 


510 


dudgeoni Theob. 
durbitans Leices. 


Ekrinomyia Leices. 

elegans James 

eminentia Leices. 

epidesmus Theob. 

error Theob. o6 
eruthrosops Theob. ee 
Etiovleptiomyia Theob. ae 


fasciata F. are o% 
fatigans W. Se 

Ficalbia Theob. 

Finlaya Theob. 

flava Leices. 

flava Ludl. 

fluviatilis Leices. 

foochowensis Theob. Be 
formosaensis Tsuz. Ae iG 
formosus Ludl. Pe se 
fragilis Leices. oye ste 
fragilis Ludl. 

fragilis Theob. 

frater R. Desv. .. 

fraudatrix Theob, 

freerae Banks 

fuliginosus Giles 

fulleri Ludl. 

funerea Leices. ae 
funerea Theob. var. ornata Theob. 
funesta Giles 

funesta Leices. 

fusca Leices. 

fusca Leices. 

fusca Leices. 

fusca Theob. 

fusca Theob. 

fuscanus W. 

fuscocephalus Theob. 

fuscus Theob. 


gardneri Ludl. 
Gettonomyta Leices. 
gelidus Theob. 


id. id. var. cuneata Theob. 
id. id. var. sinensis id. 
genurostris Leices. ; 
gigas Giles 


gilesii Theob. 
gnophodes Theob. 
gracilis Leices. 
gracilis Leices. 
gracilis Leices. 
gracilis Theob. 
Grabhamia Theob, 
graminis Leices. 
greenii Theob. 
greenii Theob. 
gubernatoris Giles 


Index. 


Neocellia 
Topomyia 


* 


Neomyzomyia : 


Lophoceratomyia .. 


Taeniorhynchus 


Culicada 


Aldrichinella es 


* 


Stegomyia 


Culex é 


Verrallina 
Culex 


Stethomyia Ai 
=Steg. fasciata F. . 


Lophoceratomyia 
Pyretophorus 
Nyssorhynchus 
Oculeomyia 
Wyeomyia 
Skusea 
Myzomyia 
Teromyia 
Dasymyia 
Skeiromyia 
Stegomyia 
Uranotaenia 


Leicesteriomyia (Chaetomyia) .. 
Cellia bo 
Hulecoetom yia : 
Culex ; 
Anopheles 
Anopheles 


Blanchardiomyia (Desvoidya) i 


Culex 
Culex 


Trichorhynchomyia 


* 


Quasistegomyia 
= Pecomyia Theob. 


Theobaldiomyia (Le 


id. 
id. 


Malaya 


Anopheles 
Toxorhynchites 


Culex 
Stegomyia 
Topomyia 
Zeugnomyia 
Bironella 


Culex 

Howardina 
Wyeomyia 
Phagomyia 


* 


ucomyia) 
id. 


id. 


halifaxii Theob. 
halli James 
Harpagomyta Meij. 
Heinemannia Ludl. 
himalayana Giles 
hirsuteron Theob. 
Hodgesia Theob. 
Howardina Theob. 
Hulecoetomyia Theob. 
hybrida Leices. 
hybrida Leices. 


imitator Leices. 
imitator Leices. 
immaculatus Theob. 
immisericors WIk. 
impellens WIk. 
imprimens WIk. 
indecorabilis Leices. 
indefinita Ludl. 
indica Theob. 

indica Theob. 

indica Theob. var. simplex Theob. 
indica Theob. 
indiensis Theob. 
inepta Annandale 
infula Theob. 
inornata Theob. 
inornata Theob. 
inornatus WI1k. 
intermedia Rothwell 


jamesii Theob. 
japonicus Theob. 
jehafi Patton 

joloensis Ludl. 
jugraensis Leices. 
jugraensis Leices. 
juxtapallidiceps Theob. 


karwari James 

Kerteszia Theob. 

Kingia Theob. 

kochi Don. 

kumasi Chalmers (Anopheles) 


laniger W. 

lateralis Ludl. 

lateralis Theob. 

Letcesteria Theob. 

leicesteri Theob. 

Leicesteriomyia Brun. nom, nov. 

Leptdotomyia Theob. (I) 

Lepidotomyia Theob. (11) 

leptomeres Theob. 

Leptosomatomyia Theob. 

leucomeres Giles. 

Leucomyia Theob. 

leucophyrus Don, 

lewaldii Ludl. 

lindsayi Giles. 
ide 1d. 


var. maculata Theob. 


Index. 


Culex St oe 
Christophersia mC aie 


= —Dendromyia Theob. 
Howardina 
Culex 


Colonemyia 
Conopomyia 


* 


Stegomyia 
Verrallina 
Anopheles 
Toxorhynchites 
Culex Sy 

id. ie 
Verrallina ve 
Myzomyia 56 
Neocellia 
Neomacleaya 

id. 

Phoniomyia are 
Nyssorhynchus Sc Sc 
Ramcia ate ve 
Culex 
Culiciomyia 
Squamomyia 
Toxorhynchites 
Neocellia 


* 


Nyssorhynchus 

Culex 

Myzomyia ae 

Blanchardiomyia (Desv oidya) .. 
id. id. 

Hulecoetomyia 

Melanoconion 


* 


Nyssorhynchus 


Cellia 


=Myzomyia funesta, Giles 


* 


Mucidus 
Uranotaenia 
Leptosomatomyia .. 


Toxorhynchites _ 
for Chaetomyita Leices 
= Se Ludi. 


Myzomy ja 


Stegomyia 
=Theobaldiomyia mihi nom. nov. 
Neomyzomyia 
‘Yoxorhynchites 
Anopheles 
id. 


512 


lineata Ludl. .. 
lineatopennis Ludl. 

listoni List. 

longifurcatus Theob. ne 
longipalpis Leices, in Theob. 
longipalpis Wulp. : 
longipes Theob. 

longirostris Leices. 
longirostris Leices. 
Lophoceratomyia Theob. 
Lophomyia Giles 
Lophoscelomyia Theob. 
lophoventralis Theob. 
loricatus Leices. 

lowisii Theob. 

ludlowi Blanch. 


ludlowi Theob. is 
Ludlowia Theob. 

lutea Ludl. 2 
luteoabdominalis Theob. 
luteola Theob. 

luteolateralis Theob. 
luteolateralis Theob. a 
lutescens Leices. oe 
luzonensis Ludl. 


macropus Blanch. 
maculata Theob. 
maculata Theob. 
maculata Theob. 
maculata Theob. 
maculatus Theob. 
maculipalpis Theob. 
maculipes Theob. 
maculipes Theob. 
maculipes arabiensis Patton. . 
maculipleura Leices. 
magna Tkeob 
magnifica Leices. 
magnus Theob. 
Malaya Leices. 

malayi Leices. 

malayi Leices. 

malayi Leices. 

malayi Theob. 
mammilifer Leices. 
Mansonia Blanch. 
Mansonioides Theob. 
maxima Leices. 
megtegori Banks 
mediofasciata Theob. 
mediolineatus Theob. 
mediopunctata Theob. 
MEGARHINAE (MEGARHININAE) 
Megarhinus Rob. Desv. 
Melanoconion Theob. 
mendacis Daniels. 
metallica Leices. 
metallica Leices. 
metallicus Leices. 
micans Leices. 
microannulatus Theob. 
microptera Giles 
mimeticus Noé 
Mimomyia Theob. 


Index. 


Calvertina ae 
Taeniorhynchus .. 50 
Myzomyia ao 
v. Culex pseudolongifurcatus a0 
Leicesteria xc 

Culex 


v. Culex macropus Blanch. 
Ficalbia she 
Uranotaenia 


= Lophoscelomyia 1 “Theob. 
Pseudocarrollia 


Culex 
Reedomyia 


nom. nov. for Cale annuliferus 


Ludl. 
Myzomyia 


Popea 
Taeniorhynchus 
Culex 9¢ 
Banksiella 

Culex 0 
Uranotaenia ate 
Etiorleptiomyia 


* 


nom. nov. for C. longipes Theob. 


Orthopodomyia 
Pecomyia 2 
Pseudograbhamia .. 
var. Anoph. lindsayi Giles 
Nyssorhynchus 

id. 50 
Orthopodomyia .. 
var. Taenio. tenax 
var. id. id. 
Uranotaenia 
Lepidotomyia 
Teromyia 
Brevirhynchus 


Aedes 

Hodgesia 
Verrallina 
Uranotaenia 
Lophoceratomyia 


Uranotaenia 
Kertezia 
Skusea 
Culex 
Stegomyia 


Colonemyia 
Conopomyia 
Wyeomyia 
Toxorhynchites 
Uranotaenia 
Culex 
Stegomyia 
Culex 


minima Ludl. (Mimomyia) 
minima Theob. 

minimus Leices. 

minimus Theob. 

minimus Theob. 

minor Leices. 

minor Leices. 

minor Theob. 

minuta Theob. 

minuta Theob. 

minutissima Theob. 
minutissima Theob. até 
minutus Theob. 

modesta Leices. 

Mucidus Theob. ae 
mucidus Karsch. we 
multimaculosus Leices. 
multiplex Theob. 

Myzomyia Blanch. 
Myzorhynchus Blanch. 


Neocellia Theob. 
Neomacleaya Theob. 
Neomyzomyia Theob. 


Neostethopheles James and List. 


nepenthicola Banks 
nero Doles. (Mansonia) 
niger Leices. 
nigerrima Theob. 
nigerrimus Giles 

nigra Leices. 
nigrescens Theob. 
nigricephalus Leices. 
nigrifasciatus Theob. 
nigritarsis (auct. ?) var. 
nigrotarsis Ludl. 
nitidoventer Giles 
nivea Leices. 

nivea Ludl. 
niveoscutellata Theob. 
niveitaeniata Theob. 
nivipes Theob. 
nivipleura Leices. 
notoscripta Skuse 
nursei Theob. 
Nyssomyzomyia James 
Nyssorhynchus Blanch 


obscura Leices. 
obturbans WIk. 
ocellata Theob. 


ochracea Theob. aie 
ochraceus Theob. 

Oculeomyia Theob. ac 
O’ keillia Tudl. fe 
ornata Theob. 

ornatus Theob. 
Orthopodomyia Theob. 
ostentatio Leices. 


pagei Ludl, 
pallida Ludi. ‘ 
pallidiceps Theob. 


Ludlowia 

Ficalbia 

Culex Ne ae 
Toxorhynchites .. oe 
Pyretophorus 

Lophoceratomyia .. : 
Topomyia 30 st 
Culex 

Mimomyia 

Culicada 

Culiciomyia 

Stegomyia 

Myzorhynchus 

Uranotaenia 


Mucidus ; 
Culex are 
Pseudoskusea Se 


Wyeomyia 
M. annulipes WIk. . 
Lophoceratomyia . ee 
Culiciomyia ; 
Myzorhynchus we a 
Topomyia ae SC 
Aedes 
Culex 
Pyretophorus 
Orthopodomyia 
Pseudoskusea 
Uranotaenia 

id. 
Scutomyia 
Reedomyia 
Pseudotheobaldia .. 
Nyssorhynchus Be 
Uranotaenia AG 
Scutomyia 
Pyretophorus 


* 


Acalleomyia 

Blanchardiomyia (Desvoidya) 

var. of Taeniorhynchus tenax 
Theob. 

Grabhamia 

Chrysoconops 


= Etiorleptiomyia Theob. 
var. of Skusea funerea Theob... 
Melanoconion 


Aioretomyia 
* 


Taeniorhynchus .. 
Stethomyia 
Melanoconion 


514 


pallidostriatus Theob. 
pallidothorax Theob. 
pampangensis Ludl. 


panalectros Giles (‘* Desvoidea ”’) 


parangensis Ludl. 

parascelos Theob 

Pardomyia Theob. 
Patagiamvia James 

Pecomyia Theob. 
peditaeniatus Leices. 

perdita Leices. 

periskelata Giles 

perplexa Leices. 

perplexus Leices. 

pettigrewii Theob. 
Phagomyia Theob. a6 
philippinensis Ludl. “Je 
philippinensis Giles : 
Philodendromyia Theob. 


Phoniomyia Theob. 50 


pictus Lw. one a0 
pipersalata Giles 

plumiger Don. (Anoph.) 
poicilia Theob. 
Polylepidomyia Theob. 

Popea Ludl. 

powelli Ludl. ; 
pseudobarbirostris Ludl. 
Pseudocarvollia Theob. 
pseudodiurna Theob. oie 
Pseudograbhamia Theob.. .. 
Pseudograhamia Theob 
Pseudohowardina Theob. 
pseudolongifurcatus Theob. 
pseudomediofasciata Theob. 
pseudonivea Theob. 
pseudopictus Grassi ae 
Pseudoskusea Theob_ - ore 
pseudostenoetrus Theob. 
pseudotaeniata Giles 
Pseudotheobaldia Theob. 
Pseuaouranotaenia Theob. 
pseudowillmori Theob. 
pulcherrima Theob. 
pulchriventer Giles 

pulla Theob. 

pullus Theob. (Culex) 
punctolateralis Theob 
puuctulata Don. 


punctulata ‘+ James and List.’? 


pungens R. Desv 
pyginaeus Theob. sti 
Pyretophorus Blanch. 


quadripuuctis Ludl. 
quasiferox Leices. 
quasipipiens Theob. 
ouasisanguinae Leices. 
Quasistegomyta Theob. 
quasiunivittatus Theob, 


Rachionotomyia Theob. 
Radioculex Theob. 
Ramcia Annandale 
ratis Leices. 


Culex ; 30 
id. ais 

Reedomyia 

=Culex fatigans Ww. 

Pseudouranotaenia 

Culex 


Myzorhynchus 
Aioretomyia 
Stegomyia 
Stegomyia 
Culex 

Culex 


N yssorhynchus 
Runchomyia 


«* Anopheles ”’ 50 
Stegomyia 

=Myzorhynchus sinensis W: 
Finlaya Ae 


Uranotaenia 
Myzorhynchus 


Skusea i 


nom. nov. for Culex longifurcatus 474 


Skusea . 
Stegomyia 
Myzcrhynchus 
Culex 
Hulecoetomyia 


Nyssorhynchus 
Cellia 

Culex 

Culiciomyia 
=Culiciomyia pulla 
Stegomyia 

Cellia 

Myzomyia 

= Culex fatigans W. 
Chrysoconops 


* 


Pardomyvia 
Teromyia 
Culex 
Hodgesia 


Culex 


Teromyia 


456 


451 


* Index, 515 


Page 
Reedomyia Lud. “ip ise wa ae AGO 
reesii Theob.- ate he Culex oe ae AZ 4: 
rizali Banks ak fs id. 474 
rolandi D’Emm.d.Ch. -.. ee — Culex microannulatus Theob. 472 
rossii Giles .. Myzomyia Si ne ace 
rubithoracis Leices. as Fe Lophoceratomyia .. See. (ey! 
tubithoracis Leices. ae ..  Topomyia a SOO 
Runchomyia Theob. 2 ae are bea) 408 
ne * * 
samarensis Ludl. ae .. sub. sp. Scutomyia notoscripta 452 
samarensis Ludl. ; 7 Subesp. precmyiae scutellaris Wik. 449 
sanguinea Theob, oA -.  Hodgesia ee SOL 
sarawaki Theob. = ..  Oculeomyia is vet 405 
Sayomyia Coq. x. ss ate ae eS OS 
scatophagoides Theob. a ..  Mucidus te 440 
scintillans Ludl. (Heinzmannia) ..  Dendromyia Ric Sn 503 
scutellaris Wlk. Sie ae Stegomyia an wet AAS 
Scutomyia Theob 56 e a ume ABS 
seguini Laveran (Panoplites) ..Mansonia Ag ae eAG 2 
separatus Leices. : .. . Myzorhynchus ots BA Eee 
sepositus Leices. ag at Culex Be eA 
septemguttata Theob. Se ..- Mansonioides ae a" AS3 
septempunctata Theob. fe .. Mansonia 3c a AS 
sericeus Theob. be vee Culex 3 Sen ATS 
sexlineata Theob. : ..  Stegomyia ie -s 449 
simlensisJames and List. .. .. Anopheles 5A oe) 416 
simplex Theob. ae : Ficalbia 494 
simplex Theob. as ee vat. OF Neomacleaya indica Theob. 458 
sinensis Wied. Sc -- Myzorhynchus jot 4 ZO 
sinensis Theob. 2. .. Theobaldiomyia .. ~« - 463 
singularis Leices. ie ..  Aioretomyia i 3 490 
sitiens Wied. oe s- | Gulex Be ag. ee GAG 
Skeivomyia Leices ue Le fe a3 srt OO 
Skusea Yheob. aie a 5c ar Ao 4 cO 
sollicitans WIk. e ..  Grabhamia ne Pe AGL 
spathipalpis Rond. ie ..  Theobaldia oe -. 459 
spenceri Theob. y .. Grabhamia 4¢ a) 401 
splendens Meij. Age ..  Harpagomyia x3 Bee my eloy:! 
splendens Wied. a ..Toxorhynchites... Aa 
squamipenna Arrib. i ..  Aedeomyia cas et PASS 
Squamomyia ‘Theob. ae Ac Sc oe SS Ee) 
Stegomyia Theob. ae AD bie se) SAAS 
stenoetrus Theob. o .. Culex * et ATS 
stephensi Liston wa ..  Nyssorhynchus es Ago 
Stethomvia Theob. me ae “ Stale 
striocrura Giles i ..  Stegomyia xe -. 449 
subpictus Grassi a «* Anopheles ’’ nasa 
subulifer Dol. Be ..  TToxorhynchites.. 58 7437 
sugens W. sc ..  Seutomyia nag 152 
suknaensis Theob. rx ..  Culicada Ke 55-402 
sylvestris Leices. Be: .. Lophoceratomyia .. ne le 
; * * * 
taeniarostris Theob, Ae ..  Grabhamia a: ve 4O 
taeniata Leices. 5c ..  Aioretomyia fe 490 
taeniata Leices. -.  Lepidotomyia a se, 459 
taeniata Leices. ae Lophoceratomyia sae 405 
Taeniorhynchus Arrib =i ae a a Lo VE: 
taytayensis Banks Ss eee Culex: 4: =e egy As 
tenax Theob. af ..  Taeniorhynchus .. .. 480 
id. var. ocellata Theob. .. at id. ate ASO 
Tevomyta Leices. ar a a4 wip 438 
tessellata Theob. .. Myzomyia re | 422 
tessellatum Theob. (nom, nud. ) sc =Cellia punctulata Don. ea 432 
testacea Theob. Ap % Uranotaenia rs : 497 
theobaldi Giles ha .. Nyssorhynchus re he AZO 


Theobaldia Ney. Lemaire .. ae a ss an 1459 


516 Index. 


Theobaldinella Blanch. : re =Theobaldia 
Theobaldiomyvia Brun. nom. nov, for 
Leucomyta ae 
thomsoni Theob. Stegomyia % 
thorntoni Ludl, Myzomyia 56 
tigripes Grandpré AG Culex : 
tipuliformis Leices. ee Topomyia 
tipuliformis Theob. Culex 
Topomyia Leices. 
Toxorhynchites Theob. oe 
treacherii Leices. Anopheles as 
triangulata Ludi. abe ..  Pseudouranotaenia 
Trichopronomyia Theob. Me sts 
Trichorhynchomyia Brun. nom. nov. fo 
Trichorhynchus D0 

Trichorhynchus Theob. je 
trilineata Leices. in Theob. .. Hulecoetomyia 
trilineata Leices. Uranotaenia 
trilineatus Lheob. Culex 
trimaculatus Theob. Culex 
tripunctata Theob. Stegomyia 
tritaeniorhynchus Giles Culex 
turkhudi Liston Myzomyia 

* * * 
umbrosus Theob. Myzorhynchus 
uncus Theob. Culex 
uniformis Leices. Culex Be 
uniformis Theob. Lophoceratomyia .. 
uniformis Theob. Mansonia a6 
uniformis Theob. Melanoconion 
uniformis Theob, Skusea .. 
unilineata Leices. Uranotaenia 
unimaculata Leices. id. 
univittatus Theob. oni aCUlexe ye. 
Uvanotaenia Arrib. a sf 

* * * 
vagans Wied. Culex ~~. 
vanus Wlk. Myzorhynchus 
variata Leices. Lophoceratomyia 
varietas [,eices, Aioretomyia of 
Verrallina Theob ay 55 
vincenti Laveran ** Anopheles ’”’ 
viridiventer Giles Culex 
virilis Leices. Verrallina 
vishnui Theob. oe Culex 

“ * * 
w-alba Theob. Stegomyia 
watsoni Leices. Pyretophorus 
wellcomei Theob. Anopheles 
willmori James in Theob. Nyssorhynchus 
W orcestevia Banks ae 
Wyeomyia Theob. 

* * * 
Zeugnomyta Leices, : 

ERRATA. 


Page 
459 


462 
450 
422 
475 
500 
475 
499 
435 
417 
494 
477 


477 
477 
455 
498 
476 
476 
450 
476 
422 


427 
476 
476 
465 
483 
484 
451 
498 
498 
476 
495 


476 
427 
465 
491 
491 
434 
476 
492 
476 


450 
424 
417 
430 
439 
501 


498 


Page 438, line 10 from bottom. Foy ‘* magnificus’’ vead ‘* magnifica ’’ and line 
16 from bottom for ‘* funestus ’’ vead ‘* funesta.’’ 


Page 487, line 9 from bottom. Foy ‘“«Aedeomyinae ’’ vead ‘‘Aedinae.’’ 


1912. | E. Brunetti: Catalogue of Oriental Culicide. 517 


ADDENDA. 


The following species were unfortunately omitted from the 
foregoing Supplement to my Annotated Catalogue of Oriental 
Culicide : 
DACT YLOMYIA, gen. nov., Newstead and Carter, Ann. Trop. 

Med. iv. 377. ‘‘ near Anopheles.”’ 


D. ceylonica, sp. nov., id. id. loc. cit. 377. Ceylon. 


Myzomyia aurirostris, Watson, Ann. Trop. Med. iv. 251. Malay 
States. 


Stegomyia nigritia, Ludlow, Can Ent. xlii. 194 @. Philippine 
Islands. 


Scutomyia treubi, Meij. Ann. Jard. bot. Buitenzorg (1910) 3rd 
Supp., p. 922. Java. 


Culex aureopunctis, Ludlow, Can. Ent. xlii. 195, @ , a unique. 
Philippine Islands. 

CYATHOMYIA, gen. nov., near Finlaya. Meij. Ann. Jard. 
bot. Buitenzorg (1g10) 3rd Supp., p. 922. Java. 

@, jenseni, sp. nov,; id. loc..cit.gz2. Java. 


Popea lutea, Ludl., Can. Ent. xlii. 193, 2, described from a 
unique. 


Ficalbia tenax, id. loc. cit. 928. Java. 


Uranotaenia ascidiicola, id. loc. cit. g25. Java. 


3-4-1972. 


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SS wear = 7 ; us Su onaS 
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Culicidae and Corethridae in the Indian Museum. M¢scellanea :—Measurements 
of the skeletons of two large Indian elephants in the Indian Museum. The 
young of Aelurus fulgens. Some Batrachia recently added to the collection of 
the Indian Museum. Breeding habits of Tylototriton verrucosus. The occurrence 
of Rhinodon typicus at the head of the Bay of Bengal. Note on Ephydatia 
meyeni (Carter). 


Part IV.—Aquatic animals from Tibet, I. Aculeate Hymenoptera in the Indian 
Museum, I. Indian Psychodidae. A new species of mouse from the Madura 
District, Madras. Some Cleridae of the Indian Museum. ‘The Fauna of Brackish 
Ponds at Port Canning, Lower Bengal, XII. A new species of Saw-Fish captured 
off the Burma Coast. A new Sting Ray of the genus Trygon from the Bay of 
Bengal. New Microlepidoptera from India and Burma. Chrysomelid Beetles in 
the Indian Museum. Six new Cicindelinae from the Oriental Region, A new 
Slug from Tibet. 


Part V.—Revision of the Oriental Leptidae. Revised and annotated Catalogue of 
Oriental Bombylidae. 


Vol. TT 1909. 


Part I.—The races of Indian rats. 


Part II.—Freshwater Sponges, X. Aquatic animals from Tibet, II. Some amphibious 
Cockroaches. Quelques nouvelles Cécidomyies des Indes. New land and marine 
shells from Ceylon and S. India. Two new species of Cavanx from the Bay of 
Bengal. Some little known Indian Ophidia. Some forms of Dipsadomorphus. 
A pelagic Sea-Anemone without tentacles. Rhynchota Malayana, II. 


Part ITI.—The Neuroptera in the Indian Museum. New Indian Leptidae and Bom- 
bylidae. The Trichoptera in the Indian Museum. New species and varieties 
of Freshwater Crabs, 1—3. Lizards from Travancore. Three new Cicindelinae 
from Borneo. The relation between fertility and normality in Rats. A 
Barnacle of the genus Scalpellum from Malaysia. The Hemipterous family 
Polyctenidae. Freshwater Sponges, XI. Two new shells from S. India. A 
new genus of Phylactolaematous Polyzoa. Miscellanea :—Major Wall on some 
forms of Dipsadomorphus. Notes on Indian Batrachia. Notes on Indian Fresh- 
water Fish. Field notes on Indian Insects. The habits of Indian King-Crabs. 
The rate of growth in Conchoderyma and Lepas. large colonies of Hislopia lacus- 
tyis. Bvanchiocerianthus tmperatoy von der Kiiste von Oman und Baluchistan. 


Part IV.—A minute Hymenopterous insect from Calcutta. The Insect Fauna of 
Tirhut, No. 1. New species of Botia and Nemachilus. New Oriental Sepsinae. 
A new species of Fyredericella fr6m Indian lakes. New species and varieties of 
freshwater crabs, 4. Some new or little known Mygalomorph spiders from the 
Oriental region and Australasia. 


Vol. IV, 1910-1911. 


No. I.—Second report on the collection of Culicidae in the Indian Museum. 


Nos. II and III.—The Indian species of Papataci Fly (Phlebotomus). Taxonomic 
values in Culicidae. 


No. IV.—Revision of the Oriental blood-sucking Muscidae. 
No. V.—A new arrangement of the Indian Anophelinae. 


No. VI.—A revision of the species of Tabanus from the Oriental Region, including 
notes on species from surrounding countries. 


No. VII.—New Oriental Nemocera. Miscellanea :—Synonymy in Corethrinae. 
Indian Phlebotomi. 


Nos. VIII and I1X.—A revision of the Oriental species of the genera of the family 
Tabanidae other than Tabanus. Contributions to the fauna of Yunnan, Part VII. 


Vol. V, 1910. 


Part I.—The Hydroids of the Indian Museum, I. Freshwater Sponges, XII. New 
Shells in the Indian Museum from Burma, Siam and the Bay of Bengal. Mate. 
tials for a revision of the Phylactolaematous Polyzoa of India. Studies on the 
aquatic Oligochaeta of the Punjab. An undescribed Burmese Frog allied to Rana 
tigvina. Miscellanea :—The occurrence of Vultury monachus in Caleutta. An 
albino Owl. ‘‘ Matla bengalensis’’ : a correction. 


Part II.—Description d’Ophiures nouvelles provenant des derniéres campagnes de 
‘‘l’Investigator’’ dans 1l’Océan Indien. Description d’Holothuries nouvelles 
appartenant au Musée Indien. The races of Indian rats, II, A new species of 
Scalpellum from the Andaman sea. Five new species of marine shells from the 
Bay of Bengal. Fish from India and Persia. 


Pavt III.—A new genus of Psychodid Diptera from the Himalayas and‘ Travancore. 
The Indian barnacles of the subgenus Smiliwm, with remarks on the classification 
of the genus Scalpellum. A subspecies of Scutigerella unguiculata, Hansen, found 
in Calcutta. The distribution of the Oriental Scolopendridae. Decapoda in 
the Indian Museum, I. A new species of Nemachilus from Northern India. 
The larvae of Toxorhynchites immisericors, Wlk. A South Indian frog allied 
to Rana corrugata of Ceylon. Contributions to the fauna of Yunnan, 
Introduction and Part I. Miscellanea :—she Darjiling skink (Lygosoma sikkim- 
ense). Cockroaches as predatory insects. Note on Aedeomyia squammipenna, 
Arribalzaga. Named specimens of Chrysomelidae in the Indian Museum. - Two 
barnac'es of the genus Dichelaspis new to Indian seas. Slugs from the Eastern 
Himalayas. 


Part IV.—Indian Microlepidoptera. Some aquatic oligochaete worms commensal in 
Spongilla carteri. Bothrioneuvum ivis, Beddard. Nudibranchs from the Indian 
Museum. ‘The classification of the Potamonidae (Telphusidae). Catalogue of the 
pheasants, peafowl, jungle fowl and spur fowl in the Indian Museum. Species of 
Palaemon from South India. Alluaudella himalayensis, a new species of degener- 
‘ate (d') cockroach, with an account of the venation found in the genera Cardax 
and Alluatdella. Rhynchota Malayana, III. Ws 


Vol. Vi, 1911: 


Part I.—A Rhizocephalous Crustacean from fresh water aud on some specimens of 
the order from Indian seas. Decapoda in the Indian Museum, II. Contri- 
butions to the fauna of Yunnan, Parts II to V. Pedipalpi in the Indian 
Museum, I and II. Six new species of shells from Bengal and Madras. Mzis- 
cellanea :—Fleas from India and China. Flies found associated with cattle in 
the neighbourhood of Calcutta. Mosquito sucked by a midge. Large egg laid 
by a beetle. 


Pari II.—Some sponges associated with gregarious molluscs of the family Vermetidae. 
Aquatic animals from Tibet, III. Cyprinidae from Tibet and the Chumbi Valley, 
with a description of a new species oi Gymnocypris. New species and varieties 
of Crustacea Stomatopoda in the Indian Museum. The development of some 
Indian Ascalaphidae and Myrmeleonidae. Miscellanea:—The occurrence of 
Dactylopius cityi, Risso, in the Himalayas. Note on Aquatic Rhynchota. 


Part III,—Nouveaux Chironomides de 1’Indian Museum de Calcutta. 


Part IV.—Indian Isopods. Systematic notes,on the Ctenostomatous Polyzoa of 
fresh water. Some aquatic Oligochaeta in the Indian Museum. Contributions 
to the fauna of Yunnan, Part VI. Asiatic species of Crustacea Anostraca in the 
Indian Museum. Freshwater sponges, XIII]. Muscellanea:—Synonymy in 
Corethrinae. The distribution of the different forms of the genus Jd/a. 


Pari V.—Revision of the Oriental Tipulidae. Five new Indian species of Clavicorn 
Coleoptera. Les Chironomides (Tendipedidae) de |’Himalaya et d’Assam, The 
occurrence of Apus in Eastern Asia. 


Yol. VII, 1912. 


Part I.—Deep-sea Fish obtained by the R.I.M.S.S. ‘‘ Investigator ’’ during 1910-TT. 
Decapoda in the Indian Museum, III. Fauna of Paresnath Hill. Observations 
on the shallow-water Fauna of the Bay of Bengal made on the ‘‘ Golden Crown,’’ 
1908-09. Coccidae in the Indian Museum, I. On the nervous system of 
Ampullaria globosa. Description of a species of Phoridae that causes Myasis in 
man. Miscellanea:—Mimicry of a Mutillid by a spider. Capture of Limulus on. 
the surface. Development of the larva of Lingula, Distribution of some Indian 
and Burmese Lizards. 


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